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                    <text>Vocational Heavy Construction Technology Program
A Comprehensive Plan including Program Needs and Future
Directions
Matt Kondratieff
Committee Members:
Rod Van Velson,
Tom Bowen,
Larry Strohl
Colorado Division of Wildlife – Aquatic Research Section
April 2007

�EXECUTIVE SUMMARY
In 1997 Warren Diesslin, former Warden of the Buena Vista Correctional Facility, and Eddie
Kochman, former Colorado Division of Wildlife (CDOW) Aquatic Section Manager, met and
discussed a joint venture to rehabilitate degraded stream habitats while providing heavy
construction training for inmates sincere about changing the direction of their lives. These men
conceived and supported the vision of what is now known as the Vocational Heavy Construction
Technology (VHCT) program. Tom Bowen, once a prison guard and later a Colorado
Department of Corrections (CDOC) vocational educational instructor with years of practical
heavy construction experience, developed and coordinated this program with the support and
approval of Warden Diesslin. Tom contacted the Colorado Contractors Association (CCA) and
they agreed to serve as a program sponsor. The CCA has since become an integral part of the
program, serving as the advisory board and assisting student inmates with job placement once
they have successfully completed the program. Through the VHCT program, two state agencies
(CDOW and CDOC) and private industry have formed a rare partnership with different missions:
to help redirect human lives while restoring river natural processes and aquatic habitats within
driving distance of the Buena Vista Correctional Facility.
Student inmates have rehabilitated 8.7 miles of degraded aquatic stream habitats on CDOW
properties located along the South Platte River in South Park. South Park was identified as an
ideal location to implement the program because CDOW owns or leases over 25 miles of public
fishing waters in the Upper South Platte River drainage and its close proximity to the Buena
Vista Correctional Facility. Much of the South Platte River in South Park is degraded due to
excessive livestock grazing and mining. To date, 127 student inmates have graduated from the
VHCT program. Program recidivism rate is 12% compared to a 60% overall recidivism rate in
the Colorado penal system.
Through FY 2006-2007, CDOW/CDOC river restoration projects in South Park have cost a total
of $21/linear foot. A survey of six recent river restoration projects in Colorado conducted by
private companies range in cost from $61-$390/linear foot. The average cost for these privateindustry projects was $218/linear foot. The VHCT program realizes on average a 90% cost
savings, or up to 20 times less expensive than private industry. Private industry is also benefiting
by being able to hire from a pool of well-trained, highly qualified heavy equipment operators.
Cost of CDOW/CDOC projects in South Park range from $47,000-$148,000/year, with an
average cost of $108,865/year. The total cost for eight years of construction is $979,785.
Over 20 different habitat treatments have been implemented in South Park that fall within three
functional categories: restoring river natural processes, reducing bank erosion, and enhancing
aquatic habitat for sport fish. Treatments include the use of rock, stumps, logs and riparian
plants for bank re-vegetation.
CDOW personnel will work with the VHCT program again this year (FY 2007-2008), as well as
in the future, continuing ongoing efforts aimed at restoring degraded stream habitats in South
Park. However, a new research phase will begin that includes evaluation and monitoring of
restored aquatic habitats, quantifying how various stream habitat improvements translate into
positive changes in sport fish biomass, carrying capacity, improved angler opportunities, as well
as addressing other complex research questions.

i

�EXECUTIVE SUMMARY ............................................................................................................. i
TABLE OF CONTENTS................................................................................................................ ii
I.

INTRODUCTION ...............................................................................................................1
A.
Purpose of the Vocational Heavy Construction Technology Program....................1
B.
Why Restore Rivers? ...............................................................................................1

II.

VHCT PROGRAM BACKGROUND.................................................................................3
A.
History......................................................................................................................3
B.
Student Inmate Rehabilitation Program...................................................................5

III.

VHCT PROGRAM RESULTS............................................................................................5
A.
River Restoration. ....................................................................................................5
B.
Restoration of Inmate Lives...................................................................................10

IV.

PROGRAM COSTS AND BENEFITS .............................................................................10
A.
Inmate Incarceration ..............................................................................................10
B.
River Restoration Costs .........................................................................................10

IV.

COMMITTEE RECOMMENDATIONS ..........................................................................11
A.
Background ............................................................................................................11
B.
VHCT Program Expansion ....................................................................................11
C.
Research Opportunities..........................................................................................12
Sport Fish Enhancement ........................................................................................12
Improvements in Angler Use..................................................................................12
D.
Heavy Construction Equipment Needs..................................................................13
E.
Conclusions............................................................................................................14

VI.

CITATIONS ....................................................................................................................15

Appendix A: VHCT program informational brochure .................................................................16

ii

�I.

INTRODUCTION

A. Purpose of the Vocational Heavy Construction Technology Program
The Vocational Heavy Construction Technology (VHCT) program exists to provide student
inmates with education and training that will equip them with the basic life and work skills
necessary to obtain employment with a construction company once they have completed their
sentences. This program is a cooperative program between the Colorado Department of
Corrections (CDOC) and the Colorado Contractors Association (CCA). The Colorado Division
of Wildlife (CDOW) has been the major customer of the program, particularly in South Park
where natural river processes and aquatic habitats have been restored nearly nine miles of the
South Platte River.

B. Why Restore Rivers?
Overall health of river aquatic habitats generally deteriorate because of poor land use in the
watershed and riparian areas, accelerated stream bank erosion, poor water quality and stream
flow regimes altered by water use such as irrigation diversions or transport of water via the
stream for domestic purposes. When this occurs fish and aquatic habitats are degraded.
Past land practices have caused many portions of the South Platte River to become degraded in
South Park. Some of the major factors leading to the South Platte River’s current degraded state
include historic mining operations, channelization for irrigation and to accommodate railroads,
extermination of beaver, changes in the natural stream flow regime due to water impoundments
and long-term affects due to overgrazing.
River restoration projects can improve river bank stability, natural river processes and in-channel
aquatic and fish habitats. Our river restoration experience and studies indicate the most severe
cause of aquatic and trout habitat degradation can be traced to eroding stream banks and over
width river channels. Our river restoration projects concentrated in these two areas, encouraging
natural river processes.
Completed channel improvements in South Park have nearly doubled adult and juvenile brown
trout WUA (weighted useable area, Milhous et al. 1984). Trout biomass also increased almost
2.5 times with channel improvements. The increase in biomass was from new trout production
and migration of fish into newly created habitat.
In addition to these direct measures of success, the Statewide Fish Management Policy states
several principles related to river restoration.
1.

The long-term health of aquatic systems, including both habitat and
fisheries, is paramount.

2.

Providing recreational fishing opportunity will be an important objective of
the Colorado Division of Wildlife.

1

�3.

The protection of native species and their habitats is a priority.

Thus, it follows that both the protection of existing healthy fisheries habitat as well as restoring
degraded habitats back to a healthy state should be a high priority in regards to fisheries
management. River restoration activities not only rehabilitate existing degraded aquatic habitats
for fish, but also can potentially create new fish habitat where formerly no fish were present.
The goal of river restoration work is to repair degraded aquatic habitats and thus influence the
long-term health of aquatic systems. New recreational fishing opportunities are created either
through rehabilitation of degraded fish habitat (marginal habitats), enhancement of aquatic
habitat to address factors that limit fish populations, and/or creation of new aquatic fish habitat
where there once was no habitat at all. Therefore, the Statewide Fish Management Policy
provides a clear mandate for restoring streams and rivers in Colorado.
In addition to the Statewide Fish Management Policy, the Colorado Division of Wildlife 20022007 Strategic Plan states the following under F-1, Fishery Habitat Quantity and Quality:
“Healthy aquatic environments are essential to maintain healthy and viable fisheries, and
critical for self-sustaining populations. The Division desires to protect and enhance the
quality and quantity of aquatic habitats.”
The desired achievement, F-1.1, is to “protect and enhance existing quantity and quality of
habitats available to support fish populations.” One of the performance measures listed under
this desired achievement is to “Quantify performance targets for in-stream flows, conservation
pools, purchase or lease of water for aquatic habitat and habitat improvements.” Included under
Recommended Means are two statements:
1.

Protect the quality of habitat through working partnerships to achieve pollution
abatement, environmental protection, and physical habitat improvements.

2.

Take actions to minimize the negative impacts upon aquatic habitats
resulting from human activity.

The most recent statewide resident Angler Survey conducted in 2004 helps to direct priorities of
aquatic habitat restoration. The following information is from the 2004 Colorado Angler Survey
Summary Report (March 2006).
The 2004 Angler Survey reaffirms the importance of trout fishing in Colorado since seventy-six
percent of resident anglers list “trout” as their preferred fish. All of Colorado’s coldwater
habitats, including seasonally cold waters, are important in meeting the statewide demand for
trout fishing. Most of the restoration efforts conducted in Colorado have focused on coldwater
habitats containing trout, including all work conducted in South Park by through the VHCT
program.
When queried about where anglers prefer to fish, the majority of anglers of all license types say
they most often fish in mountain lakes, followed by coldwater streams and lakes in lower

2

�elevations. Almost all restoration work conducted through the VHCT program has involved
restoration of coldwater streams.
When licensed anglers were presented with a list of potential steps the CDOW could take to
encourage them and other anglers to fish more, the top most frequently mentioned steps (in order
of preference) by anglers were:
1. Improve the quality and size of fish.
2. Increase access to fishing locations on private land.
3. Improve fish habitat.
4. Stock more catchable trout.
5. Provide better information on where to fish.
Thus, the angling public recognizes the importance of fish habitat improvement as a way to
encourage themselves (and other anglers) to fish more. They even ranked fish habitat
improvement as more important than increasing the numbers of stocked catchable trout.

II.

VHCT PROGRAM BACKGROUND

A. History
The VHCT program began during the mid 1990’s after Warren Diesslin, former Warden of the
Buena Vista Correctional Facility, and Eddie Kochman, former CDOW Aquatic Section
Manager met several times, starting in 1997, and discussed a joint venture to rehabilitate
degraded stream habitats in South Park, CO while simultaneously providing heavy construction
training for inmates sincere about changing the direction of their lives.
Tom Bowen coordinated and developed the new VHCT program with the support and approval
of Warden Diesslin. While creating the program, Tom benefited from his past work experience,
including working as a prison guard and later as a DOC vocational educational instructor with
years of practical heavy construction experience. Tom had a vision for the program that would
enable inmates to more effectively transition from prison back to society with the hope of a
secure, well-paying job once they had completed the program. Tom’s belief was that out of such
a program, lives would ultimately be changed and recidivism reduced. Tom located a willing
program sponsor in the Colorado Contractors Association (CCA). Among other important
functions, the CCA serves as an advisory board to assist students with job placement once they
have completed the program. The VHCT program brought together two state agencies (CDOW
and CDOC) with different missions and a major trade association, the CCA, in a rare
partnership: to help redirect human lives while restoring river natural processes and aquatic
habitats within driving distance of Buena Vista (see VHCT program brochure, Appendix A).

3

�The first CDOW/CDOC river restoration project was completed in 1998 (Figure 1). The CDOW
paid for restoration materials including boulders, trees, and cobble; heavy equipment rentals;
inmate salaries (at $.60 per hour) and provided on-the-job technical assistance. This first river
restoration project using student inmates was highly successful in terms of enhancing aquatic
habitat for sport fish and restoring natural river processes. More importantly when these inmates
(about 20 students) were released from prison, they all found jobs in the construction industry
and were successful in turning their lives around.

Figure 1. Photo of VHCT inmates involved in the first restoration project in South Park, 1998.
Based on the early success of river restoration projects through the VHCT program, former
CDOW Director, John Mumma, earmarked $1,000,000 of capital construction monies for river
restoration projects in South Park over a five-year period. CDOW has continued funding river
restoration work in South Park through 2007. CDOW invested monies for restoration projects
because of the significant cost savings resulting from this program (up to 90% cost savings) and
because it allowed CDOW aquatic researchers (technical managers) opportunities to implement
on-the-ground creative ideas without expensive change-order charges during construction. This
project cost savings plus the valuable vocational training during river restoration projects made
this a win-win situation for inmates, the CDOW, CDOC and subsequently the private sector
contractors who hire heavy equipment operators.

4

�B. Student Inmate Rehabilitation Program
Only student inmates sincere about changing their lives and lacking heavy equipment experience
are admitted as candidates for the VHCT program. Student applicants must have a high school
education or equivalent and be approved through an interview process with the Program
Advisory Board (CCA). The program provides hands-on training related to operating heavy
equipment and obtaining necessary experience to gain employment within the construction
industry at a salary considerably higher than employment available to them prior to
incarceration. On-the-job training includes heavy equipment operation, heavy equipment
maintenance, surveying skills, blueprint reading, development of various additional construction
job skills, teamwork development, inmate behavioral changes and leadership training. The
VHCT program aids inmates in redirecting their lives once they are released from prison, as
evidenced by the reduced recidivism rates of inmates successfully graduating from the program.
The VHCT program is self supporting using the monies generated from contracts with
customers, primarily public entities.
The VHCT program works because it instills a work ethic which enables inmates to develop self
confidence on the job and in their lives, and it helps students develop individual job skills.
Alumni provide an important safety net to recent graduates during the first critical months after
leaving prison. The program has also served to build interpersonal relationships as alumni assist
recently released graduates by loaning them money to buy work clothing and safety equipment
for their jobs. They also assist newly released students to find jobs. Once inmates graduate from
the program and are released from prison, they live in a halfway house. CDOC program staff,
VHCT program alumni, and CCA members work together to assist inmates with securing a job
in the construction industry.
Besides the CDOW, the VHCT program also contracts for natural resource construction projects
with Federal agencies, the U.S. Fish and Wildlife Service, and U.S. Forest Service and public
entities including the Denver Water Department, Trout Unlimited and Park County-Upper South
Platte Coalition.

III.
A.

VHCT PROGRAM RESULTS
River Restoration

A major component of the VHCT program has been to restore natural river processes and instream aquatic habitats. To date, student inmates have rehabilitated 8.7 miles of degraded
aquatic stream habitats on CDOW properties located along the South Platte River in South Park
(Table 1). Stream habitat rehabilitation utilized 22 different habitat treatments that fall within
three functional categories: restoring river natural processes, reducing bank erosion, and
enhancing aquatic habitat for sport fish (Table 2). Treatments include the use of rock, stumps,
logs and riparian plants for bank revegetation. South Park was identified as an ideal location to
implement the program because CDOW owns or leases over 25 miles of public fishing waters in

5

�the Upper South Platte River drainage and because of its close proximity to the Buena Vista
Correctional Facility. Much of the South Platte River in South Park is degraded due to excessive
livestock grazing and mining. Through FY 2006-2007, CDOW/CDOC river restoration projects
in South Park have cost an average of $21/linear foot. A survey of six recent river restoration
projects conducted in Colorado by private companies range in cost from $61-$390/linear foot
(Table 3). The average cost for these private industry projects was $218/linear foot. Stream
restoration cost savings utilizing the VHCT program result in an average cost savings of 90%, or
up to 20 times less expensive than private industry.

6

�Table 1. CDOW stream restoration projects completed using the VHCT program including year of project completion, restoration
location, county, length of stream restored, costs, major treatments implemented, land ownership, and primary funding sources.
Length
(miles)

Project
Cost (total/per
Major Treatments
linear ft)

Ownership

Funding

Park

.2

$97,850 / $92.66

Colo. State Parks

CDOW/Cap. Const.

Park

.7

$139,610 / $37.77

Denver Water Dept.

2000

South Fork of South Platte
River
Threemile Creek

Park

.5

$138,000 / $52.27

DWD/CDOW Cap.
Const.
CDOW/Cap Const.

200l

South Platte River

Park

.9

$148,000 / $31.14

2002

South Fork of South Platte
River
South Fork of South Platte
River

Park

1.2

$146,000 / $12.57

Park

1.0

2003

South Platte River

Park

1.0

$128,725 / $24.38

2004

South Platte River

Park

.3

$47,000 / $29.67

2005

South Fork of South Platte
River

Park

1.7

2005

Tarryall Creek

Park

.6

2006

South Fork of Middle South
Platte River

Park

.6

Year

Stream

County

1998

South Platte River

1999

2002

Total

All Streams

$84,000 / $6.92

$50,600 / $15.97

Park

8.7

$979,785 / $21.33
7

Reduce channel width,
excavate pools, boulder &amp; log
placement
Excavate new channel,
Boulder and log placement
Constructed new flood
channel for Threemile Creek,
dam and retention lake.
Reduce channel width,
excavate pools, boulder &amp; log
placement
reduced channel width,
excavate pools
reduced channel width,
excavate pools , boulder &amp; log
placement
reduce channel width,
excavate pools, boulder &amp; log
placement
reduce channel width,
excavate pools, boulder
placement
Installed streamflow structures
and developed existing
channels, excavate pools
Reduce channel width,
excavate pools, willow &amp; log
placement
Reduce channel width,
excavate pools, boulder &amp; log
placement

&gt; 20 total treatments
applied

CDOW/Lower
Spinney SWA
CDOW/Lower
Spinney SWA

CDOW/Cap Const.

CDOW/
Knight/Imler SWA
CDOW/ Badger
Basin SWA

CDOW/Cap.
Const.
CDOW/Cap. Const.

Aurora/Colo. State
Park

CDOW/Cap. Const.

CDOW/Lower
Spinney SWA

CDOW/Cap. Const.

CDOW/Upper
Spinney SWA

CDOW/Cap Const.

CDOW/ Tarryall
SWA

CDOW/Cap Const.

CDOW/ Badger
Basin SWA

CDOW/Cap Const.

-

-

�Table 2. Benefits assigned to river channel and aquatic/trout habitat treatments used in
restoration projects.
Treatments to Improve Natural River Processes
Benefits
River Channel Treatment
Natural processes Reduces bank erosion
Aquatic habitats
Reduce river channel width
Primary
Secondary
Primary
Pool excavation
Primary
Secondary
Primary
Elevate riffle substrate
Primary
Limited
Primary
Woody overhead trout cover
Primary
Secondary
Primary
Riparian vegetation
Primary
Primary
Secondary
Riparian bench
Primary
Primary
Secondary
Woody Material Treatments Used to Reduce River Bank Erosion
Benefits
River Bank Treatments
Natural processes Reduces bank erosion
Aquatic habitats
Log spur
Secondary
Primary
Secondary
Log vane
Secondary
Primary
Secondary
Horizontal log
Secondary
Primary
Primary
River bank root wad
Secondary
Primary
Primary
Channel-edge log/root wad
Secondary
Primary
Primary
Boulder Treatments Used to Reduce River Bank Erosion
Benefits
River Channel Treatments
Natural processes
Reduces bank erosion Aquatic habitats
Cross vane
Secondary
Primary
Primary
Single boulder deflector
Secondary
Primary
Secondary
Hard point
Secondary
Primary
Limited
Boulder J hook
Secondary
Primary
Primary
Boulder vane
Secondary
Primary
Secondary
Treatments to Enhance Mid-Channel Aquatic and Trout Habitats
Benefits
Aquatic Habitat Treatments
Natural processes Reduces bank erosion
Aquatic habitats
Random boulders
Limited
Limited
Primary
Boulder clusters
Rock garden
Stumps
Mid-channel root wads
Off bank root wads

Limited
Limited
Limited
Limited
Limited

Limited
Limited
Limited
Limited
Secondary

8

Primary
Primary
Primary
Primary
Primary

�Table 3. Cost comparison of six major river restoration projects from Colorado with river
restoration costs using the VHCT program including stream name, river restoration
collaborators, miles of river restored, restoration cost, and reasons for restoration.

Stream name

Blue River

River restoration company/
organization
Northwest Colorado Council of
Governments (NWCCOG), Town
of Silverthorne, T.U., National
Forest Foundation, CDOW, and
Denver Water

Miles
restored

0.6

Cost per
linear foot

$61

Little Snake River

Dave Rosgen, Wildland Hydrology

10.5

$90

San Miguel River
(Phase I)

Town of Telluride (Public Works)

0.7

$200

Eagle River
(Edwards Eagle
River restoration)

Eagle River Watershed Council

1.6

$236

Lefthand Creek

CDOW, City of Longmont (Public
Works), Parks and Open Space,
CDOT, Longmont Power and
Communications, Carter &amp;
Burgess, Duran Excavating,
Aquatic and Wetlands Company,
and Property Owners

0.9

$333

0.4

$390

8.7

$21

West Ten-mile
Creek
South Platte River

CDOW/CDOC (VHCT program)

9

Reason for restoration
Enhance aquatic habitats
and channel
reconstruction

Enhance aquatic
habitats, channel
reconstruction, and
riparian revegetation
Restoration included:
creation of an instream
sedimentation basin,
implementing bank
stabilization treatments,
creating and improving
wetlands, developing
riparian habitats,
enhancing aquatic
habitat, and placing
instream hydraulic
structures.
Enhance aquatic
habitats, channel
reconstruction, and
riparian revegetation
Channel reconstruction,
floodplain reconnection,
and riparian revegetation

Channel reconstruction
and riparian revegetation
Restore natural river
processes, reduce bank
erosion, enhance
aquatic habitats

�B.

Restoration of Inmate Lives

Another part of the program’s mission is to develop student inmate work ethics necessary to
succeed in the industry as well as society. The VHCT program fosters an environment that
promotes integrity, trust, responsibility and confidence. The program encourages changes in
behavior, changes in the way students think and helps them to establish priorities and goals
necessary to get a “fresh start” in life. Inmates receive a minimum of 18 months of on-the-job
training in heavy equipment and maintenance. To date, 127 student inmates have graduated
from the VHCT program. Only 15 students have returned to prison because they violated
program rules. The program recidivism rate is 12% compared to a 60% overall recidivism rate
in the Colorado penal system. A major accomplishment that has taken place with this program is
the mentoring of recent VHCT program graduates from VHCT alumni. This is particularly true
for those students when they reach the halfway house.
Typical salary range for program graduates is $15.00-$18.00/hour with an annual salary of
$40,000-$50,000 (including overtime wages). For example, one program graduate is now a
project superintendent in Denver for a large construction company and has an annual salary of
$72,000/year. Another graduate started his own roofing company in Montrose and hires inmates
on release from local jails.

IV.

PROGRAM COSTS AND BENEFITS

A. Inmate Incarceration
On average, each inmate returning to prison costs taxpayers about $35,000 per year. The breakeven point of a $600,000 expenditure (cost for four pieces of heavy equipment) would be
reached when 17 former inmates were returned to prison for one year. The VHCT program has a
12% recidivism rate compared to 60% for the statewide penal system. A one-time cost of
$600,000 over a 10-year period appears to be a sound investment for this program because it has
reduced recidivism by 48%. The program has graduated 112 inmates that are now tax-paying
citizens living crime-free lives.

B.

River Restoration Costs

River restoration projects completed by this CDOW/CDOC program have provided savings
ranging from 31-90% over private industry costs. The actual improvement to the fishery, in
terms of increased biomass per mile or increase in the proportion of quality-sized fish has not
been well-estimated or studied. However, the number of angler hours that restored river
segments provide over un-restored segments is greatly increased (personal communication, Jeff
Spohn CDOW fisheries biologist). This change in the total number of angler hours (recreation
opportunity) from before and after river restoration projects has also not been well-studied and
only weak data exist to document changes in angler use in these areas. However, anglers have
expressed high satisfaction with their experience in fishing restored segments of the South Platte
River.

10

�IV.

COMMITTEE RECOMMENDATIONS

A. Background
During the winter and spring of 2007, committee members met on several occasions to discuss
VHCT program needs and recommendations for the future. The committee identified the
following conditions as necessary to ensure the VHCT program’s continued ongoing success. In
addition to the VHCT program’s contributions to restoring portions of the South Platte River,
CDOW research will focus on quantifying how habitat improvement work has increased angling
opportunities, increased the level of angler use and satisfaction, and how specific habitat
improvement treatments perform over time.

B.

VHCT Program Expansion

If the program owns its own heavy equipment, the cost savings could be passed on to other state
agencies like Colorado State Parks and Colorado Bureau of Mines and Reclamation. The Aurora
Water Department and Superfund sites in the Leadville area may also have an interest in the
services provided by this program. The program could also potentially be expanded to other
locations in Colorado that have candidate river restoration sites nearby a state correctional
facility (Rifle or Delta). While this paper focuses on aquatic program benefits, there are similar
opportunities for terrestrial programs to benefit from the VHCT program. The CDOC facility at
Sterling could be a prime example where medium security inmates can receive similar training.
A small program currently exists at Sterling, but CDOW has not yet contracted for services.
Future projects in South Park will continue to focus on the most degraded sections of the South
Platte River. A total of 7.2 miles of degraded river segments have been identified as candidate
sites that would benefit from CDOW/CDOC river restoration projects (Table 5).

11

�Table 5. Prioritized South Park stream channels segments and projects that would benefit from
CDOW/DOC restoration projects.

Stream

Length
(mile)

Primary
Treatment

2.5
1.0

Reduce channel width,
Excavate pools
Reduce channel width,
Excavate pools

Middle Fork/ South
Platte River
South Fork of
South Platte River
South Fork of
South Platte River
South Platte River

1.0

South Platte River

1.0

Tarryall Creek
(upstream from
Tarryall Res.)
Tarryall Creek
(Upper SWA
segment)
Total

.5

C.

1.0

.2

7.2

Reduce channel width,
Excavate pools
Reduce channel width,
Excavate pools
Reduce channel width,
Excavate pools
Design new stream
channel &amp; irrigation
diversion
Design trout passage
around an irrigation
diversion structure
-

Project
Description
Upper Spinney SWA/Lower end of
Badger Basin perpetual easement
River reach upstream of Badger
Basin HQ - Lower end of Badger
Basin perpetual easement
Badger Basin perpetual easement
adjacent to Hartsel town site
Lower Spinney SWA (Dream
Stream)
River segment downstream of Park
Co. Rd 59.
Construct new stream channel and
irrigation diversion.
Construct trout passage structure
over irrigation diversion
-

Research Opportunites

Sport Fish Enhancement
Future research opportunities associated with river restoration work in the South Platte include
the following: a comprehensive study that would seek to quantify how habitat improvements
have increased carrying capacity in streams, increased sport fish biomass, changed the number of
quality-sized sport fish and assisted in addressing limiting factors/conditions that might
otherwise preclude sport fish enhancement (in terms of total biomass). Determining whether
particular habitat treatments are more appropriate than others to maximize the benefits to sport
fish, whether particular habitat treatments would favor rainbow trout over brown trout (or favor
one particular fish species over another, including non-desirable species). Developing a
comprehensive plan for monitoring stream habitat improvements over time and developing
modeling techniques that would assist in addressing these questions (simulating various habitat
treatment scenarios under varying flow conditions, etc.).
Improvements in Angler Use
Another research objective would be to quantify how habitat improvements increase recreational
angling use and increase the total number of angler hours per stream mile. A program creel

12

�study may be appropriate to document angler use of restored versus un-restored stream segments.
This is important in determining how habitat improvement projects might increase angler use
annually, an important economic incentive to consider when planning future restoration projects.

D. Heavy Construction Equipment Needs
Field CDOC staff and CDOW biologists have identified four pieces of heavy construction
equipment that are needed to insure the program’s continued success. Currently, the most
expensive part of the river restoration projects (annually) is heavy construction equipment rentals
(Figure 3). Specific equipment needs include an excavator with an attached hydraulic thumb, a
front end loader, a backhoe with attached hydraulic thumb and a road grader. The estimated life
of this equipment could be as high as 10 years. The total cost for these four pieces of heavy
equipment would be approximately $600,000, depending upon price quotes and contributions by
the construction industry (Table 4).

Excavator w/ thumb
Front end loader
(tracked)
Backhoe w/ thumb
Motor grader
Dump truck
Cobble
Fuel
Labor

Figure 3. Typical costs associated with 3 month restoration project (based on FY 05-06 values).
Cost percentages include the following: heavy equipment rental = 81%, treatment materials =
9.5%, fuel = 8.5% and labor = 1%.

13

�Table 4. Equipment costs (purchase and lease) for 4 pieces of heavy construction equipment
necessary to perform stream restoration projects.
Heavy equipment

Annual lease

Purchase

Excavator w/ attached hydraulic thumb

$68,100

$189,750

Front-end loader

$58,000

$125,400

Road grader

$58,000

$200,000

Backhoe w/ attached hydraulic thumb

$36,000

$89,825

Total

$220,100

$604,925

If the VHCT program had its own heavy equipment, seat time for inmates would not be limited
by the short-time period that heavy equipment is leased. Inmates could spend more time learning
operational and maintenance skills as well as developing additional skills if the equipment was
available on-site and on a year-round basis.
E.

Conclusions

The VHCT program consists of a rare partnership between the CDOC, CDOW, and private
industry. The program has endured for nearly 10 years in spite of numerous obstacles and
challenges. Each party involved realizes a tremendous benefit from this relationship, as well as
society as a whole. The CDOC ultimately benefits by successfully rehabilitating inmates and
lowering the overall recidivism rate for prisoners in the Colorado penal system, the CDOW
benefits by improving degraded stream habitat and increasing angling opportunity and
satisfaction completed at a fraction of the expense required if using private contractors, the
private industry benefits from a pool of well-trained individuals from which to hire from, and
society as a whole benefits from the program when inmates are successfully rehabilitated and
reintegrated into society as law-abiding, tax-paying citizens holding down a steady job. A
unique program such as this deserves the attention and financial support necessary to ensure its
continued success.

14

�VI.

CITATIONS

Milhous, R. T., D. L. Wegner, and T. Waddle. 1984. User’s Guide to the Physical Habitat
Simulation System. Instream Flow Information Paper 11. U.S. Fish Wildl. Serv. FWS/OBS81/43 Revised. [475 pp.]

15

�Appendix A.

16

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                <text>In 1997 Warren Diesslin, former Warden of the Buena Vista Correctional Facility, and Eddie Kochman, former Colorado Division of Wildlife (CDOW) Aquatic Section Manager, met and discussed a joint venture to rehabilitate degraded stream habitats while providing heavy construction training for inmates sincere about changing the direction of their lives. These men conceived and supported the vision of what is now known as the Vocational Heavy Construction Technology (VHCT) program. Tom Bowen, once a prison guard and later a Colorado Department of Corrections (CDOC) vocational educational instructor with years of practical heavy construction experience, developed and coordinated this program with the support and approval of Warden Diesslin. Tom contacted the Colorado Contractors Association (CCA) and they agreed to serve as a program sponsor. The CCA has since become an integral part of the program, serving as the advisory board and assisting student inmates with job placement once they have successfully completed the program. Through the VHCT program, two state agencies (CDOW and CDOC) and private industry have formed a rare partnership with different missions: to help redirect human lives while restoring river natural processes and aquatic habitats within driving distance of the Buena Vista Correctional Facility.</text>
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                <text>Monitoring activities to evaluate restoration effectiveness for the upper Arkansas River Habitat Restoration Project were conducted by Colorado Parks and Wildlife (CPW) and contractors during 2020 and 2021. Efforts were primarily focused on data analysis and publication of results for monitoring targets, including fish populations, riparian vegetation, benthic macroinvertebrates, instream habitat structures, and water quality. Some fish population, benthic macroinvertebrate, riparian vegetation and habitat metrics improved following restoration, although not all metrics have achieved target goals. Significant improvements in Brown Trout Salmo trutta density, biomass and condition were encouraging, but apparent declines in quality trout could be indicative of increased competition or limited forage. The abundance of benthic macroinvertebrates increased, but not to the level of project goals. Woody riparian vegetation increased significantly, and encroachment of riparian vegetation has outpaced bank erosion, which suggests that bank stability has improved and the channel is moving towards dynamic equilibrium. Multiple metrics indicate that ecosystem health within the California Gulch Superfund Site continues to improve.</text>
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                    <text>DRAFT PROPOSED
Upper Colorado River Headwaters Project Monitoring Plan

Prepared by
Colorado Parks and Wildlife

March 19, 2018

1

�Table of Contents
Chapter 1: Project Overview

4

1.1

Introduction

4

1.2

Windy Gap Connectivity Channel Project

4

1.3

Kemp Breeze State Wildlife Area (SWA) Habitat Restoration Project

6

1.4

Irrigators of Lands in the Vicinity of Kremmling (ILVK) Project

8

1.5

Monitoring Overview

9

Chapter 2: Fish Populations
2.1

10

Introduction

10

Sportfish Evaluations

10

Native Fish Evaluations

12

2.2

Methods

13

Population Estimates for Trout Fry

13

Population Estimates for Adult Trout

14

Population Estimates for Mottled Sculpin

14

2.3

Study Sites

14

Trout Fry

14

Adult Trout

15

Mottled Sculpin

15

2.4

Monitoring Schedule

18

Trout Fry

18

Adult Trout

18

Mottled Sculpin

18

Chapter 3: Benthic Macroinvertebrates

18

3.1

Introduction

18

3.2

Methods

20

3.3

Study Sites

20

3.4

Monitoring Schedule

21

Chapter 4: Hydrology, Hydraulics, and Geomorphology

21

4.1

Introduction

21

4.2

Methods

22
2

�Hydrology

22

Hydraulics

22

Geomorphology

23

4.3

Study Sites

24

4.4

Monitoring Schedule

24

Chapter 5: Monitoring Budget

24

References

25

Appendix A: Proposed Geomorphic Monitoring of the Connectivity Channel

28

A.1

Introduction

28

A.2

Methods

28

Hydrology

28

Hydraulics

29

Geomorphology

29

A.3

Study Sites

30

A.4

Monitoring Schedule

30

A.5

References

30

Appendix B: Proposed Fish Movement Study

32

B.1

Introduction

32

B.2

Methods

33

B.3

Study Sites

36

B.4

Monitoring Schedule

37

B.5

References

38

3

�Chapter 1: Project Overview
1.1

Introduction

In December 2016, a group of partners including American Rivers, Colorado Parks and Wildlife
(CPW), Colorado River Water Conservation District (CRWCD), Colorado Water Conservation
Board (CWCB), Denver Water, Grand County, Irrigators of Lands in the Vicinity of Kremmling
(ILVK), Municipal Subdistrict of Northern Colorado Water Conservancy District (Northern
Water), Trout Unlimited, and the Upper Colorado River Alliance was awarded $7.75 million by
the U.S. Department of Agriculture Natural Resources Conservation Service (NRCS) through their
Regional Conservation Partnership Program (RCPP) for the Upper Colorado River Headwaters
Project (Headwaters Project). The Headwaters Project is comprised of three endeavors within
Grand County that will collectively restore fish and wildlife habitat, and improve water quality
and agricultural water management on a regional scale. The RCPP funding will be utilized to
focus on two specific project components: 1) reconnecting the Colorado River upstream and
downstream of Windy Gap Reservoir (referenced as the Windy Gap Connectivity Channel
Project); and 2) restoration of the Colorado River channel to be resilient to hydrological
modifications, while sustaining agriculture, and aquatic and riparian habitat (referenced as the
Irrigators of Lands in the Vicinity of Kremmling Project). The third endeavor, the Kemp Breeze
State Wildlife Area (SWA) Habitat Restoration Project, is expected to be funded by the partners,
and other interested parties. These three project components are further described in the following
sections.
1.2

Windy Gap Connectivity Channel Project

The Windy Gap Connectivity Channel Project was selected among several alternatives evaluated
to improve degraded habitat of the Colorado River downstream of Windy Gap Reservoir (Tetra
Tech 2015). This project will create a new river channel around Windy Gap Reservoir,
reconnecting the Colorado River upstream and downstream of the reservoir, while simultaneously
preserving water storage for Northern Water’s municipal water supply. The new channel will be
designed to mimic natural conditions, including increasing habitat heterogeneity with the addition
of riffles, pools, and runs that accommodate upstream and downstream passage of juvenile and
adult life stages of Rainbow Trout (Oncorhynchus mykiss), Brown Trout (Salmo trutta) and
Mottled Sculpin (Cottus bairdii). Reconnection will also expand available wildlife habitat, reestablish sediment transport, and improve water quality conditions downstream of the project area.
The Windy Gap Connectivity Channel Project will be primarily managed by Northern Water, in
collaboration with other partners, and will entail a three-phase approach to plan, design, and
implement/monitor/maintain within a five to six year period, beginning in 2017. The details and
projected timeline are described below in Table 1.1.

4

�Table 1.1. Proposed process and schedule for the Windy Gap Connectivity Channel Project.
Step
1. Planning

Tasks

Timeline

Identify goals and objectives

2017-2018

Evaluate goals and limiting factors

2017-2018

Develop watershed plan and NEPA 2017-2018
Easement and ROW

2018-2019

Tasks

Timeline

Step
2. Design

3. Permitting

Survey and analysis

2017-2019

Evaluate alternatives

2017-2019

Conceptual design

2017-2019

Preliminary design

2017-2019

Final design

2017-2019

404 permit and others

2017-2020

Step
4. Implementation

Tasks

Timeline

Contract documents

2019-2020

Construction

2020-2021

Baseline

1980-2020

Implementation

2020-2021

Effectiveness

2021-2026

6. Adaptive Management Maintenance

2021-2026

5. Monitoring

Information dissemination

5

2021-2026

�1.3

Kemp Breeze State Wildlife Area (SWA) Habitat Restoration Project

The original “Habitat Project” was designed in coordination with Northern Water to address
concerns raised by CPW and other stakeholders regarding conditions of the aquatic ecosystem in
the Colorado River downstream of Windy Gap Reservoir. The goal of the Habitat Project is to
design and implement a stream restoration program to improve the existing aquatic environment
in the Colorado River from the Windy Gap diversion to the lower terminus of CPW’s Kemp Breeze
State Wildlife Area (SWA) by returning the river to a more functional system considering current
and future hydrology. The intent is for Denver Water and Northern Water to join with CPW and
other stakeholders in a cooperative effort to identify and address desired improvements to the
stream environment. To guide project implementation, the Denver Board of Water Commissioners
and CPW signed an Intergovernmental Grant Agreement (IGA) to implement the Habitat Project
on March 26, 2014. Northern Water and CPW signed a parallel IGA on April 3, 2014.
Though the IGAs describe the large-scale Habitat Project (including a project area of
approximately 16.7 miles of the Colorado River), the 2017 NRCS RCPP Headwaters Project will
focus entirely on CPW’s Kemp Breeze SWA Habitat Restoration Project. This project area will
include a smaller scale reach (approximately 2.7 miles) contained within the larger geographical
area of the overall Habitat Project, where CPW can complete planning, design, and implementation
of the Kemp Breeze Habitat Restoration Project within the five to six year time frame required by
the NRCS. The lowermost segment within the Kemp Breeze Habitat Restoration Project includes
the Williams Fork River, the largest tributary in the project area. The Colorado River through the
Kemp Breeze SWA is owned almost entirely by the State of Colorado, and is likely the most
heavily fished section of the Colorado River in Grand County. Improving habitat conditions within
the Kemp Breeze SWA reach will include defining the thalweg, targeting channel narrowing
through point-bar enhancement, de-armoring riffles, and increasing habitat heterogeneity. This
section will likely also benefit from some of the most intensive channel work required in the largescale Habitat Project upstream.
The Kemp Breeze SWA segment has been identified as the highest priority for habitat
improvement when compared to seven other reaches within the Habitat Project. The rationale for
this assessment was based on several factors, including the need for restoration, increased public
fishing access, lack of complex landowner agreements, proximity to Windy Gap Reservoir, and
preliminary cost estimates. The location of the Kemp Breeze SWA as the segment farthest away
from potential unknown impacts of the proposed Windy Gap connectivity channel was a huge
consideration in the prioritization process. Construction of the proposed Windy Gap connectivity
channel will likely be on a similar schedule as the Kemp Breeze Habitat Restoration Project.

6

�Table 1.2. Proposed process and schedule for the Kemp Breeze SWA Habitat Restoration
Project*.
Step
1. Planning

2. Prioritize actions

Tasks
Identify goals and objectives

2017-2018

Evaluate goals and limiting factors

2017-2018

Site selection

2018-2019

Step
3. Design

4. Permitting

Tasks

6. Monitoring

7. Adaptive
Management

Timeline

Survey and analysis

2018-2019

Evaluate alternatives

2018-2019

Conceptual design

2018-2019

Preliminary design

2019-2020

Final design

2019-2020

404 permit

2020

Floodplain permit

2020

Landowner agreements

2020

Step
5. Implementation

Timeline

Tasks

Timeline

Contract documents

2019-2020

Construction

2020-2021

Baseline

1980-2020

Implementation

2021-2023

Effectiveness

2023-2028

Maintenance

2023-2028

Information dissemination

2023-2028

* This table reflects implementation of habitat restoration on the Kemp Breeze SWA only, and not
the entire Habitat Project.

7

�The Kemp Breeze Habitat Restoration Project will be primarily managed by CPW in collaboration
with the large-scale Habitat Project Stream Team, which includes members from the CRWCD,
Denver Water, Grand County, Northern Water, Trout Unlimited, and other parties including but
not limited to private landowners. The restoration process will entail a three-phase approach to
plan, design, and implement/monitor/maintain the Kemp Breeze Habitat Restoration Project
within a five to six year period, beginning in 2017. The details and projected timeline are described
below in Table 1.2.
1.4

Irrigators of Lands in the Vicinity of Kremmling (ILVK) Project

In general, most Irrigators of Lands in the Vicinity of Kremmling (ILVK) experience difficulty in
retrieving or using water during low flow periods in late summer, especially during drought
periods. Low water level creates a lack of positive pressure in ditch heads, making irrigation
systems more difficult to operate. Measures to address these issues, including the installation of
pumps and cobble dams, have failed to fully address the problem, and have resulted in several
unintended consequences, including increased soil erosion and instream habitat modification.
Eleven private ranches are participating in the ILVK Project by improving the Colorado River
channel through a series of practices utilizing the NRCS Environmental Quality Incentives
Program (EQIP). These practices will create and install innovative in-stream structures designed
to improve water levels for irrigation, while rebuilding riffle and pool structure to enhance river
habitat. Structures include grade control riffles, riffle vanes, sustainable habitat riffles, point bar
creations, habitat structures, and pool enhancements. Several pilot projects within the ILVK area
have already been completed, and the results to date indicate success beyond expectations.
The ILVK Project will be primarily managed by Trout Unlimited in collaboration with other
partners, and will entail a three-phase approach to plan, design, and implement/monitor/maintain
within a five to six year period, beginning in 2017. The details and projected timeline are described
below in Table 1.3.
Table 1.3. Proposed process and schedule for the ILVK Project.
Step
1. Planning

2. Producer outreach
and recruitment

Tasks

Timeline

Identify goals and objectives

2017-2018

Evaluate goals and limiting factors

2017-2018

Site selection

2017-2018

8

�Step
3. Producer participant
selection;
contracting with
producers

4. Design

5. Permitting

Tasks
Survey and analysis

2018-2019

Evaluate alternatives

2018-2019

Landowner agreements and
contracts

2018-2019

Conceptual design

2017-2019

Preliminary design

2017-2018

Final design

2018-2019

404 permit and others

2018-2019

Step

Tasks

Timeline

6. Implementation

Construction

2019-2022

7. Monitoring

Baseline

2017-2019

Implementation

2019-2022

Effectiveness

2019-2024

Maintenance

2019-2024

Information dissemination

2019-2024

8. Adaptive
Management

1.5

Timeline

Monitoring Overview

The NRCS requests a Monitoring Plan be developed as part of the RCPP process. This document
has been developed by CPW in collaboration with the other partners to comply with this
requirement for the Headwaters Project. The Monitoring Plan is considered a “living document”
with the flexibility to incorporate necessary schedule updates and other relevant modifications as
the Headwaters Project proceeds.
Multiple partners are completing various monitoring activities (primarily of the chemical and
physical attributes of the upper Colorado River drainage) related to their respective Headwaters
Project components and permitting requirements. Those efforts, some of which may be directly
connected to the Headwaters Project, are not included within this document. Rather, this plan
9

�focuses on the monitoring that CPW has committed to with regard to labor and equipment within
the RCPP. Though monitoring is included for each of the three components of the Headwaters
Project, most of these efforts by CPW will be related to the Windy Gap Connectivity Channel and
Kemp Breeze Habitat Restoration Project, with limited fish monitoring within the ILVK Project
area. Pre-project (baseline) and post-project evaluations are incorporated within the plan.
Specifically, CPW will primarily be concentrating on fish populations (Chapter 2), benthic
macroinvertebrates (Chapter 3), and hydrology, hydraulics, and geomorphology (Chapter 4) of the
upper Colorado River. CPW will evaluate elements included within Chapter 4 with an emphasis
on the Kemp Breeze Habitat Restoration Project; hydraulic modeling, sediment transport surveys,
and assessment of the Windy Gap connectivity channel can be investigated should supplementary
funds be secured or if monitoring priorities are changed during discussions with project
stakeholders. Thus, CPW has included Proposed Geomorphic Monitoring of the Windy Gap
connectivity channel in Appendix A. Additionally, CPW has developed a Proposed Fish
Movement Study in Appendix B that may also be pursued should additional funding and staff time
become available.

Chapter 2: Fish Populations
2.1

Introduction

Sportfish Evaluations
Prior to the introduction of whirling disease, caused by the parasite Myxobolus cerebralis, in the
upper Colorado River, adult Colorado River Rainbow Trout (CRR) had an average abundance of
689 fish/mile while adult Brown Trout averaged 385 fish/mile (Nehring and Thompson 2001),
resulting in a ratio of Rainbow Trout to Brown Trout of 2:1. Rainbow Trout fry abundance ranged
from 9,012 to 13,518 fry/mile of stream bank and Brown Trout fry ranged from 4,184 to 9,173
fry/mile (Walker and Nehring 1995). Traditionally, eggs were harvested by CPW from this wild
CRR brood stock, reared in state hatcheries, and used to stock many rivers across the state.
However, the CRR was one of the most susceptible strains of Rainbow Trout to whirling disease,
developing over 150,000 myxospores/fish (Fetherman et al. 2012).
Myxobolus cerebralis was unintentionally introduced to the upper Colorado River in the 1980s
when privately-reared Rainbow Trout previously exposed to M. cerebralis were stocked into three
private water bodies located upstream of Windy Gap Reservoir. Fish downstream of Windy Gap
Reservoir tested positive for M. cerebralis in 1988, and a subsequent decline in the younger age
classes of Rainbow Trout was observed in the early 1990s (Nehring 2006). While several reasons
for the declines were investigated (Schisler et al. 1999a; Schisler et al. 1999b; Schisler et al. 2000),
exposure to M. cerebralis was determined to be the primary cause for the disappearance of the
younger age classes (Nehring and Thompson 2001). In an effort to restore the Rainbow Trout
fishery, tens of thousands of CRR were stocked annually between 1994 and 2008. Despite these
repeated stocking efforts, the CRR exhibited low survival and little recruitment success, resulting
10

�in a Rainbow Trout abundance that was approximately 90% lower than the Rainbow Trout
abundance observed prior to the establishment of M. cerebralis (Nehring 2006).
Whirling disease-resistant Rainbow Trout were first introduced to the upper Colorado River in
2006, with an additional introduction occurring in 2010, using a strain of Rainbow Trout known
as the HxC. The HxC is a cross between the whirling disease-resistant, but domesticated Rainbow
Trout strain known as the German Rainbow (GR), or Hofer, strain and the CRR. The HxC cross
was chosen for these introductions because they exhibited resistance characteristics similar to the
GR strain (Schisler et al. 2006; Fetherman et al. 2012), and were capable of attaining critical
swimming velocities similar to those of the CRR strain (Fetherman et al. 2011). Larger Rainbow
Trout (&gt; 6.69 inches TL) were used in these first introductions because larger fish were: 1) less
susceptible to M. cerebralis infection (Ryce et al. 2005), and 2) less susceptible to Brown Trout
predation. However, despite these potential survival advantages, the apparent survival rate of the
introduced HxC cross over the entire study period between 2006 and 2011 was estimated to be
only 0.7%, and the adult Rainbow Trout population in the upper Colorado River continued to
decline. Although survival of the introduced HxC cross was low, these fish did reproduce in the
river, resulting in genetic shifts in the fry population toward GR-cross fish and a decrease in
average myxospore count of the fry population over time (Fetherman et al. 2014).
One potential reason for the low survival in the introduced HxC cross was the length of time these
fish were held in the hatchery to reach the larger sizes needed prior to stocking. It was thought
that the longer these fish were held in the hatchery, the more they acted like the domestic GR rather
than the wild CRR upon stocking. Stocking fish as fry was thought to potentially counteract these
effects because fish would be held in the hatchery for a shorter period of time prior to stocking.
The HxC cross was first stocked into the Colorado River as fry in 2013, with additional stocking
occasions in 2014 and 2015. Since fry stocking began, survival and recruitment from fry to adult
life stages have increased in the upper Colorado River, and the adult Rainbow Trout population
has exponentially increased from a low of three adult fish/mile in 2013 to 165 adult fish/mile in
2017. In recent years (2016 and 2017), fry stocking has continued using pure GR strain fish since
these fish exhibit similar apparent survival rates to the HxC cross, and to reduce resistance issues
due to backcrossing and outcrossing that could occur when using the HxC cross for Rainbow Trout
reintroductions (Avila et al. In review).
The objective of this project component is to re-establish a wild, and ultimately, self-sustaining
Rainbow Trout population in the upper Colorado River. A variety of stocking techniques,
including manipulating size-at-stocking and strain stocked, have been used to achieve this
objective. The goal of the fry stocking in recent years has been to determine if stocking GR and
GR-cross fish as fry increases post-stocking survival, and recruitment to the adult Rainbow Trout
population, and if overall M. cerebralis infection prevalence and severity decreases with an
increase in abundance of whirling disease-resistant Rainbow Trout.

11

�Native Fish Evaluations
Sculpin are an ecologically important part of freshwater ecosystems because they can occur in high
densities in depauperate coldwater mountain streams (Adams and Schmetterling 2007).
Additionally, sculpin can exert a large influence on aquatic food webs through their diverse trophic
positions. The Mottled Sculpin (Cottus bairdii) is common in coldwater western Colorado streams
where they occur in sympatry with important sport and native trout species. Mottled Sculpin prefer
cool, high gradient mountain streams with cobble habitat and are rarely found in stream reaches
where substrate is embedded with silt (Sigler and Miller 1973; Woodling 1985; Nehring et al.
2011). As such, their habitat preferences for cobble substrate and high quality riffle-run habitat
make Mottled Sculpin a good ecological indicator of stream health (Adams and Schmetterling
2007; Nehring et al. 2011).
Dams are known to drastically alter river habitat and have many diverse effects on fish and
invertebrate habitat and populations (Ward and Stanford 1979). Dams can radically alter stream
temperature and substrate composition, which are considered the primary influences of sculpin
habitat suitability (Scott and Crossman 1973). In the upper Colorado River basin, stream reaches
below many dams and water projects have reduced the density of Mottled Sculpin (Nehring et al.
2011). The decline in sculpin distribution appears to be both temporally and spatially related to
impoundments. Mottled Sculpin were common in the main stem Colorado River before Windy
Gap Reservoir was built, but are rare or absent after construction (Erickson 1983; Nehring et al.
2011). A survey conducted in 1975-1976 on the Colorado River before Windy Gap Reservoir was
constructed documented Mottled Sculpin at all sampling sites (Dames and Moore 1977). In 2010,
a project investigating the sculpin distribution and density throughout the upper Colorado River
revealed that sculpin density was on average 15 times higher in sites upstream of impoundments
than downstream (Nehring et al. 2011). In the main stem of the Colorado River between Windy
Gap Reservoir and the confluence with the Williams Fork River, a single Mottled Sculpin was
encountered in 3,200 ft of river sampled by electrofishing. This study attributed the decline of
Mottled Sculpin in the upper Colorado River downstream of Windy Gap Reservoir to habitat and
flow changes associated with the reservoir. Surveys conducted by CPW in 2013 confirmed these
patterns, finding that sculpin were common upstream of impoundments on the upper Colorado
River, but rare or absent downstream (Kowalski 2014). As part of this study, three sites were
sampled on the Colorado River between Windy Gap Reservoir and the Williams Fork River
confluence, and no Mottled Sculpin were found.
While Mottled Sculpin were once common downstream of Windy Gap Reservoir, and remain
common upstream, habitat alterations associated with the reservoir seem to have reduced the
habitat quality in the river to a point where viable populations are rare or nonexistent. Restoring
connectivity and addressing habitat limitations associated with the flow and sediment regimes
should improve conditions for this important native fish in the upper Colorado River. The
objective of this segment of the project is to document how the Windy Gap connectivity channel

12

�affects the distribution and density of Mottled Sculpin in the Colorado River downstream of the
Fraser River confluence.
Goals
● Re-establish a wild, self-sustaining Rainbow Trout population in the upper Colorado River
● Re-establish a wild, self-sustaining population of native Mottled Sculpin in the Colorado
River downstream of Windy Gap Reservoir
Objectives
● Evaluate the effects of fish stocking techniques on Rainbow Trout post-stocking survival
and recruitment to the adult population
● Determine if the overall M. cerebralis infection prevalence and severity decreases with an
increase in abundance of whirling disease-resistant Rainbow Trout
● Document the distribution and density of Mottled Sculpin over time in the Colorado River
downstream of the confluence with the Fraser River
2.2

Methods

Population Estimates for Trout Fry
GR and GR-cross fish are stocked by CPW as post-swim-up fry in the margins of the river between
Hitching Post Bridge and either the Chimney Rock Ranch diversion structure near the confluence
with Drowsy Water Creek or the Sheriff Ranch, dependent upon annual fry availability in July. In
previous years, 70,000 to 250,000 fry have been stocked into this section of the river. Fry are
loaded into a well-aerated tank on a raft and walked or slowly floated downriver and evenly
distributed throughout optimal fry habitats on both sides of the river. Seven fry sites are sampled
in the Colorado River annually by CPW to obtain fry abundance estimates, genetic composition,
and for myxospore enumeration (see Table 2.1, Sections 2.3 and 2.4, and Figures 2.1, 2.2, and
2.3). All seven fry sites are sampled at least once prior to fry stocking to determine if wild
reproduction of Rainbow Trout and Brown Trout has occurred. Genetic samples in the form of a
caudal fin clip (top lobe) are collected from all wild Rainbow Trout fry encountered to determine
the percentage of GR genes in the wild fry population. The same seven fry sites are sampled once
a month from July through October to determine abundance and survival of wild Brown Trout and
stocked Rainbow Trout fry in the Chimney Rock Ranch/Sheriff Ranch reach, or wild Brown Trout
and Rainbow Trout fry in the sites downstream of Byers Canyon. Three-pass depletion population
estimates are completed using two, Smith-Root LR-24 backpack electrofishing units running sideby-side to cover all available fry habitat at each site. Genetic samples are collected from all
Rainbow Trout fry that can be identified as wild. In October, five Brown Trout and five Rainbow
Trout fry are collected from each site for myxospore enumeration via the pepsin-trypsin digest
(PTD) method using whole fish samples.

13

�Population Estimates for Adult Trout
Adult trout populations in the Colorado and Fraser rivers are monitored by CPW at several sites
on a varying schedule (see Table 2.1, Sections 2.3 and 2.4, and Figure 2.1). Two-pass markrecapture adult population estimates are conducted using raft-mounted, fixed-boom electrofishing
units on the Chimney Rock Ranch, Paul Gilbert-Lone Buck, Parshall-Sunset, and ILVK reaches.
All fish captured on the mark run are given a caudal fin punch, measured to the nearest millimeter,
and returned to the river. At least one day is left between the mark and recapture runs to allow
redistribution of marked fish. On the recapture run, fish are examined for the presence of caudal
fin punches, measured to the nearest millimeter, and weighed to the nearest gram. Up to sixty
genetic samples are collected from Rainbow Trout encountered on the Chimney Rock Ranch reach
to determine the proportion of GR genes present in the adult spawning population, and to correlate
this proportion with that observed in the wild fry population. On rivers that are too small to float
a raft, such as the Fraser and the Colorado rivers upstream of Windy Gap Reservoir, two-pass
depletion population estimates are conducted using a Smith-Root 2.5 GPP bank electrofishing unit
and wade electrofishing crews.
Population Estimates for Mottled Sculpin
Mottled Sculpin will be sampled in the fall annually by CPW at eight sites utilizing presenceabsence and three-pass depletion electrofishing (see Table 2.1, Sections 2.3 and 2.4, and Figures
2.2 and 2.3). Sampling will be completed using either Smith-Root LR-24 backpack electrofishing
units or a Smith-Root 2.5 GPP bank electrofishing unit. Presence-absence sampling sites will
range from 50-500 feet long depending on channel dimensions and fish density. Sites in which
Mottled Sculpin are present will range from 50-100 feet long, and will be sampled using threepass depletion electrofishing for abundance determination. At sites where Mottled Sculpin are not
documented in the first pass, sampling will continue upstream for up to 500 feet in an attempt to
detect sculpin over larger river reaches.
2.3

Study Sites

Trout Fry
GR and/or GR-cross fry stocking by CPW occurs between Hitching Post Bridge and either the
diversion structure on the Chimney Rock Ranch near the confluence with Drowsy Water Creek or
the Sheriff Ranch, dependent upon annual fry availability. CPW completes fry population
estimates at seven sites, four upstream of Byers Canyon, and three downstream. Sites upstream
of Byers Canyon include: Hitching Post (13T 414502, 4440208), Upper Red Barn (13T 412749,
4439680), Lower Red Barn (13T 412287, 4439359), and Sheriff Ranch (13T 409063, 4438050).
Sites downstream of Byers Canyon include: Paul Gilbert (13T 403510, 4433781), Lone Buck
(13T 402804, 4433786), and Breeze Bridge (13T 398436, 4435431; Figures 2.1, 2.2, and 2.3).

14

�Adult Trout
Adult trout population sampling by CPW on the Colorado River from the Chimney Rock Ranch
to the Sheriff Ranch occurs from just upstream of the Hitching Post Bridge at CR 57 (13T 414569,
4440307) to just upstream of the Sheriff Ranch (13T 409216, 4438019; Figure 2.1). In 2016 and
2017, this section was broken into two sections, upstream and downstream of the diversion
structure on the Chimney Rock Ranch (13T 412203, 4439300), to evaluate effects of the diversion
structure on trout abundance and distribution.
Mottled Sculpin
Mottled Sculpin sampling sites (Figures 2.2 and 2.3) include the following locations: Windy Gap
connectivity channel (13T 416671, 4439947), Hitching Post (13T 414502, 4440208), Upper Red
Barn (13T 412749, 4439680), Sheriff Ranch (13T 409063, 4438050), Paul Gilbert SWA (13T
403398, 4434163), and Breeze Bridge SWA (13T 398436, 4435431). The number and location
of Mottled Sculpin sampling sites may be adjusted in response to river conditions and budget
constraints; additional sites could be surveyed opportunistically.
Table 2.1. Fish monitoring locations in the upper Colorado River and Fraser River, monitoring
period and monitoring frequency for trout fry, adult trout, and Mottled Sculpin.

Monitoring Target
Trout fry
population
estimates, genetic
composition, and
myxospore
enumeration
Monitoring Target

Adult trout
population
estimates

Site
Colorado River, Hitching Post
Colorado River, Upper Red Barn
Colorado River, Lower Red Barn
Colorado River, Sheriff Ranch
Colorado River, Paul Gilbert
Colorado River, Lone Buck
Colorado River, Breeze Bridge
Site
Colorado River, Chimney Rock Ranch to
Sheriff Ranch
Colorado River, ILVK reach
Colorado River, Parshall-Sunset reach on
Kemp-Breeze SWA
Colorado River,
Gilbert-Lone Buck SWA
Colorado River, Town of Granby property
(2 sites at Granby Trails)
Fraser River, 4-7 sites
15

Monitoring
Period

Monitoring
Frequency

June-October

Monthly

Monitoring
Period

Monitoring
Frequency

Spring

Annually

Spring

Annually

Fall

Annually

Spring

Biannually

Fall

Biannually

Fall

Some annually,
some biannually

�Monitoring Target

Site

Mottled Sculpin
population
estimates

Confluence of Fraser and Colorado rivers
Colorado River, Windy Gap connectivity
channel
Colorado River, Hitching Post
Colorado River, Chimney Rock Ranch
Colorado River, Pioneer Park SWA
Colorado River, Hot Sulphur SWA
Colorado River, Kemp Breeze SWA
Colorado River, Powers BLM

Monitoring
Period
Fall

Monitoring
Frequency
Annually

Fall

Annually

Fall
Fall
Fall
Fall
Fall
Fall

Annually
Annually
Annually
Annually
Annually
Annually

Figure 2.1. Map of the adult trout population sampling location in the Colorado River from the
upstream terminus at the CR 57 bridge on the Chimney Rock Ranch to the downstream terminus
on the Sheriff Ranch (from Fetherman et al. 2014).

16

�Figure 2.2. Map of the upper Colorado River trout fry, Mottled Sculpin and benthic
macroinvertebrate sampling sites downstream of Windy Gap Reservoir.

Figure 2.3. Map of the upper Colorado River trout fry, Mottled Sculpin and benthic
macroinvertebrate sampling sites downstream of Byers Canyon.
17

�2.4

Monitoring Schedule

Trout Fry
GR fry stocking by CPW will continue into July 2018. Fry stocking using the Hofer by Gunnison
River Rainbow (HxG) will occur from 2019-2021. CPW will complete fry population estimates,
genetic composition, and myxospore enumeration monthly, June-October, at all seven sites from
2018-2022.
Adult Trout
CPW will conduct annual adult trout population estimates in the spring in the Colorado River from
Chimney Rock Ranch to Sheriff Ranch and also within the ILVK reach from 2018-2022. The
Parshall-Sunset reach of the Colorado River on Kemp Breeze SWA will be sampled annually in
the fall, beginning in 2018 and continuing through 2022. Three additional sites on the Colorado
River (two on the Town of Granby property at Granby Trails) and one at the Gilbert-Lone Buck
SWA will be sampled biannually in the fall and spring, respectively, beginning in 2018 and
continuing through 2022. Multiple sites (four to seven) on the Fraser River will be sampled in the
fall on rotating annual and biannual schedules, beginning in 2018 and continuing through 2022.
Mottled Sculpin
CPW will conduct annual Mottled Sculpin sampling at eight sites along the Colorado River in the
fall, beginning in 2018 and continuing through 2022. Additional sites may be sampled
opportunistically depending on river conditions, and budgetary and personnel constraints.

Chapter 3: Benthic Macroinvertebrates
3.1

Introduction

Dams are known to drastically alter river habitat and have many diverse effects on aquatic
invertebrates (Ward and Stanford 1979). Those effects can be large and result in long term changes
in invertebrate communities (Vinson 2001). In the upper Colorado River basin, previous work
documented the dramatic change of the aquatic invertebrate community in the upper Colorado
River downstream of Windy Gap Reservoir since construction of the reservoir and that these
changes may be associated with flow alterations (Nehring et al. 2011). Nehring et al. (2011)
documented a 38% reduction in the diversity of aquatic invertebrates below Windy Gap Reservoir
between 1980-2011. Nineteen species of mayflies, four species of stoneflies, and eight species of
caddisflies have been extirpated from the sampling sites since 1982. In addition to the changes
temporal changes in the invertebrate community, there was a spatial pattern of increasing diversity
downstream of Windy Gap Reservoir that indicated current effects of the reservoir on invertebrate
18

�habitat and communities. Sensitive species including Drunella grandis, Pteronarcella badia, and
Pteronarcys californica were reduced or eliminated from sites close to Windy Gap Reservoir and
replaced by tolerant species including Ephemerella sp, Baetis sp, and Hydropsyche sp.
The salmonfly (P. californica) is a large aquatic invertebrate that can reach high densities in some
Colorado rivers. These invertebrates play an important ecological role as grazers in stream systems
and have been documented as extremely important to stream dwelling trout as a food resource.
Nehring (1987) reported in a diet study of trout in the Colorado River that P. californica was the
most common food item, comprising 64-75% of the mean stomach content over the four year
study. Because of their high biomass and hatching behavior, salmonflies also play an important
role in supplementing terrestrial food webs and riparian communities with stream-derived nutrients
(Baxter et al. 2005; Walters et al. 2018).
While ecologically important and found in high abundance at some sites, the salmonfly has
relatively specific environmental requirements and is considered intolerant of disturbance in
bioassessment protocols (Erickson 1983; Fore et al. 1996; Barbour et al. 1999). Salmonflies are
sensitive to habitat alterations in part because of their lifespan; they are one of the longest lived
aquatic insects in the Nearctic (DeWalt and Stewart 1995). Previous work indicates that the range
and density of P. californica have declined in the Colorado River, and that these declines may be
associated with flow alterations (Nehring et al. 2011). Once common in the upper Colorado River
(USFWS 1951; Dames and Moore 1977; Erickson 1983), the abundance of salmonflies has
declined, especially downstream of Windy Gap Reservoir where flow alterations associated with
trans-mountain water diversions are greatest (Nehring et al. 2011). This pattern has been observed
in other rivers. Richards (2000) documented six to eight times lower density of salmonflies
downstream of a reservoir compared to upstream and found a negative correlation between their
density and substrate embeddedness.
Aquatic invertebrate diversity, as well as salmonfly abundance and distribution, have been reduced
in the Colorado River downstream of Windy Gap Reservoir. Habitat alterations associated with
the project seem to have reduced benthic aquatic habitat quality in the river. Restoring connectivity
in the upper Colorado River and addressing habitat limitations associated with the flow and
sediment regimes should improve conditions for, and the diversity of, invertebrates in the Colorado
River. The objective of this project is to document the distribution and density of P. californica
in the upper Colorado River and investigate changes over time after the construction of the Windy
Gap connectivity channel.

19

�●
●

●
●
3.2

Goals
Increase the abundance and diversity of benthic macroinvertebrates in the upper Colorado
River and re-establish historically common species
Increase the distribution and abundance of salmonflies downstream of Windy Gap
Reservoir
Objectives
Evaluate how the Windy Gap connectivity channel affects the aquatic invertebrates of the
upper Colorado River
Evaluate the distribution and density of salmonflies in the upper Colorado River
Methods

Replicate macroinvertebrate samples (n = 5) will be collected by CPW at each site using a 0.92 ft2
Hess sampler with a 350-µm mesh net. The replicate samples will be collected from the same
riffle with predominantly cobble substrate by disturbing the streambed to a depth of approximately
3.9 inches. Field samples will be washed through a 350-µm sieve and organisms preserved in 80%
ethanol. Velocity and depth will be taken at each Hess sample site to ensure samples were taken
from similar riffle habitat. Macroinvertebrate samples will be sorted and sub-sampled in the
laboratory using a standard USGS 300-count protocol, except that replicates will not be
composited and each one will undergo sub sampling and identification protocols (Moulton et al.
2000). All organisms, except for chironomids and non-insects, will be identified to genus or
species. Chironomids will be identified to subfamily and non-insects (e.g., oligochaetes,
amphipods) identified to class.
Salmonfly emergence and density will be monitored by CPW during the emergence period of late
June through early July. Multiple pass removal estimates will be completed by searching 98.6 foot
sections of stream bank for P. californica exuvia adjacent to riffle habitat. If possible, each site
will be visited multiple times to encompass the entire emergence. If a site is visited only once,
this will occur as soon as possible after the emergence is complete. Three to seven people will
intensively search the riparian area from 3.3 to 65.6 feet from the water’s edge depending on exuvia
distribution. On a single sampling occasion, each area will be searched two to four times with
identical search areas, effort, and personnel. A multiple-pass depletion model will be used to
estimate the total density of exuvia at each site (Zippin 1956).
3.3

Study Sites

Aquatic invertebrate sampling sites (Figures 2.2 and 2.3) include the following locations: Windy
Gap connectivity channel (13T 416671, 4439947), Hitching Post (13T 414502, 4440208), Upper
Red Barn (13T 412749, 4439680), Sheriff Ranch (13T 409063, 4438050), Paul Gilbert #2 (13T
403398, 4434163), and Breeze Bridge #2 (13T 398341, 4435450). Exuvia sampling sites include
the following locations: Windy Gap connectivity channel (13T 416671, 4439947), Hitching Post
(13T 414502, 4440208), Sheriff Ranch (13T 409063, 4438050), Paul Gilbert SWA (13T 403398,
20

�4434163), and Breeze Bridge SWA (13T 398436, 4435431). The number and location of
invertebrate sampling sites may be adjusted in response to river conditions and budget constraints;
additional sites could be surveyed opportunistically.
3.4

Monitoring Schedule

Benthic macroinvertebrate samples will be collected in August-September of each year, and
salmonfly density and emergence will be monitored in June-July of each year. Sampling will
begin in 2018 and continue through 2022. If construction of the Windy Gap connectivity channel
is delayed, the proposed monitoring schedule will be adjusted accordingly.

Chapter 4: Hydrology, Hydraulics, and Geomorphology
4.1

Introduction

Aquatic habitat in the upper Colorado River has been altered by changes in the flow regime, water
depletions, sedimentation, and armoring of the stream bed (Municipal Subdistrict 2011). Flow
alteration has impacted a myriad of processes related to hydrologic, hydraulic, geomorphic,
physicochemical, and biological stream functions (Harman et al. 2012). The reduced capacity to
mobilize sediment has embedded riffle habitats important for benthic organisms, such as the
salmonfly and Mottled Sculpin. Reconnecting flows from the Colorado River through the Windy
Gap connectivity channel should improve sediment transport supply to reaches downstream of
Windy Gap Reservoir. However, reconnecting reaches upstream and downstream of Windy Gap
Reservoir may not be sufficient to restore benthic organisms without improving habitat conditions.
Habitat restoration will reduce channel dimensions to improve the frequency of sediment transport
and floodplain activation under current and future flow regimes. Riffle de-armoring techniques
will also be applied to improve habitat for benthic macroinvertebrates and fish in reaches
associated with the Kemp Breeze Habitat Restoration Project.
This chapter presents a monitoring approach for CPW to evaluate the effectiveness of the Kemp
Breeze Habitat Restoration Project on instream hydraulics and geomorphology. Hydrology will
be evaluated by investigating the channel-forming discharge, flood frequency, and flow duration
for historical, current, and future flow regimes. Analysis of hydrology will also inform design of
the Kemp Breeze Habitat Restoration Project. Assessment of floodplain connectivity and flow
dynamics will be used to evaluate changes in hydraulics. Geomorphology monitoring will include
evaluation of sediment transport, lateral stability, riparian vegetation, bedform diversity, and bed
material characterization. The Kemp Breeze Habitat Restoration Project may utilize experimental
riffle de-armoring treatments, which will be monitored to inform the application of those
treatments in other project areas.

21

�Monitoring and evaluation of hydrology, hydraulics, and geomorphology for the Windy Gap
connectivity channel is beyond the scope of this monitoring plan, but should be conducted if
additional funding becomes available or if monitoring priorities are changed during discussions
with project stakeholders. An approach for geomorphic assessment including evaluation of
channel hydraulics and sediment transport is presented in Appendix A to support project
development and fundraising. Although CPW cannot commit to geomorphic assessments for the
connectivity channel without additional resources, CPW will continue to provide technical design
assistance and review for the Windy Gap Connectivity Channel Project.
Goals
● Improve sediment transport processes in upper Colorado River reaches associated with the
Kemp Breeze Habitat Restoration Project
● Improve floodplain connectivity in upper Colorado River reaches associated with the
Kemp Breeze Habitat Restoration Project
● Restore and enhance riparian corridors of the upper Colorado River to improve wildlife
habitat and increase flood resilience
Objectives
● Increase sediment transport capacity and competence by manipulating channel dimensions
● Decrease the prevalence of fine sediment in embedded riffle habitats
● Increase the suitability of aquatic habitat for Rainbow Trout, Brown Trout, and Mottled
Sculpin by improving instream hydraulics
● Increase the abundance of native riparian vegetation along streambanks and floodplains
● Increase the frequency of floodplain inundation under current and future flow regimes
4.2

Methods

Hydrology
Average daily and annual peak discharge records will be obtained by CPW for existing stream
gauges and used to evaluate flow alteration and inform restoration designs. Design flows will be
selected from hydrologic analysis to support hydraulic modeling. If the location or availability of
streamflow data from existing stream gauges is insufficient to support assessment and design,
streamflow measurement may be conducted to support evaluation of project reaches. As there are
a number of stream gauges located throughout the upper Colorado River basin, collecting
additional streamflow measurements may not be needed.
Hydraulics
Floodplain connectivity and flow dynamics will be evaluated by CPW to support monitoring of
hydraulics within the Habitat Restoration Project reach on the Kemp Breeze SWA. Topographic
surveys will be conducted with survey-grade GPS and an Acoustic Doppler Current Profiler
(ADCP) to support hydraulic modeling and project design. Hydraulic models will be configured
22

�and calibrated for existing, proposed, and as-built conditions. The Hydrologic Engineering Center
- River Analysis System (HEC-RAS) model will be used for assessment, design, and evaluation
of the Kemp Breeze Habitat Restoration Project. Stream velocity, water depth, shear stress, and
stream power will be derived from model outputs to investigate channel hydraulics and habitat
suitability. HEC-RAS models will also be used to investigate changes in floodplain connectivity
by determining the extent and frequency of floodplain inundation across a range of flows. Channel
narrowing activities associated with the Kemp Breeze Habitat Restoration Project should increase
the frequency of floodplain inundation under current and future flow regimes.
Geomorphology
Geomorphology monitoring will include evaluation by CPW of sediment transport, lateral
stability, riparian vegetation, bedform diversity, embeddedness, and bed material characterization.
Sediment transport will be evaluated within the Kemp Breeze SWA by monitoring the movement
of 200-600 Passive Integrated Transponder (PIT) tagged rocks. Using PIT tags to monitor
sediment transport is an ideal technique for tracking movements of individual sediment particles
in gravel-bed rivers (Lamarre et al. 2005). Pebble counts will be conducted to characterize bed
material and select sediment sizes for tagging. PIT-tagged rocks will be placed in riffles, and
movement of rocks will be monitored using stationary and mobile radio frequency identification
(RFID) antennas. Mobile antennas will be used to relocate PIT-tagged rocks and locations will be
recorded with survey-grade GPS. Stationary antennas will be placed downstream of select areas
where PIT-tagged rocks have been deployed to monitor the sediment transport across a range of
flows. Measurements of sediment movement will be combined with results from hydraulic
modeling to evaluate changes in sediment transport associated with implementation of the Kemp
Breeze Habitat Restoration Project. Sediment monitoring will target locations where riffle dearmoring occurs to evaluate the effectiveness of these experimental treatments, including
assessments of fine sediment and embeddedness.
Channel morphology will be monitored with repeat surveys of the longitudinal profile and
monumented cross-section locations. Geomorphic surveys will be conducted with survey-grade
GPS and an ADCP. Additional survey points will be collected throughout the active stream
channel, on islands, and along the floodplain. Lateral stability and bedform diversity will be
evaluated for existing, proposed, and as-built conditions within the Kemp Breeze SWA using data
derived from longitudinal profile and cross-section surveys. The greenline stability rating
(Winward 2000; ACOE 2017) will be used to evaluate changes in riparian vegetation and lateral
stability for the Kemp Breeze Habitat Restoration Project. Identification of riparian vegetation will
be limited to the following classes: anchored rock/logs, trees (coniferous and deciduous), willows,
other shrubs (sagebrush, cinquefoil, etc.), wet sedges and rushes, other sedges, wet grasses, other
grasses, and unvegetated areas (sandbars, loose rock, and bare soil). As CPW aquatic biologists
and researchers would not typically be tasked with vegetation identification, additional funding
could be used to support a more rigorous approach to riparian vegetation monitoring.

23

�4.3

Study Sites

The primary study area is the Habitat Restoration Project at the Kemp Breeze SWA. Additional
study areas could include sites associated with the future, larger scale Habitat Project, such as
Pioneer Park SWA, Hot Sulphur Springs SWA, and Chimney Rock Ranch. Reference reaches on
the Fraser River could also be monitored to compare habitat conditions in project areas to relatively
undisturbed reference areas, but reference-reach surveys on the Fraser River are currently beyond
the scope of this monitoring plan.
4.4

Monitoring Schedule

The proposed schedule for the Kemp Breeze Habitat Restoration Project is presented in Table 4.1.
Baseline monitoring will be conducted in 2018 and 2019, project construction is scheduled for
2020, implementation monitoring will occur in 2021, and effectiveness monitoring will take place
in 2022. The schedule assumes that the Kemp Breeze Habitat Restoration Project will be
constructed in 2020, but the construction schedule is dependent upon various factors and is subject
to change. If construction of the project is delayed, the proposed monitoring schedule will be
adjusted accordingly.
Table 4.1. Proposed schedule for the Kemp Breeze SWA Habitat Restoration Project.
Task

2018

Project Design

2019

2020

2021

2022

X

X

X

Project Construction

X

Geomorphic Surveys and Assessment

X

Hydraulic Modeling

X

X

X

Sediment Transport Surveys and Assessment

X

X

X

Riparian Vegetation Surveys and Assessment

X

X

X

Chapter 5: Monitoring Budget
CPW has committed within the Headwaters Project to a budget of $115,000 annually beginning in
2017, across five years of effort for a total of $575,000. Securing additional funds could be
beneficial for more extensive evaluation of the Headwaters Project, i.e. proposed hydrology,
hydraulics, and geomorphology assessments specifically related to the Windy Gap Connectivity
Channel Project (Appendix A), and the proposed fish movement study (Appendix B).
24

�References
ACOE (U.S. Army Corps of Engineers). 2017. Wyoming Stream Quantification Tool (WSQT)
User Manual and Spreadsheet. Beta Version.
Avila, B. W., D. L. Winkelman, and E. R. Fetherman. In review. Survival of whirling disease
resistant Rainbow Trout fry: a comparison of two strains. Submitted to the Journal of Aquatic
Animal Health.
Baxter, C. V., K. D. Fausch, and W.C. Saunders. 2005. Tangled webs: reciprocal flows of
invertebrate prey link streams and riparian zones. Freshwater Biology 50:201-220.
Dames and Moore. 1977. Environmental assessment report Windy Gap project Grand County,
Colorado for municipal subdistrict, Northern Water Conservancy District. Denver, CO.
DeWalt, R. E., and K. W. Stewart. 1995. Life-histories of stoneflies (Plecoptera) in the Rio
Conejos of southern Colorado. Great Basin Naturalist 55:1-18.
Fetherman, E. R., D. L. Winkelman, M. R. Baerwald, and G. J. Schisler. 2014. Survival and
reproduction of Myxobolus cerebralis resistant rainbow trout in the Colorado River and
increased survival of age-0 progeny. PLoS ONE 9(5):e96954.
Fetherman, E. R., D. L. Winkelman, G. J. Schisler, and M. F. Antolin. 2012. Genetic basis of
differences in myxospore count between whirling disease-resistant and -susceptible strains of
Rainbow Trout. Diseases of Aquatic Organisms 102:97-106.
Fetherman, E. R., D. L. Winkelman, G. J. Schisler, and C. A. Myrick. 2011. The effects of
Myxobolus cerebralis on the physiological performance of whirling disease resistant and
susceptible strains of Rainbow Trout. Journal of Aquatic Animal Health 23:169-177.
Fore, L. S., J. R. Karr, and R. W. Wisseman. 1996. Assessing invertebrate responses to human
activities: Evaluating alternative approaches. Journal of the North American Benthological
Society 15(2):212-231.
Harman, W., R. Starr, M. Carter, K. Tweedy, K. Suggs, and C. Miller. 2012. A function-based
framework for stream assessment and restoration projects. US Environmental Protection
Agency, Office of Wetlands, Oceans, and Watersheds, Washington, DC. EPA 843-K-12-006.
Kowalski, D. A. 2014. Colorado River aquatic resources investigations. Federal Aid Project F237-R21. Federal Aid in Fish and Wildlife Restoration, Job Progress Report. Colorado Parks
and Wildlife, Aquatic Wildlife Research Section. Fort Collins, Colorado.
Lamarre, H., B. MacVicar, and A. G. Roy. 2005. Using passive integrated transponder (PIT) tags
to investigate sediment transport in gravel-bed rivers. Journal of Sedimentary Research
75:736-741.
Municipal Subdistrict. 2011. Windy Gap Firming Project Fish and Wildlife Enhancement Plan.
Prepared by the Municipal Subdistrict, Northern Colorado Water Conservancy District in
partnership with Denver Water. 15 pp.
Nehring, R. B. 1987. Stream fisheries investigations. Federal Aid Project F-51-R. Federal Aid in
Fish and Wildlife Restoration, Job Progress Report. Colorado Division of Wildlife, Fish
Research Section. Fort Collins, Colorado.
25

�Nehring, R. B. 2006. Colorado’s cold water fisheries: whirling disease case histories and insights
for risk management. Colorado Division of Wildlife Aquatic Wildlife Research Special Report
Number 79. Denver, Colorado. 46 pp.
Nehring, R. B., B. Heinhold, and J. Pomeranz. 2011. Colorado River aquatic resources
investigations. Federal Aid Project F-237-R18. Federal Aid in Fish and Wildlife Restoration,
Final Report. Colorado Division of Wildlife, Aquatic Wildlife Research Section. Fort Collins,
Colorado.
Nehring, R. B., and K. G. Thompson. 2001. Impact assessment of some physical and biological
factors in the whirling disease epizootic among wild trout in Colorado. Colorado Division of
Wildlife Aquatic Research Special Report Number 76. Denver, Colorado. 78 pp.
Richards, D. C., M. G. Rolston, and F. V. Dunkle. 2000. A comparison of salmonfly density
upstream and downstream of Ennis Reservoir. Intermountain Journal of Sciences 6(1):1-9.
Ryce, E. K. N., A. V. Zale, E. MacConnell, and M. Nelson. 2005. Effects of fish age versus size
on the development of whirling disease in Rainbow Trout. Diseases of Aquatic Organisms
63:69-76.
Schisler, G. J., E. P. Bergersen, and P. G. Walker. 1999a. Evaluation of chronic gas supersaturation
on growth, morbidity, and mortality of fingerling Rainbow Trout infected with Myxobolus
cerebralis. North American Journal of Aquaculture 61:175-183.
Schisler, G. J., E. P. Bergersen, and P. G. Walker. 2000. Effects of multiple stressors on morbidity
and mortality of fingerling Rainbow Trout infected with Myxobolus cerebralis. Transactions
of the American Fisheries Society 129:859-865.
Schisler, G. J., K. A. Myklebust, and R. P. Hedrick. 2006. Inheritance of Myxobolus cerebralis
resistance among F1-generation crosses of whirling disease resistant and susceptible Rainbow
Trout strains. Journal of Aquatic Animal Health 18:109-115.
Schisler, G. J., P. G. Walker, L. A. Chittum, and E. P. Bergersen. 1999b. Gill ectoparasites of
juvenile Rainbow Trout and Brown Trout in the upper Colorado River. Journal of Aquatic
Animal Health 11:170-174.
Scott, W. B., and E. J. Crossman. 1973. Freshwater fishes of Canada. Bulletin of the Fisheries
Research Board of Canada Number 184.
Sigler, F. F., and R. R. Miller. 1963. Fishes of Utah. Utah Department of Fish and Game. Salt Lake
City, Utah.
Tetra Tech. 2015. Final Report Windy Gap Reservoir Modification Study. Fort Collins, Colorado.
USFWS (U.S. Fish and Wildlife Service). 1951. Recreational use and water requirements of the
Colorado River fishery below Granby Dam in relation to the Colorado-Big Thompson
diversion project. U.S. Fish and Wildlife Service, Region 2. Albuquerque, New Mexico.
Vinson, M. R. 2001. Long-term dynamics of an invertebrate assemblage downstream from a large
dam. Ecological Applications 11:711-720.
Walters, D. M., J. S. Wesner, R. E. Zuellig, D. A. Kowalski, and M. C. Kondratieff. 2018. Holy
flux: spatial and temporal variation in massive pulses of emerging insect biomass from western
U.S. rivers. Ecology 99(1):238-240.

26

�Ward, J. V. 1998. Riverine landscapes: biodiversity patterns, disturbance regimes, and aquatic
conservation. Biological Conservation 83:269-278.
Ward, J. V., and J. A. Stanford. 1979. The ecology of regulated streams. Plenum Press, New York.
Woodling, J. 1985. Colorado’s little fish, a guide to the minnows and other lesser known fishes in
the state of Colorado. Colorado Division of Wildlife. Denver, Colorado.
Walker, P. G., and R. B. Nehring. 1995. An investigation to determine the cause(s) of the
disappearance of young wild rainbow trout in the Upper Colorado River, in Middle Park,
Colorado. Colorado Division of Wildlife Report. Denver, Colorado. 134 pp.
Windward, A. H. 2000. Monitoring the vegetation resources in riparian areas. Gen. Tech. Rep.
RMRS-GTR-47. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky
Mountain Research Station. 49 p.

27

�Appendix A: Proposed Geomorphic Monitoring of the Connectivity Channel
The following proposal describes a geomorphic monitoring study that could be conducted in
conjunction with the construction and evaluation of the Windy Gap connectivity channel in the
Colorado River, if additional funding and staff time are added for these evaluations.
A.1

Introduction

Aquatic habitat in the upper Colorado River has been altered by changes in the flow regime, water
depletions, sedimentation, and armoring of the stream bed (Municipal Subdistrict 2011).
Reconnecting flows from the Colorado River through the Windy Gap connectivity channel should
improve sediment transport processes for reaches downstream of Windy Gap Reservoir. The
connectivity channel should also support fish passage for target species and life stages across a
range of flows and operational scenarios. This document presents a monitoring approach to
evaluate the effectiveness of the Windy Gap connectivity channel. Hydrology should be
monitored to support evaluation of fish passage and channel evolution. Flow monitoring within
the connectivity channel may also be necessary for water rights administration. Geomorphology
monitoring will include evaluation of sediment transport, lateral stability, riparian vegetation,
bedform diversity, and bed material characterization. Theses monitoring targets were selected to
support evaluation of project goals and objectives.
Goals
● Restore sediment transport in the upper Colorado River around Windy Gap Reservoir
● Restore fish passage in the upper Colorado River around Windy Gap Reservoir
Objectives
● Transport sediment loads from the Colorado River through the Windy Gap diversion
structure and connectivity channel
● Convey flood flows through the connectivity channel and adjacent floodplain
● Validate that hydraulics within the connectivity channel are within the range of target fish
passage criteria
● Evaluate evolution of the connectivity channel to ensure it maintains a target range of
dimensions that support fish passage
● Improve the resilience and stability of streambanks along the connectivity channel by
increasing the abundance of native riparian vegetation
A.2

Methods

Hydrology
Flows through the connectivity channel should be monitored to support evaluation of the Windy
Gap Connectivity Channel Project. Discharge records will be obtained for existing stream gauges
and diversion structures to evaluate flows within the connectivity channel. If the location or
28

�availability of streamflow data from existing stream gauges and diversions is insufficient to
support assessment and design, streamflow measurements may be conducted to support project
evaluation. The establishment of any streamflow stations should be coordinated with Northern
Water to ensure that monitoring is adequate to support water rights administration and project
evaluation.
Hydraulics
Floodplain connectivity and flows dynamics will be evaluated to support monitoring of hydraulics
within the connectivity channel. Topographic surveys will be conducted with survey-grade GPS
and an Acoustic Doppler Current Profiler (ADCP) to support hydraulic modeling and project
evaluation. Hydraulic models will be configured and calibrated for as-built (implementation
monitoring) and post-runoff (effectiveness monitoring) conditions with the Hydrologic
Engineering Center - River Analysis System (HEC-RAS). Stream velocity and water depth will
be derived from model outputs to compare channel hydraulics and fish passage criteria. HECRAS models will also be used to investigate sediment transport and floodplain connectivity.
Geomorphology
Geomorphology monitoring will include evaluation of sediment transport, channel evolution,
riparian vegetation, bedform diversity, and bed material characterization. Sediment transport will
be evaluated within the connectivity channel by monitoring the movement of 200-600 Passive
Integrated Transponder (PIT) tagged rocks. Using PIT tags to monitor sediment transport is an
ideal technique for tracking movements of individual sediment particles in gravel-bed rivers
(Lamarre et al. 2005). Pebble counts will be conducted to characterize bed material and select
sediment sizes for tagging. PIT-tagged rocks will be placed in riffles, and movement of rocks will
be monitored using stationary and mobile radio frequency identification (RFID) antennas (see
Appendix A). Mobile antennas will be used to relocate PIT-tagged rocks, and locations will be
recorded with survey-grade GPS. Stationary antennas will be placed downstream of select areas
where PIT-tagged rocks have been deployed to monitor sediment transport across a range of flows.
Connectivity channel morphology will be monitored with repeat surveys of the longitudinal profile
and cross-section locations. Geomorphic surveys will be conducted with survey-grade GPS and
an ADCP. Lateral stability and bedform diversity will be evaluated for as-built and post-runoff
conditions using data derived from longitudinal profile and cross-section surveys. The greenline
stability rating (Winward 2000; ACOE 2017) will be used to evaluate changes in riparian
vegetation and lateral stability. Identification of riparian vegetation will be limited to the following
classes: anchored rock/logs, trees (coniferous and deciduous), willows, other shrubs (sagebrush,
cinquefoil, etc.), wet sedges and rushes, other sedges, wet grasses, other grasses, and unvegetated
areas (sandbars, loose rock, and bare soil). Additional funding could support a more rigorous
approach to riparian vegetation monitoring if desired.

29

�A.3

Study Sites

The primary study area is the Windy Gap connectivity channel associated with the Windy Gap
Connectivity Channel Project. Topographic surveys will include the connectivity channel and
adjacent floodplain from the upstream diversion point to the downstream confluence with the
Colorado River.
A.4

Monitoring Schedule

Baseline monitoring will not be conducted prior to construction of the Windy Gap connectivity
channel, but survey data collected during project design should be sufficient to document changes
associated with project implementation. Implementation and effectiveness monitoring for the
connectivity channel would occur in 2021 and 2022, respectively. The schedule assumes that the
Windy Gap connectivity channel will be constructed in 2020, but the construction schedule is
dependent upon various factors and is subject to change. If construction of the project is delayed,
the proposed monitoring schedule will be adjusted accordingly.
Table A.1. Proposed geomorphic monitoring schedule for the Windy Gap Connectivity Channel
Project.
Task

2021

2022

Geomorphic Surveys and Assessment

X

X

Hydraulic Modeling

X

X

Sediment Transport Surveys and Assessment

X

X

Riparian Vegetation Surveys and Assessment

X

X

Project Design
Project Construction

A.5

2018

2019

X

X

2020

X

References

ACOE (U.S. Army Corps of Engineers). 2017. Wyoming Stream Quantification Tool (WSQT)
User Manual and Spreadsheet. Beta Version.
Lamarre, H., B. MacVicar, and A. G. Roy. 2005. Using passive integrated transponder (PIT) tags
to investigate sediment transport in gravel-bed rivers. Journal of Sedimentary Research
75:736-741.

30

�Municipal Subdistrict. 2011. Windy Gap Firming Project Fish and Wildlife Enhancement Plan.
Prepared by the Municipal Subdistrict, Northern Colorado Water Conservancy District in
partnership with Denver Water. 15 pp.
Windward, A. H. 2000. Monitoring the vegetation resources in riparian areas. Gen. Tech. Rep.
RMRS-GTR-47. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky
Mountain Research Station. 49 p.

31

�Appendix B: Proposed Fish Movement Study
The following draft proposal describes a fish movement study that could be conducted in
conjunction with the construction and evaluation of the Windy Gap connectivity channel in the
Colorado River, if additional funding and staff time are added for these evaluations. The proposal
is still in draft form and is subject to change.
B.1

Introduction

Connectivity is a fundamental element of landscape structure and ecological processes, and
longitudinal connectivity is especially important in rivers (Taylor et al. 1993; Fausch et al. 2002).
Loss of free passage due to artificial barriers can lead to habitat fragmentation and limit fish
distributions, such as those of Mottled Sculpin (Cottus bairdii) in the Colorado River, by reducing
access to key habitats (Fausch et al. 2002; Lucas et al. 2009). In general, fish require three major
habitat types: 1) feeding habitats with favorable growth conditions, 2) refugia from harsh
environmental conditions with unfavorable growth conditions, and 3) spawning habitat with
necessary flow conditions for egg incubation. Movements among these various habitat types
occurs continuously throughout the year based on spawn timing (e.g., spring for Rainbow Trout
[Oncorhynchus mykiss] and fall for Brown Trout [Salmo trutta]), environmental and flow
conditions, physical or habitat conditions and availability, and growth or life stage (Schlosser and
Angermeier 1995). The construction of the connectivity channel around Windy Gap Reservoir
will help restore connectivity for these types of movements, and provide access to favorable
habitats, such as upstream spawning locations, that had been previously unavailable for fish
populations in the Colorado River downstream of Windy Gap Reservoir. Conversely, fish such as
Mottled Sculpin, which are currently absent immediately downstream of Windy Gap Reservoir,
will be able to distribute downstream increasing the diversity of the riverine ecosystem
downstream of the reservoir. This study will be focused on evaluating fish movement rates through
the connectivity channel and validating that the connectivity channel is being used for fish passage
in both the upstream and downstream directions.
Passive integrated transponder (PIT) tags are an important tool for evaluating fish growth,
movement, and mortality due to their relatively low cost, longevity, ability to identify unique
individuals, ease of application, and minimal effects on fish survival, growth, feeding behavior,
and swimming performance (Zydlewski et al. 2006; Newby et al. 2007; Ficke et al 2012).
Antennas constructed of copper wire anchored to the bottom of the river (stationary antenna) or
actively passing over the fish (portable antennas) are used to detect PIT tags that have been inserted
internally in target fish species. The antennas create an electromagnetic field that activates the tag
and records the unique identification number returned from the tag to an interrogation system or
reader. Since the tags are activated by an electromagnetic field rather than battery, they are not
only considered passive, but also have an infinite life. In recent years, Colorado Parks and Wildlife
(CPW) has utilized PIT technology to monitor movement of salmonids in and out of specific
management sections (Fetherman et al. 2014; Fetherman et al. 2015), passage of salmonids,
32

�suckers, and dace at whitewater park structures (Fox et al. 2016), and evaluate Brown Trout habitat
utilization in mountain streams (Richer et al. 2017). The focus species for this study include
Rainbow Trout and Brown Trout, the dominant sport fish species in the Fraser River and Colorado
River upstream and downstream of Windy Gap Reservoir. Mottled Sculpin distribution in the
downstream direction through the connectivity channel will also be an important focus of the fish
movement monitoring efforts. Although most Mottled Sculpin are relatively sedentary, recent
research has shown that a small percentage of these fish exhibit considerable movement capability
(Breen et al. 2009; Hudy and Shiflet 2009).
Using PIT tags and stationary and portable PIT tag antennas, the overall objectives of this study
are to evaluate Rainbow Trout, Brown Trout, and Mottled Sculpin movement both upstream and
downstream through the connectivity channel, adequacy of attraction flows from the connectivity
channel, and large-scale movement patterns of various age classes of target fish species throughout
the upper Colorado River.

●
●
●
●
●
●
●
●
●
B.2

Goals
Restore longitudinal connectivity for fish populations around Windy Gap Reservoir
Primary Objectives
Evaluate fish movement patterns under existing conditions upstream and downstream of
Windy Gap Reservoir
Validate that the connectivity channel is being utilized for fish passage by Brown Trout,
Rainbow Trout, and Mottled Sculpin
Secondary Objectives
Determine if the stream gage on the Fraser River upstream of Windy Gap Reservoir is an
obstacle for fish movement
Determine if the diversion structure on the Fraser River below Highway 40 is an obstacle
for fish movement
Determine if the diversion structure on the Chimney Rock Ranch is an obstacle for fish
movement
Evaluate the relative proportion of the fish population that is utilizing the Colorado and
Fraser rivers upstream of Windy Gap Reservoir
Evaluate fish entrainment into Windy Gap Reservoir following construction of the Windy
Gap connectivity channel diversion structure
Evaluate utilization of spawning habitat in the headwaters of the Fraser River
Methods

The study will include two distinct phases. First, a baseline study of fish movement patterns and
rates will be conducted for reaches upstream and downstream of Windy Gap Reservoir. Second,
the baseline study design will be expanded to evaluate the efficiency of fish movement through
the Windy Gap connectivity channel. To achieve the primary study objective of evaluating fish
movement patterns under existing conditions, stationary antennas will be installed in the Colorado
River on the Chimney Rock Ranch upstream of the Hitching Post Bridge and upstream of Windy
33

�Gap Reservoir downstream of the confluence with the Fraser River in 2019 to monitor baseline
movement rates. An additional stationary antenna will be installed on the Chimney Rock Ranch
just upstream of the confluence with Drowsy Water Creek to determine distances moved from
downstream fish tagging locations. All three antenna locations will consist of paired antenna loops
constructed from 8-gauge copper speaker wire and anchored to the substrate to prevent movement
or change in shape, maximizing read range. Paired antenna loops allow researchers to determine
directionality of movement (upstream versus downstream) when it occurs at each antenna location.
Antenna stations will be powered by multiple 12-V, 120-Ah marine deep cycle batteries housed in
a job box along with the reader(s). Additional power for each antenna station will be supplied by
solar panels installed near each antenna location. Readers will run continuously after installation
to capture fish movements during all times of the day and throughout the entire time that they are
installed. Baseline fish movement rates will be monitored in 2019 and 2020, prior to the
construction of the Windy Gap connectivity channel.
Following the completion of the Windy Gap connectivity channel, additional antenna stations will
be installed upstream and downstream of the connectivity channel to achieve the primary study
objective of validating that the connectivity channel is being used by Brown Trout, Rainbow Trout,
and Mottled Sculpin. The downstream antenna will be installed in the connectivity channel just
upstream of where the channel reconnects with the Colorado River downstream of Windy Gap
Reservoir. The upstream antenna will be incorporated into the diversion structure located at the
upstream end of the connectivity channel. Similar to the antenna design described above, each
antenna location will have paired antenna loops to determine directionality of movement into or
out of the connectivity channel. Additionally, successful movement in either direction through the
connectivity channel will be evaluated during the data analysis phase of the project by quantifying
the proportion of fish that were detected at both antenna locations within the connectivity channel.
A third antenna station may be installed on the Schmuck’s bypass channel originating from Windy
Gap Reservoir, dependent upon the amount of flow in this channel, to determine if flows attract
fish and prevent them from finding or using the connectivity channel during certain times of the
year. Efficiency of the connectivity channel will be monitored using these three antenna stations
in 2021 and 2022.
Portable antennas, antennas mounted in or on rafts and floated on the surface of the river, will be
deployed during both the baseline and efficiency monitoring phases of the study (2019-2022) to
improve detection probability of tagged fish and determine the fate of fish that are never detected
at a stationary antenna site. GPS technology will be incorporated into portable antenna designs to
get more precise locations on fish throughout the study reach and to estimate average distance
moved by fish species and size. Portable antennas will be deployed in three reaches: 1) on the
Colorado River between Hitching Post Bridge and the Sheriff Ranch; 2) on the Colorado River
upstream of Windy Gap Reservoir and the confluence with the Fraser River; and 3) on the Fraser
River between the Highway 40 diversion structure and Fraser Canyon (Figure A.1). An additional
portable antenna reach will be added within the connectivity channel following its completion,
which will be monitored during the effectiveness monitoring phase of the study (2021-2022) to
determine location and fate of fish that entered but did not exit the channel. During deployment,
34

�portable antennas will be run down the thalweg, as well as the left and right sides of the river
channel, to increase detection probabilities of fish throughout the channel. The combination of the
three runs will constitute a single detection or “mark” pass in any given reach. A second set of
three runs (thalweg, left and right sides of the river) will be used as a “recapture” pass to estimate
detection probability, tagged fish abundance, and change in detection location in the event that a
fish moved between passes. To maximize the chance of capturing fish movements during portable
antenna surveys, surveys will be conducted in the spring (March-April) and fall (SeptemberOctober) during times when fish are most likely to be moving to spawning sites.
Secondary objectives, specifically determining if the gauge on the Fraser River upstream of Windy
Gap Reservoir, the diversion structure on the Fraser River downstream of Highway 40, and the
diversion structure on the Chimney Rock Ranch are obstacles for fish movement, will be achieved
using additional antenna stations at these locations. At each location, a set of paired antennas will
be installed upstream and downstream of each structure allowing for determination of
directionality of movement as fish approach the structure. Antennas located downstream of each
structure will provide information regarding the number of fish that approached and attempted to
pass the structure, whereas antennas located upstream of the structures will provide information
regarding the number of fish that successfully passed the structure. Each of the three locations
will be monitored for at least one season during the baseline monitoring phase of the study (20192020). The secondary objective of determining the relative proportion of the fish population that
are utilizing the Colorado and Fraser rivers upstream of Windy Gap Reservoir will be achieved by
installing a paired antenna on the Colorado River upstream of the confluence with the Fraser River
to estimate fish movement rates among the two rivers. This antenna will be installed during the
baseline monitoring phase and remain intact following construction of the Windy Gap connectivity
channel (2019-2022). Additionally, an antenna station will be installed downstream of the
dynamic weir in the inlet to Windy Gap Reservoir to achieve the secondary objective of
determining if fish are entrained in Windy Gap Reservoir following construction of the Windy
Gap connectivity channel diversion structure. Finally, to further validate that the connectivity
channel is being utilized for fish passage, an antenna station will be installed roughly half way
through the connectivity channel to better ascertain the fate of fish moving within the channel but
not detected at both ends of the channel. This antenna will be used to both determine if fish move
partway down the channel and then turn around, and/or if fish are being retained and using the
habitat features constructed within the channel itself. The antennas installed in the inlet and
connectivity channel will be used to monitor fish movement at these locations in 2021 and 2022.
Antenna design, construction, and power needs for the antennas used to achieve the secondary
objectives will be similar to those described above. However, additional personnel will be needed
for antenna installation, upkeep, data collection and archiving, and data analysis.
Portable antennas will be used to obtain the secondary objective of evaluating utilization of
spawning habitat in the headwaters of the Fraser River. Similar to portable antenna deployments
described to meet the primary objectives, multiple runs will be used to increase detection
probability across the width of the channel. Mark and recapture passes will be used to estimate
detection probability, tagged fish (or tag) abundance, and location and fate of fish that have moved
35

�into spawning sites within the Fraser River. GPS data from multiple sampling occasions will be
compared to determine if detected tags remain in live fish (indicated by change in location between
passes and/or sampling occasions) or have been spawned out (no change in movement between
passes or sampling occasions). Tags determined to have been spawned out of the fish will be used
to monitor the use of and identify preferred spawning sites for both Rainbow Trout and Brown
Trout in this section of the river. Portable antennas used to evaluate spawning habitat in the Fraser
River will be deployed during or following the spring and fall spawning periods in 2019-2022.
Fish will be tagged in multiple reaches upstream and downstream of stationary antenna locations.
A large portion of the tagging on the Chimney Rock and Sheriff ranches will occur during the
spring adult population estimates conducted in May 2019-2022. Additional tagging events will be
needed to tag fish in the Colorado and Fraser rivers upstream of Windy Gap Reservoir, likely
concurrent with standard sampling sites conducted in these locations. Additionally, fish may be
tagged as far downstream as Pioneer Park to look at long-distance movements within the upper
Colorado River. Target species for tagging include Rainbow Trout, Brown Trout, and Mottled
Sculpin. Multiple age classes will be tagged of all three target species using 12 mm, 23 mm, and
32 mm tags as appropriate for fish body size and desired detection distance. The number of tags
used will be dependent upon target species and age class abundances. The goal is to tag at least
250 Rainbow Trout and 250 Brown Trout (500 fish total) in the Chimney Rock Ranch reach, with
an additional 250 fish of each species tagged in the Colorado and Fraser rivers upstream of Windy
Gap Reservoir (combined) annually between 2019 and 2022. In the Fraser and Colorado rivers
upstream of Windy Gap Reservoir, up to 250 Mottled Sculpin will be tagged annually. Additional
numbers of fish of each species will be tagged opportunistically dependent upon tag availability
and number of fish captured during electrofishing efforts in each reach. To achieve the secondary
objective of evaluating utilization of spawning habitat in the headwaters of the Fraser River, an
additional 100 Brown Trout and 100 Rainbow Trout will be tagged annually in headwater sections
of the Fraser River near the towns of Fraser and Winter Park between 2019 and 2022. All fish will
be anesthetized using AQUI-S 20E (clove oil), tagged in the interperitoneal cavity using a tagging
gun and needle, and secondarily fin clipped to estimate tag loss. Fish will be held in net pens
following tag insertion to ensure recovery from tagging and anesthetization prior to release back
into the river.
B.3

Study Sites

The project study area will extend from the Fraser and Colorado rivers upstream of the connectivity
channel downstream through the Chimney Rock Ranch to the Sheriff Ranch, incorporating
stationary and portable antenna deployment locations described above to meet the primary and
secondary objectives of the study. Additionally, portable antenna stations may extend further
upstream on the Fraser River to evaluate utilization of spawning habitat in the headwaters of the
Fraser River. Primary fish tagging reaches will include the Chimney Rock Ranch, the Colorado
River upstream of the confluence with the Fraser River, and the Fraser River near Highway 40.
Additional tagging locations for evaluating the utilization of spawning habitat may be located in
36

�the Fraser River closer to the towns of Fraser and Winter Park. An overview of stationary antenna
locations, mobile antenna reaches, and electrofishing sites is presented in Figure B.1.

Figure B.1. Location of stationary sites, mobile antenna reaches, and electrofishing sites for the
proposed fish movement study.
B.4

Monitoring Schedule

Pre-construction baseline movement rates will be monitored in the Colorado and Fraser rivers
upstream of Windy Gap Reservoir and in Chimney Rock Ranch beginning in 2019 and continuing
through the completion of the connectivity channel in 2020. Tagging will begin on the Chimney
Rock Ranch during the spring 2019 adult population estimates and continue annually during these
estimates through 2022. Antenna installation for baseline monitoring would occur in 2019, with
additional antennas being installed in and around the connectivity channel for monitoring the
efficiency of the channel to pass fish in fall 2020 or spring 2021, dependent upon when
construction of the channel is complete. Antennas will remain in place through fall 2022, with
additional tagging events occurring in the Colorado and Fraser rivers in 2020, 2021, and 2022.

37

�B.5

References

Breen, M. J., C. R. Ruetz III, K. J. Thompson, and S. L. Kohler. 2009. Movements of Mottled
Sculpin (Cottus bairdii) in a Michigan stream: how restricted are they? Canadian Journal of
Fisheries and Aquatic Sciences 66:31-41.
Fausch, K. D., C. E. Torgersen, C. V. Baxter, and L. W. Hiram. 2002. Landscapes to riverscapes:
bridging the gap between research and conservation of stream fishes. Bioscience 52:483-498.
Fetherman, E. R., B. W. Avila, and D. L. Winkelman. 2014. Raft and floating radio frequency
identification (RFID) systems for detecting and estimating abundance of PIT-tagged fish in
rivers. North American Journal of Fisheries Management 34:1065-1077.
Fetherman, E. R., D. L. Winkelman, L. L. Bailey, G. J. Schisler, and K. Davies. 2015. Brown trout
removal effects on short-term survival and movement of Myxobolus cerebralis-resistant
rainbow trout. Transactions of the American Fisheries Society 144:610-626.
Ficke, A. D., C. A. Myrick, and M. C. Kondratieff. 2012. The effects of PIT tagging on the
swimming performance and survival of three nonsalmonid freshwater fishes. Ecological
Engineering 48: 86-91.
Fox, B. D., B. P. Bledsoe, E. Kolden, M. C. Kondratieff, and C. A. Myrick. 2016. Eco-hydraulic
evaluation of a whitewater park as a fish passage barrier. Journal of the American Water
Resources Association 52(2):1-23.
Hudy, M., and J. Schiflet. 2009. Movement and recolonization of Potomac sculpin in a Virginia
stream. North American Journal of Fisheries Management 29:196-204.
Lamarre, H., B. MacVicar, and A. G. Roy. 2005. Using passive integrated transponder (PIT) tags
to investigate sediment transport in gravel-bed rivers. Journal of Sedimentary Research
75:736-741.
Lucas, M. C., D. H. Hubb, M.-H. Jang, K. Ha, and J. E. C. Masters. 2009. Availability of and
access to critical habitats in regulated rivers: effects of low-head barriers on threatened
lampreys. Freshwater Biology 54:621-634.
Newby, N. C., T. R. Binder, and E. D. Stevens. 2007. Passive integrated transponder (PIT) tagging
did not negatively affect the short-term feeding behavior or swimming performance of juvenile
Rainbow Trout. Transactions of the American Fisheries Society 136:341-345.
Richer, E. E., E. R. Fetherman, M. C. Kondratieff, and T. A. Barnes. 2017. Incorporating GPS and
mobile radio frequency identification to detect PIT-tagged fish and evaluate habitat utilization
in streams. North American Journal of Fisheries Management 37(6):1249-1264.
Schlosser, I. J., and P. L. Angermeier. 1995. Spatial variation in demographic processes of lotic
fishes: conceptual models, empirical evidence, and implications for conservation. American
Fisheries Society Symposium 17:392-401.
Taylor, P. D., L. Fahrig, K. Henein, and G. Merriam. 1993. Connectivity is a vital element of
landscape structure. Oikos 68:571-573.
Zydlewski, G. B., G. Horton, T. Dubreuil, B. Letcher, S. Casey, and J. Zydlewski. 2006. Remote
monitoring of fish in small streams: a unified approach using PIT tags. Fisheries 31(10):492502.
38

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                <text>In December 2016, a group of partners including American Rivers, Colorado Parks and Wildlife (CPW), Colorado River Water Conservation District (CRWCD), Colorado Water Conservation Board (CWCB), Denver Water, Grand County, Irrigators of Lands in the Vicinity of Kremmling (ILVK), Municipal Subdistrict of Northern Colorado Water Conservancy District (Northern Water), Trout Unlimited, and the Upper Colorado River Alliance was awarded $7.75 million by the U.S. Department of Agriculture Natural Resources Conservation Service (NRCS) through their Regional Conservation Partnership Program (RCPP) for the Upper Colorado River Headwaters Project (Headwaters Project). The Headwaters Project is comprised of three endeavors within Grand County that will collectively restore fish and wildlife habitat, and improve water quality and agricultural water management on a regional scale. The RCPP funding will be utilized to focus on two specific project components: 1) reconnecting the Colorado River upstream and downstream of Windy Gap Reservoir (referenced as the Windy Gap Connectivity Channel Project); and 2) restoration of the Colorado River channel to be resilient to hydrological modifications, while sustaining agriculture, and aquatic and riparian habitat (referenced as the Irrigators of Lands in the Vicinity of Kremmling Project). The third endeavor, the Kemp Breeze State Wildlife Area (SWA) Habitat Restoration Project, is expected to be funded by the partners, and other interested parties. These three project components are further described in the following sections.</text>
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                    <text>Formalin Sensitivity in Rainbow Trout
Adrian Gehr
In collaboration with:
Lindsay Rencoret
Soo-Young Kim
Charlie Vollmer
Departments of Mathematics and Statistics
Colorado State University
Eric Fetherman
Aquatic Research Section
Colorado Parks and Wildlife
Version: May 8, 2014
Abstract
Formalin is a commonly used prophalyctic antifungal and antiparasitic treatment
of fish and fish eggs, yet little is known about the differential sensitivity among strains
after exposure as eggs. This study seeks to determine the sensitivities (measured by
mortality) of four rainbow trout strains, after first exposure to formalin as eggs, and
a later exposure as fingerlings. The data is analyzed using logistic regression and a
Cox proportional hazard model. Both models yield consistent conclusions; the different
strains do die at different rates as fingerlings, but the egg treatment does not contribute
to these differences.

1

Introduction

1.1

Background

Formalin is among the most effective and commonly used antifungal and antiparasitic
treatments in fish and fish eggs (Bills et. al 1977). As such, a better understanding of
the sensitivities of various strains to treatment conditions commonly used in hatcheries
has commercial relevance. Past research has demonstrated different sensitivities among
strains exposed to formalin post-hatch (Piper and Smith 1973), yet little to no research has
explored the effects of exposure as eggs. Therefore, the purpose of this study is to determine
whether different formalin exposure levels as eggs affects mortality later as fingerlings, after
secondary exposure conditions. Four strains of rainbow trout are considered here: pure
Hofer, pure Harrison Lake, 50:50 cross, and 75:25 Hofer:Harrison cross.

1.2

Questions of Interest

1. Does dosage as eggs affect mortality as fingerlings?
1

�2. Are there sensitivity differences among the different strains?
3. Does dosage as fingerlings affect mortality, among fish previously exposed as eggs?
4. Does duration of exposure as fingerlings affect mortality, among fish previously exposed as eggs?
5. How does fish size affect sensitivity to formalin?

1.3

Experimental Design

The experiment is composed of two stages. In the first stage, eggs are treated with formalin for 15 minutes, at two different levels: 1667 or 5000 ppm. Subsequently, the surviving
eggs are allowed to grow to the fingerling stage (approx. 3 inches in size), whereupon they
are re-exposed to one of eight treatment conditions, according to a complete randomized design. The eight fingerling treatment conditions consist of the combinations of four exposure
dosages (0, 167, 250, 500 ppm) at two possible durations (30 or 60 minutes). These levels
are chosen to be in line with common hatchery treatment conditions, which, according to a
survey of Colorado Parks and Wildlife hatchery managers, range from 130-250 ppm, with
167 ppm for 30 minutes being the most common. The 500 ppm condition is included to test
for toxicity at extraordinarily high dosages. After treatment, the fish are observed over five
days, and time of death is recorded. Following the observation period to test for delayed
mortality effects, the the fish are sacrificed (i.e. the data are censored), and the weight,
length, and strain of each fingerling is recorded.

2

Exploratory Data Analysis

We begin by viewing the overall structure of the data, and then move on to visualizations that highlight the effects of different explanatory variables. We consider all possible
explanatory variables that were measured, except length. Length was excluded due to its
high colinearity with weight (r = .958). Weight serves as a general proxy for size. For ease
of reference, the explanatory variables under consideration follow:
X1
X2
X3
X4
X5
X6
X7

=
=
=
=
=
=
=

Duration of exposure
Exposure concentration as fingerlings
Weight as fingerlings
Indicator for pure Hofer strain
Indicator for 50:50 cross
Indicator for 75:25 Hofer:Harrison hybrid
Exposure concentration as eggs

The measured response variable is survival time. It will also be convenient to create a
response vector of zeros and ones, where a one corresponds to a fish that survived and a
zero corresponds to a fish that died. Figure 1 shows that fish tended to either die quickly or
survive until censored, suggesting that we are not losing a much information if we replace
time of death with this binary response vector. Doing so will allow us to analyze the data
with a logistic regression model.

2

�While the majority of fish survived the full duration of the experiment, a significant
fraction did not. To get an idea of which treatments were having an effect on the proportion
of survivors, figure 2 shows a bar graph broken down by treatment.
There are three main things to notice from figure 2:
1. The blueish blocks tend to show substantially higher mortality rates than the reddish
blocks, suggesting that the 60 min group experienced higher rates of death than the
30 min group (i.e. longer duration of exposure as fingerlings appears to increase the
probability of death.)
2. The mortality rate tends to increase within each block as fingerling dosage increases,
suggesting that increased dosage as fingerlings is associated with higher mortality.
3. The two reddish blocks look roughly the same, as do the two blueish blocks. This
suggests that egg treatment may have no significant effect on mortality rate as fingerlings.
Figure 3 attempts to illuminate the other two research questions (2 and 5). There are
couple things to notice from figure 3. The points on the left side (representing fish that
died) are somewhat more densely clumped near the low weight side of the spectrum, and
get more sparse as weight increases, possibly suggesting that increased weight tends to
reduce mortality. However, this requires that the points on the right side of the plot (those
that survived) are not also more clumped on the low weight side of the spectrum, which is
difficult to tell from this plot. Secondly, there appears to be relatively fewer green points,
and relatively more purple and blue points toward the left side of the plot, suggesting that
Harrison Lake strain may be less sensitive, and Hofer may be relatively more sensitive.
We have now provided suggestive answers to our questions of interest. Next we turn to
formal analysis to quantify our results.

3

Formal Analysis

We built two models to analyze the data, both commonly used in survival analysis. First
we will describe a logistic regression model, and follow up with a Cox proportional hazard
(PH) model. The logistic regression model has advantage of simplicity and more intuitive
interpretations, but at the cost of ignoring time of death, and instead treating survival as
an indicator variable. The Cox PH model has the advantage of accounting for the time of
death information, while still appropriately handling the censored nature of the data. Both
models yield consistent results, providing additional confidence for our conclusions.

3.1

Logistic Regression Model

The logistic regression model treats survival as an indicator variable, where any fingerling
that survived for more than 70 hours was coded as ”success” (Y=1) and all others as
”failure” (Y=0). The choice of the 70 hour time cutoff is appropriate because all fish that
survived past 70 hours, did in fact survive until censored (see Figure 1).
In this model, each Yi is assumed to be a Bernouli random variable with probability
of survival pi that depends on the values of the covariates for the ith fish. The value of pi
depends on the covariates according to the following relation:
3

�logit(pi ) = ln(

pi
) = β0 + β1 x1i + β2 x2i + β3 x3i + β4 x4i + β5 x5i + β6 x6i + β7 x7i
1 − pi

where pi = probability of surviving, and the covariates are those outlined above.
In other words, the ln(odds of survival) are assumed to follow a linear relationship with
the covariates.
Equivalently, the model can be stated as:
E[Yi ] = pi = logit−1 (β0 + β1 X1 + ... + β7 X7 ) =

1
1+

e−(β0 +β1 X1 +...+β1 X7 )

This function has a sigmoidal shape and assymptotically approaches 0 and 1, making it
an appropriate choice for modeling a probability measure.
For model selection, we started with all the listed covariates and used successive loglikelihood ratio tests to check for significant effects (i.e Ha : βi 6= 0). By this procedure,
exposure concentration as eggs did not have a statistically significant effect (p = .1297 in
the model with all covariates), while all other covariates were significant at the .05-level
(see Table 1).
It is important to note that the likelihood ratio test works by comparing the goodnessof-fit of a full model to a reduced model that drops the covariate(s) of interest. As such,
the results of likelihood-ratio test depend critically on which covariates are included in the
full model. Nonetheless, these conclusions were robust to model selection effects, as long
as interaction terms were not considered. Specifically, concentration of exposure as eggs
was consistently not significant and all the other covariates consistently were, across many
choices of full model. Interaction terms were ignored for three reasons: because there are
so many possible interactions that could potentially be considered (almost 27 ), because
they complicate the interpretation of the model and in many cases have no straightfoward
interpretation at all, and most importantly, because they are not necessary to answer our
questions of interest. The best model, according to our criteria of parsimony and significant
covariates is given in table 1.

(Intercept)
fingerling concentration
50:50 Hofer:Harrison
75:25 Hofer:Harrison
Hofer
weight
duration

Estimate
7.53
-0.01
-1.63
-0.53
-1.74
0.03
-0.06

Std. Error
0.36
0.00
0.22
0.25
0.23
0.01
0.01

exp(coef)
1863.106
0.990
0.196
0.589
0.176
1.030
.942

p-value
0.00
0.00
0.00
0.03
0.00
0.00
0.00

Table 1: Here, the reported p-values are actually generated using the Wald test, which
approximates the likelihood ratio test for one covariate. The results are very close to what
is given by the likelihood ratio test where the full model has these six covariates, and the
reduced model excludes just the covariate of interest.

4

�The estimated βi s indicate the estimated change in ln(odds of survival) associated with
an increase of one unit in Xi , while holding all other covariates constant. For ease of
interpretation, it is convenient to take eβi which gives the estimated change in odds of
survival associated with an increase of one unit in Xi . Thus, if βi is significantly less than
zero, then increasing Xi tends to harm the odds of survival, while if βi is significantly greater
than zero, then increasing Xi tends to improve odds of survival. Note, though, that we did
not standardize the covariates. Therefore, the magnitude of the βi s can only be compared
directly across the three indicator variables. Other direct comparisons do not make sense,
because the meaning of a one unit increase differs across the variables.
This model allows us to make predictions of the probability of survival for a fingerling
at any level of the covariates. For example, a roughly average weight (10g) Hofer strain
fingerling, treated at 167 ppm for 30 min, has an estimated probability of survival given by:
pi = logit−1 (7.53 + (−.01 ∗ 167) + (−1.74) + (.03 ∗ 10) + (−.06 ∗ 30)) = .932

3.2

Cox Proportional Hazard Model

The Cox proportional hazard (PH) model is concerned with modeling the time to some
event (in our case, the time to death). The model utilitizes the concept of a hazard function,
which intuitively, can be thought of as the instantaneous risk of death at time t. If we define
a random variable T to be the time to death, with a probability density function f(t), and
cumulative density function F(t) = P(T&lt;t), then the hazard function is given by:
h(t) = limδt→0

P (t&lt;T ≤t+δt|T &gt;t)
δt

=

f (t)
1−F (t)

=

f (t)
S(t)

where S(t) = 1 - F(t) = P(T ≥ t) is the survivor function.
The Cox PH model makes the assumption that the hazard function at each level of the
covariates is proportional to some baseline hazard h0 (t). Specifically, the Cox PH model is:
h(t|X1 , ..., X7 ) = h0 (t)e(β0 +β1 X1 +···+β7 X7 )
assuming we consider the same covariates as before.
Equivalently, we can say that the natural log of the hazard ratio is a linear combination
of the covariates. That is,
�
�
7)
ln h(t|Xh10,...,X
= β0 + β1 X1 + β2 X2 + · · · + β7 X7
(t)
The Cox PH model is known as a semiparametric model because it does not require
specification of the baseline hazard; it only assumes that the baseline hazard is nowhere
negative (because a negative hazard would imply immortality). This is acceptable as long
as we only care about the hazard ratio between to levels of the covariates, because in
calculating the ratio, the baseline hazard cancels out, as shown:
h0 (t)e(β0 +β1 x1 +···+β7 x7 )
h(t|x1 , ..., x7 )
=
h(t|z1 , ..., z7 )
h0 (t)e(β0 +β1 z1 +···+β7 z7 )
where x and z represent two different levels of the covariates.
Since no underlying distribution for the hazard function is assumed, the βs must be
estimated using non-parametric methods (specifically, the estimates can be calculated by
using Newton’s method to maximize the partial log-likelihood function.)
5

�Like before, we can use likelihood ratio tests for significance of the covariates. Doing so
tends to yeild p-values very close to that given by the logistic regression model. Again we
find that egg dosage is not significant (p=.1493 in the model with all other covariates.) By
the same criteria as before, we get the same six covariates in the best model. The model is
given in table 2.
dur
treat ppm
factor(color)O
factor(color)P
factor(color)R
weight

coef
0.05
0.00
1.47
0.48
1.55
-0.03

exp(coef)
1.05
1.00
4.37
1.62
4.72
0.97

se(coef)
0.00
0.00
0.20
0.23
0.20
0.01

z
10.73
14.14
7.44
2.11
7.62
-3.09

Pr(&gt;|z|)
0.00
0.00
0.00
0.04
0.00
0.00

Table 2: Here again the p-values are actually given by the Wald test, but are very close to
that given by the likelihood ratio test where the full model has all six covariates and the
reduced model has all but the covariate of interest.
Here, the interpretation of the estimated βs is slightly different than before. In this case,
βi corresponds to the change in the ln(hazard ratio) (instead of ln(odds of survival)) that
is associated with an increase in one unit of Xi , while holding all other covariates constant.
Therefore, unlike before, in the Cox PH model, positive βs correspond to an increase in
sensitivity, and negative βs correspond to a decrease in sensitivity. Note that this would
have been the case in the logistic regression model too if we had instead considered death
as ”success” and survive as ”failure”. In any case, the magnitude of the βs should not be
compared directly across the models, because they mean different things.
Again, it is convenient for interpretation to take eβi , which corresponds to the change in
the hazard ratio for every increase of one unit in Xi , while holding other covariates constant.
An advantage of the Cox PH model is that it allows us to generate survival curves
with associated confidence intervals. In all cases, we see a steep drop in survival early
and then a leveling off. Non-overlaping confidence intervals indicates a significant difference
between the groups. Figure 4 emphasizes the decreased survival among fish in the 60 minute
condition versus the 30 minute condition. It also shows that the Harrison Lake strain is
less sensitive than the Hofer strain. Figure 5 emphasizes the effect of increased dosage as
fingerlings.

4

Conclusions

Based on these results, we can return to our questions of interest, and conclude the
following:
1. There is not sufficient evidence to suggest that the exposure dosage as eggs has any
effect on mortality as fingerlings, within the range tested (p=.1296). This suggests
that hatchery managers do not need to be particularly concerned about the dosage
at which they treat eggs, assuming that they are only concerned with risk of death
later on. Note, though, that this does not imply that dosage as eggs had no effect on
mortality as eggs. That question was not tested in this experiment.
2. The different strains do express differential sensitivities to formalin treatment conditions. Specifically, pure Hofer is the most sensitive (i.e least likely to survive),
6

�followed by the 50:50 cross, then the 75:25 Hofer:Harrison cross, and finally the pure
Harrison Lake strain is the least sensitive. This result is surprising in that the 75:25
Hofer:Harrison cross reacts more like the Harrison Lake strain, despite being genetically more similar to the Hofer strain.
3. Duration of exposure affects mortality, among fingerlings previously exposed to formalin as eggs (p&lt;2e-16). Specifically, longer durations of exposure increase the probability of death.
4. Formalin dosage as fingerlings affects mortality in fingerlings previously exposed as
eggs. Specifically, increased dosage increases the mortality rate.
5. Increased size (as measured by weight) increases probability of survival (p=.0004).
That is, larger fingerlings tend to be less sensitive.

5

Bibliography

Bills, T. D., L. L. Marking, and J. H. Chandler Jr. 1977. Formalin: its toxicity to nontarget
aquatic organisms, persistence, and counteraction. Investigations in Fish Control
Number 73, U. S. Department of the Interior, Washington, D. C.
Piper, R. G., and C. E. Smith. 1973. Factors influencing formalin toxicity in trout. The
Progressive Fish-Culturist 35:78-81.

7

�Survival Time across all Treatment Groups

Frequency

1000

1500

2000

2238

500

607

308

300

221
15
0

10

0

20

81
1

1
40

0

59

0
60

80

100

120

Survival Time (hours)

Figure 1: Note that all fingerlings in the &gt;70 buckets survived the duration of the experiment. Therefore, all those on the right hand side of the histogram can be treated equally
as survivors without great loss of fidelity. The discrepancy is due to a physical constraint
of recording one fish tank at a time at the end of the observational period.

8

�Egg Treatment, Duration as Fingerlings
1667ppm, 30min
5000ppm, 30min
1667ppm, 60min
5000ppm, 60min

0.15
0.10
0.00

0.05

Proportion Died

0.20

0.25

Proportion of Death by Treatment

0

167

500

0

167

500

0

167

500

0

167

500

Fingerling Treatment (ppm)

Figure 2: Mortality rates broken down by treatment group: there is one bar for each of the
sixteen possible treatment combinations. To generate this plot, any fingerling surviving to
&gt;70 hours was coded as having survived, and all others as having died.

9

�Colored by Strain of Fish

0

10

Weight (g)

20

30

40

HL
50:50 GRxHL
75:25 GRxHL
GR

0

20

40

60

80

100

Survival Time (hours)

Figure 3: HL: Harrison Lake, GR: Hofer

10

120

�Figure 4: Note that the data for the two hybrid strains and the data for two of the fingerling
dosage groups is not shown in this figure.

11

�Comparing treatment effect for Hofer and Harrison Lake

0.8
0.4

20

40

60

80

100

120

0

20

40

60

80

100

120

Time

Hofer (duration 60min)

Harrison Lake (duration 60min)

0.8
0.6

formalin = 0 ppm
formalin = 167ppm
formalin = 250ppm
formalin = 500ppm

0.4

0.4

0.6

0.8

Proprotion survive

1.0

Time

1.0

0

Proprotion survive

0.6

Proprotion survive

0.8
0.6
0.4

Proprotion survive

1.0

Harrison Lake (duration 30min)

1.0

Hofer (duration 30min)

0

20

40

60

80

100

120

0

Time

20

40

60

80

100

120

Time

Figure 5: Note that the data for the two hybrid strains is not shown in this figure.

12

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                    <text>Colorado Parks and Wildlife (CPW) Planned Biological Monitoring and
Habitat Restoration Oversight in the Upper Colorado River
Background
The upper Colorado River is an iconic drainage in Colorado that has been severely impacted by
impoundments and reduced flows. Trans-basin and local water use divert approximately 67% of
the flow of the upper Colorado River and future projects will deplete flows further. There are
ongoing discussions on how to implement mitigation measures to reduce the impact of increased
trans-basin water diversions. One component is to reduce effects of the diversions by taking
Windy Gap Reservoir off channel and constructing a bypass around the reservoir. This would
reconnect the river and ameliorate various impacts of a large, on channel impoundment but would
not reduce the impacts of water withdrawals from the system. The planned bypass channel offers
a unique opportunity to evaluate the effects reconnecting the river through the reservoir as well as
investigate mitigation measures to offset the impacts of large water diversions on the ecology of
the river. The objectives of these projects are to evaluate the effectiveness of mitigation measures
in restoring and improving the ecological function of the upper Colorado River.
Invertebrate and Sculpin Evaluations
Current Project Description and Status
The current work on native sculpin and invertebrate species in the upper Colorado River is being
conducted by CPW aquatic research scientist Dan Kowalski. Previous work by CPW research
scientists identified ecological impacts of streamflow reductions and a main stem reservoir (Windy
Gap) on the invertebrates and fish of the river. Native mottled sculpin are currently rare or
extirpated immediately below the reservoir. The health of the invertebrate community declined
after the construction of Windy Gap Reservoir. There has been a 38% reduction in the diversity
of aquatic invertebrates from 1980 to 2011 and 19 species of mayflies, four species of stoneflies
and eight species of caddisflies had been extirpated from the sampling site below Windy Gap
(Erickson 1983, Nehring 2011). Previous work (Kowalski 2014) included mottled sculpin
sampling above and below the reservoir (as well as other impoundments of the upper Colorado
River) corroborated patterns of sculpin distributions and established that sculpin have been
functionally extirpated from the Colorado River below Windy Gap Reservoir. Once common in
this reach, sculpin are now absent for many miles downstream of the reservoir, but become
increasingly common downstream as depletions are offset by other reservoir releases to satisfy
downstream senior water rights.
Future Plans
A large amount of baseline data has already been collected previously under previous projects. To
evaluate the effects of the proposed bypass on invertebrates, sampling is planned to replicate the
historic sampling of invertebrates that occurred pre-impoundment and 25 years post impoundment
(Erickson 1983, Nehring 2011). A subset of sites from previous work will be selected to represent

�the most impacted areas below the reservoir as well as control sites. A before after control impact
study design will be employed for invertebrates using commonly accepted sampling methods.
Five replicate 0.086 m2 Hess samples will be taken from riffles at each study site.
Macroinvertebrate samples will be sorted and sub-sampled in the laboratory using a standard
USGS 300-count protocol, except that replicates will not be composited to allow for an estimate
of within site variation (Moulton et al. 2000). All organisms, except for chironomids and noninsects, will be identified to genus or species while chironomids will be identified to subfamily
and non-insects (e.g., oligochaetes, amphipods) identified to class. In addition to the analysis of
recent samples, data will be compared to previous collections six and 33 years previous.
To monitor any re-colonization of the river below Windy Gap Reservoir by the stonefly
Pteronarcys californica, methods developed by Nehring (2011) and Kowalski (2016) Heinold et
al. (in prep.) will be employed.
To evaluate the effects of the proposed bypass on native mottled sculpin, sampling is planned to
replicate the historic sampling that occurred pre- and post- impoundment (Erickson 1983,
Kowalski 2014, Nehring 2011). A before after control impact study design will be employed for
mottled sculpin using techniques developed on the Colorado River by previous work.
Colorado River Rainbow Trout Establishment
Current Project Description and Status
The current phase of the upper Colorado River rainbow trout reestablishment project, examining
fry stocking as a means to increase the adult rainbow trout population, began in 2013. Currently,
CPW aquatic research scientist Dr. Eric Fetherman is assigned to this project. Rainbow trout fry
have been stocked in to the upper Colorado River study section between Hitching Post Bridge and
the Sheriff Ranch every year since 2013. CPW aquatic biologist Jon Ewert, who is responsible for
fish population assessments in this area, simultaneously stocked the section from the Hot Sulphur
Springs office to Sunset Ranch with similar numbers of rainbow trout fry. To determine fry
survival rates, fry sites (four in the study section and three below Byers Canyon) have been
sampled once a month June through October. Fry infection rates were monitored by collecting
rainbow trout and brown trout fry samples for myxospore counts during fall sampling. Adult
population estimates have been conducted in the spring (late April-early May) to determine if fish
are recruiting to the adult population since 2013.
Future Plans
Fry stocking and evaluations will continue each year before and after the construction of the
bypass. Myxospore counts will be obtained from rainbow trout and brown trout before and after
construction of the bypass to determine if infection levels due to Myxobolus cerebralis, the parasite
that causes salmonid whirling disease, are changed due to reduction of available Tubifex tubifex
(the intermediate host of M. cerebralis) habitat.
Wild trout reproduction response from
construction of the bypass and related stream habitat work (connecting floodplain habitats and
reducing entrenchment) will be evaluated. Adult population estimates will be conducted to
determine if increased recruitment occurs after the construction of the bypass. During these

�estimates, genetic samples will be collected to determine the survival and recruitment of stocked
and naturally produced fry.
Stream Hydrology, Channel Function, and Habitat Evaluations
Current Project Description and Status
CPW hydrologist/ research scientist Eric Richer and CPW research scientist Matt Kondratieff are
involved in technical design assistance and review for the Windy Gap Bypass project and will
serve the same functions for the Upper Colorado River Habitat Project. This review and
oversight function for the entire segment of the upper Colorado River is important to integrate
the numerous aspects of stream habitat work in the drainage with Windy Gap bypass project.
Future Plans
The Windy Gap bypass project presents a unique opportunity to evaluate various aspects of
stable channel design and ecological restoration. The physical habitat characteristics for fish
species should be evaluated for post-project changes by monitoring embeddedness and substrate.
The project will create a new stream channel on a historic floodplain, and the new channel is
likely to experience some adjustments in morphology over time. Monitoring geomorphology
through repeated surveys for monumented cross-sections, the longitudinal profile of the new
bypass channel, and sediment composition will provide valuable information on channel stability
over time and inform maintenance needs. Ideally, bedload transport will be monitored to
determine if the bypass channel is achieving sediment continuity. As establishing riparian
vegetation will be critical for lateral channel stability, monitoring vegetation cover and
composition would help evaluate the need for additional vegetation work. If habitat structures are
used within the bypass channel, rapid assessment could be used to monitoring structural stability
and function which would inform maintenance needs.
Fish Movement and Habitat Use
If funding becomes available, additional work will be conducted in addition to the planned evaluation and
monitoring, which will more specifically evaluate use of the proposed Windy Gap bypass by fish and
invertebrates. RFID (PIT tagging) technology could be used to evaluate fish movement through the
bypass channel, adequacy of attraction flows from the bypass channel, and large-scale movement patterns
for various species in the Upper Colorado River watershed. Adult fish, including rainbow, brown trout,
and sculpins, will be marked upstream and downstream of Windy Gap Reservoir before construction of
the bypass. Portable PIT Tag arrays will be used to track movement of marked fish in the areas upstream
and downstream of the reservoir prior to and after construction of the bypass. Stationary PIT tag arrays
will be installed in the bypass to monitor movement through the newly constructed Windy Gap bypass
channel. Use of portable antennas will further allow the evaluation of fish use of habitat
treatments installed within the bypass. This work will be integrated with the ongoing monitoring of

invertebrates and fish to evaluate re-colonization in the bypass channel itself, as well as the
effects on species composition upstream and downstream of Windy Gap Reservoir.
References

�Erickson, R.C. 1983. Benthic field studies for the Windy Gap study reach, Colorado River,
Colorado, fall, 1980 to fall, 1981. Prepared for The Northern Colorado Water Conservancy District,
Municipal Sub-District.
Heinold, B.D., R.B. Nehring and D.A. Kowalski. In preparation. Estimating the density of benthic and
emerging Salmonflies (Pteronarcys californica) in Colorado rivers.
Kowalski, D.A. 2014. Colorado River aquatic resources investigations. Colorado Division of
Wildlife, Federal Aid in Sportfish Restoration, Project F-237-R21, Progress Report, Fort Collins.
Kowalski, D.A. 2016. Colorado River aquatic resources investigations. Colorado Division of
Wildlife, Federal Aid in Sportfish Restoration, Project F-237-R23, Progress Report, Fort Collins.
Moulton, S.R., II, Carter, J.L., Grotheer, S.A., Cuffney, T.F. &amp; Short, T.M. 2000. Methods of
analysis by the U. S. Geological Survey national water quality laboratory: processing, taxonomy,
and quality control of benthic macroinvertebrate samples. Open-File Report 00-212, U.S.
Geological Survey, Washington D.C.
Nehring, R.B. 2011. Colorado River aquatic resources investigations. Colorado Division of
Wildlife, Federal Aid in Sportfish Restoration, Project F-237R-18, Final Report, Fort Collins.

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                <text>The upper Colorado River is an iconic drainage in Colorado that has been severely impacted by impoundments and reduced flows. Trans-basin and local water use divert approximately 67% of the flow of the upper Colorado River and future projects will deplete flows further. There are ongoing discussions on how to implement mitigation measures to reduce the impact of increased trans-basin water diversions. One component is to reduce effects of the diversions by taking Windy Gap Reservoir off channel and constructing a bypass around the reservoir. This would reconnect the river and ameliorate various impacts of a large, on channel impoundment but would not reduce the impacts of water withdrawals from the system. The planned bypass channel offers a unique opportunity to evaluate the effects reconnecting the river through the reservoir as well as investigate mitigation measures to offset the impacts of large water diversions on the ecology of the river. The objectives of these projects are to evaluate the effectiveness of mitigation measures in restoring and improving the ecological function of the upper Colorado River.</text>
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                    <text>Whitewater Park Projects
Guidance for Reviewing 404 Projects

�COLORADO PARKS AND WILDLIFE
Dan Prenzlow, Director
LEADERSHIP TEAM
Reid DeWalt, Assistant Director for Aquatics, Terrestrial and Natural Resources; Heather
Dugan, Assistant Director for Field Services; Justin Rutter, Assistant Director for Financial
Services; Lauren Truitt, Assistant Director for Information and Education; Jeff Ver Steeg,
Assistant Director for Research, Policy, and Planning; Brett Ackerman, Southeast Region
Manager; Cory Chick, Southwest Region Manager; Mark Leslie, Northeast Region
Manager; JT Romatzke, Northwest Region Manager
STUDY FUNDED BY
Colorado Parks and Wildlife
SUGGESTED CITATION
Kondratieff, M. C., K. R. Bakich, E. E. Richer, D. A. Kowalski, and B. F. Atkinson. 2020.
Whitewater Park Projects: Guidance for Reviewing 404 Projects. Colorado Parks and
Wildlife Aquatic Research Section, Fort Collins, CO. 26 pp.

�Introduction
Colorado Parks and Wildlife’s (CPW) statutory mission is to perpetuate the wildlife resources
of the State, to provide a quality State Parks system, and to provide enjoyable and
sustainable outdoor recreation opportunities that educate and inspire current and future
generations to serve as strategic stewards of Colorado’s natural resources (C.R.S. § 33-9-101
(12) (b)). As CPW is responsible for the management and conservation of aquatic resources
within the State, we are asked to review projects that may affect aquatic habitats or
populations. Specifically, CPW staff is often engaged by the Army Corps of Engineers (USACE)
to review permit applications related to the design, construction, and monitoring of
whitewater parks (WWPs) regulated under Section 404 of the Clean Water Act. WWP projects
typically fall under the following permits:



NWP 27 - Aquatic Habitat Restoration, Establishment, and Enhancement Activities
IP - An individual, or standard permit, is issued when projects have more than minimal
individual or cumulative impacts, are evaluated using additional environmental
criteria, and involve a more comprehensive public interest review.

Recreational in-channel WWPs (Figure 1) are gaining popularity throughout the United States
with Colorado being the epicenter for WWP development. Although WWPs provide economic
and recreational benefits for local communities (Hagenstad et al. 2000; Loomis and McTernan
2011), they can have unintended impacts on aquatic biota, habitat, and river functions. This
is especially true when the hydraulic conditions formed by the WWP differ substantially from
those naturally found in the surrounding river. Natural unmodified river channels are not good
candidates for locating WWPs (American Whitewater 2007). Rather, WWP projects should be
located in areas that have already been substantially modified by past human activities.

A

B

Figure 1. Two typical whitewater park structures include chute-type (A) and drop-type
structures (B)
CPW recommends that adequate environmental safeguards be included in the design and
construction of WWPs to assure that impacts to river functions (Harman et al. 2012),
fisheries, and recreational angling opportunities are minimized. The intent of this document
is to provide USACE with uniform guidance from CPW with regard to project review to assure
that the least environmentally damaging practicable alternative (LEDPA) is followed when
WWPs are proposed, designed, constructed, and maintained over time. CPW offers the
following guidelines to maximize the benefits of recreational WWP opportunities while
1

�minimizing adverse impacts to fisheries and river functions. These are general
recommendations and each project should be reviewed on a case-by-case basis prior to
issuing permits. Failure to demonstrate that the following guidelines were implemented with
due diligence will result in categorical opposition to the project from CPW.
General Recommendations
WWPs are constructed in a wide variety of stream locations utilizing a diverse array of design
elements that are unique to the particular design firm and project engineer, project goals
and expectations, and river conditions. General recommendations for all WWP projects should
include:
1) Early Consultation with CPW: Contact the local CPW Area Aquatic Biologist as early as
possible in the design process to obtain information regarding the species presence, fish
populations, and fisheries management objectives for a proposed project site. CPW
conducts hundreds of fish population surveys on streams and rivers throughout Colorado
annually and uses survey results to inform fisheries population management. Instructions
for submitting formal requests for CPW fisheries data are available at the CPW Aquatics
Data Management webpage. A map of CPW management areas with contact information
for Aquatic Biologists is included in Appendix A and available at the CPW Aquatic
Management webpage.
2) Monitoring and Adaptive Management: Monitoring efforts may focus on physical aspects of
habitat, biological aspects of fish populations, or a combination of both. Monitoring
efforts should be tailored specifically with the goal of detecting undesired or unintended
impacts to the aquatic environment or community. CPW recommends a minimum
monitoring period that includes two years of baseline and five years following project
construction. Data collection should focus on documenting baseline conditions, as-built
conditions, and project effectiveness with at least two monitoring events occurring during
the five year post-construction monitoring period. Adaptive Management provides a
framework that incorporates measurable, relevant monitoring criteria and predetermined
thresholds for acceptable change to assess and address undesired or unintended impacts
to the aquatic environment and communities (Bouwes et al. 2016). Every WWP design
package should include a detailed Adaptive Management Plan (AMP). An AMP should
identify quantifiable monitoring criteria and anticipated impacts to the aquatic
environment that incorporates review and input from CPW and other management
agencies to the USACE. A robust AMP will include a remediation strategy that identifies
stakeholders, resolution processes, and funding sources to engage if project objectives are
not met and thresholds are exceeded.
3) Thresholds for Mitigation Actions: As part of the permitting process prior to issuing permits
and project implementation, regulators should work with CPW to establish the level of
allowable impairment to the natural resource and develop objective thresholds to trigger
mitigation actions, such as requiring structural modifications or off-site mitigation.
Objective and measurable thresholds for changes to river condition and function will
provide enforceable triggers for mitigation or remediation actions.
4) Cumulative Impacts: The potential for cumulative impacts exist when a WWP has two or
more structures. Projects consisting of multiple structures should be reviewed as having
the potential for cumulative impacts. Cumulative impacts should be viewed as more
serious than impacts from a single structure. WWPs have the potential to cause
2

�cumulative impacts to fish passage, fish habitat or both within a single project location or
when a project is located in proximity to other existing manmade river structures (e.g.,
diversion structures, dams, etc.). Some examples of cumulative impacts from WWP
development include: degradation of State-identified high priority habitats and creation
of fish movement obstacles or barriers that limit access by fish to critical forage, refugia,
or reproductive habitats.

Popular whitewater park recreation activities on a Colorado stream.
A CPW fact sheet that provides an overview of WWP research, impacts on fisheries, and
design guidelines has been included as Appendix B.
Fish Passage
WWP structures have the potential to negatively affect fish by fragmenting populations,
reducing migratory ranges, and limiting access to habitat for spawning, feeding, and refuge
(Schlosser and Angermeir 1995). Aquatic habitat fragmentation is ubiquitous throughout
Colorado, contributing to the decline of native aquatic species diversity and abundance (CPW
2015). The elements that create and maintain a desirable play wave (hydraulic jump,
increased velocity, decreased depth, steep-sloping long chutes, abrupt vertical drops, and
grouted smooth stream channel) can create hydraulic conditions that can impede or prevent
upstream fish passage. Suppression of upstream fish movement has been documented at WWP
structures, but the degree of impact varies by fish species, fish size, depths, velocities,
characteristics of individual structures, and variability in flow conditions (Stephens et al.
2015; Fox et al. 2016; Richer et al. 2018). As trout are among the strongest swimming and
jumping species found in Colorado, small-bodied and weaker-swimming fish native to
Colorado streams are even more susceptible to suppression of upstream movement at WWP
structures. Migratory populations of native Colorado suckers, minnows, and trout are also
adversely affected by habitat fragmentation, with some individuals moving long distances (25
or more miles) during upstream migrations to access spawning habitat (Kondratieff et al.
3

�2017; Thompson et al. 2019). To minimize loss of fish passage functions, CPW recommends
the following guidelines be incorporated into the design of WWP projects:
1) Target Species and Life Stages: Design WWP structures to allow upstream fish passage for
all species present at the project site, unless there are specific management objectives
that warrant exclusion of particular fish species. Fish passage elements are expected to
pass juvenile and adult life stages (Forty et al. 2016).
2) Design Flows: Fish passage design elements of WWP structures should be designed to
provide passage across a range of typical flows, including flows corresponding with the
timing of critical life history movement events such as spawning migrations or access to
refugia. The average daily discharge that is exceeded 95% and 5% of the time should be
selected for the low and high fishway design flows, respectively (NMFS 2008).
3) Fishway Invert Elevations: Fishways are engineered pathways specifically designed to
accommodate fish movement around or through WWP structures. Fishways should provide
passable conditions over a range of flow conditions. Fish passage design elements should
be constructed so that the upstream invert of the fishway exit is located at a lower
elevation than the upstream invert of the WWP recreation structure crest to ensure that
the fishway functions during extreme drought or low flow conditions.
4) Instream Flows: Fishways should have sufficient capacity for carrying either: 1) the
decreed instream flow (if a Colorado Water Conservation Board (CWCB) instream flow
water right exists for the stream or river in question), or 2) where no decreed instream
flow exists, a minimum flow volume that is reasonably necessary for maintaining fish
passage in the stream or river in question.
5) Attraction Flows: Sufficient attraction flows at the downstream fishway entrance is a
critical factor that will affect efficiency of the fish passage structure. The hydraulic
conditions (i.e., velocity, depth, and turbulence), quantity, and location of attraction
flows are all important design considerations. In general, increasing the amount of
attraction flow relative to the total river flow will increase the effectiveness of the
fishway for providing upstream passage. The minimum attraction flow necessary to
provide adequate attraction conditions for fish is 5-10% of the total river flow (NMFS
2008).
6) Passage Criteria: WWP designs should provide comparisons of hydraulic conditions within
the fishway to species-specific design criteria for identifying limiting swimming speeds,
water depths, and vertical drops that ultimately provide evidence for the effectiveness of
fish passage conditions. Hydraulic modeling results should include depths, velocities, and
locations of hydraulic jumps for existing and proposed conditions so that fish passage
hydraulic conditions can be evaluated before the project is implemented. Fish passage
criteria including the swimming speeds and jumping heights for Colorado fishes are
included in a CPW fact sheet on Fish Passage at River Structures (Appendix C).
7) Incorporation of Natural Channel Forms and Processes: Fish passage design elements
should function to enhance, maintain or mimic the pre-existing natural stream conditions
(gradient, depth, velocity, and channel roughness) found at the proposed site of each
recreational drop structure. This is especially important when there is a lack of speciesspecific swim passage criteria.
8) Monitoring Fish Passage: Various methods have been used to evaluate fish passage at
instream obstacles or barriers. Fish passage efficiency through WWPs has been monitored
4

�using hydraulic modeling by 3-dimensional hydraulic models (Stephens et al. 2016), 2dimensional hydraulic modeling (Hardee 2017), and a least-cost path approach combining
known swim speeds and 2-dimensional hydraulic modeling (Brubaker et al. 2018). Fishway
evaluations can also be conducted by marking individual fish and monitoring their
movements over time (i.e., PIT tag or Mark-Recapture studies; Fox et al. 2016). A
combination of hydraulic modeling and validation studies using marked fish provide the
strongest support for monitoring fishways by utilizing multiple lines of evidence. Ideally
fish passage evaluations collect fish passage data from a) the pre- project reach, b) a
nearby control site (up or downstream of the project reach) that is representative of a
natural condition, c) the post-project reach, or d) a combination of some (Before/After or
Control/Impact) or all sites (i.e., a full BACI study design). Project objectives should be
measureable and monitoring of pre- and post-project conditions should be used to
evaluate project effectiveness and inform adaptive management. CPW recommends a
minimum monitoring period of one year of baseline and three years following WWP
project construction, with an emphasis on documenting baseline conditions, as-built
conditions, and project effectiveness with at least two monitoring events during the postconstruction three-year period.
9) Adaptive Management: AMPs should evaluate proposed and post-project changes to
hydraulics and topographic conditions including water depth, velocities, hydraulic jumps,
and bed drops at each constructed WWP feature over the range of design flows. These
criteria should be used to evaluate project objectives and thresholds for requiring
mitigation actions. AMPs should be included as part of the design package for each WWP
project.

Whitewater park structure with engineered fish bypass
Fish Habitat
WWP structures and their placement within a stream channel have the potential to degrade
aquatic habitat quality. Many factors influence aquatic habitat quality and should be
5

�incorporated into WWP designs. The placement of WWPs in channels should follow published
geomorphic criteria for physical relationships related to channel width, pool spacing, and
riffle lengths to minimize the potential for channel instability and habitat impairment
(Leopold et al. 1964, Dunne and Leopold 1978). The valley type, process domain, stream
gradient, stream hydrology, and substrate characteristics should be used to inform WWP
designs and the placement of structures within the proposed reach. Fish behavioral traits, life
history characteristics, physiological tolerances, and swimming and jumping capabilities are
directly related to the physical habitat characteristics found in the natural channels which
they occupy. Low gradient stream channels in unconfined valleys or plains are typically
occupied by fish species that are incapable of jumping over vertical obstacles and have
resident fish with weaker swimming capabilities. High gradient mountain streams are typically
occupied by fish species that are capable of jumping and have swimming capabilities that
enable them to burst through high velocities and turbulence. Some weaker swimming, smallbodied fishes have behavioral traits that cause them to avoid swimming over deep pools
where they are vulnerable to predation. Instead, they utilize the lateral edges of stream
channels (Swarr 2018). Research has shown that impacts of WWPs on rivers and fisheries is
very specific and will depend on the specific conditions at a river site as well as the fish
populations present (Kowalski 2019). Ultimately, design and construction of WWP structures
should provide for fish and aquatic invertebrate habitat; such structures must take account of
the preservation of functional riverine and aquatic processes and maintain the natural
aesthetic qualities of the river to the greatest extent possible. CPW recommends the
following guidelines be incorporated into the design of WWP projects:
1) Minimize Extreme Hydraulic Conditions in WWP Pools: Natural pools located in unconfined
valleys with low channel bed slopes are characterized by predictable and relatively stable
hydraulic conditions that provide a balance between feeding and resting for fish. Fish feed
on aquatic prey drifting into pools from upstream riffles and find low velocity resting
areas close to the bed within pools that minimize energetic demands for fish swimming
and maintaining equilibrium. Although WWPs create deep pools, observed fish densities
and biomass were higher in natural pools than in WWP pools for trout and native fish
(Kolden et al. 2015). A combination of hydraulic modeling and direct field measurement
of hydraulic conditions present in WWP pools found higher turbulence (6×), vorticity (2×),
velocity (3×), surging (40×), and depth (2×) were observed in WWP pools as compared to
natural pools. Habitat suitability scores incorporating depths and velocities for Rainbow
Trout (Oncorhynchus mykiss) and Brown Trout (Salmo trutta) were higher for pools
located in WWP reaches than natural pools. However, fish abundance estimates (biomass
and densities) for WWPs were lower, providing evidence that the use of habitat suitability
scoring to quantify habitat conditions for pools located in WWPs may be inappropriate.
Direct measurements of fish abundance (biomass and densities) are preferred over habitat
suitability modeling for evaluating WWP pool quality until more research can be done to
incorporate additional information to improve model performance. Lower fish abundance
may be explained by conversion of food producing riffles to impervious grouted drops over
WWP structures, increased hydraulic variability (turbulence, vorticity, velocity, and
surging) characteristic of WWP pools, or a combination of these factors. Based on results
from Kolden et al. (2015), habitat suitability scoring through use of hydraulic models for
pools may not be as reliable an indicator of overall habitat quality as direct estimation of
fish abundance. Additional studies in Colorado have indicated that impacts to fish
populations are site-specific and can be subtle (Kowalski 2019). If structures are designed
and spaced properly, impacts to fish populations can be reduced. In some rivers, WWP
structures have been shown to increase habitat suitability and density of non-native and
6

�non-game fish species like White Sucker (Catostomus commersonii) and Longnose Sucker
(Catostomus catostomus) while large scale impacts to trout populations can be minimized.
2) Design Pool-to-Pool Spacing to Match Expected Ranges from Geomorphic Relations: Pool to
pool spacing within WWPs are often outside the range of natural variability found in
natural channels of the same valley and stream type (Leopold et al. 1964, Dunne and
Leopold 1978). Most often, pools in WWPs are more closely spaced than what would be
found in natural stream reaches of the same geomorphic context. This can result in
increased channel instability from accelerated erosion or deposition with pools filling with
sediment and the need for frequent maintenance and removal of sediments from WWP
pools. Pool spacing also can have an impact on aquatic invertebrate populations.
Improperly designed and spaced WWP structures can remove natural riffles from river
reaches and reduce the diversity of aquatic invertebrates (Kowalski 2019). Designing river
channel features that are in balance with the stream flow and sediment supply of each
specific site is important in preserving natural hydraulic and biological functions of riffles.
3) Preservation of Riffle Habitats: WWP structures are commonly constructed using concrete
grout, pre-cast concrete blocks, and large boulders used to direct flow and manipulate
hydraulics. The materials used for WWP structures either fill-in or replace interstitial
spaces normally found in coarse riffle habitat where macroinvertebrates reside, juvenile
trout find refuge, and native fish such as Mottled Sculpin (Cottus bairdii), dace, and
suckers live out most of their life cycles. Large, grouted WWP structures have been shown
to support less diverse aquatic invertebrate communities then natural riffles (Kowalski
2019). WWP designs should preserve some riffles within the project reach instead of
converting all riffles to WWP drops. Individual WWP structures require a drop in elevation
to function optimally and thus WWP drop structures often replace natural riffle features.
As a consequence, WWP reaches typically have proportionally less riffle habitat as
compared to adjacent natural stream reaches within the same valley and stream type. A
reduction in overall riffle habitat (area) could result in less macroinvertebrate habitat and
consequently less food for fishes residing in WWP reaches. Research has shown that in
some rivers in Colorado, improperly spaced structures degrade riffle habitat and reduce
the diversity of aquatic invertebrates while properly designed structures that include
riffle habitat can be as diverse and productive as natural riffles (Kowalski 2019). Riffle
habitat converted to impervious, grouted structures may result in a loss of interstitial
habitat critical for providing habitat for macroinvertebrates, native benthic fishes like
sculpin and dace, and juvenile rearing habitat for trout.
4) Monitoring Fish Habitat: A variety of methods have been used to evaluate fish habitat at
WWPs including 2-dimensional and 3-dimensional hydraulic modeling of the natural
(control) and project reach incorporating habitat suitability criteria (Kolden et al. 2015).
Fish habitat conditions can also be evaluated by directly surveying fish populations (fish
density and biomass) before and after project construction and/or within and outside of
the constructed WWP reach. Ideally fish habitat evaluations collect data from a) the preproject reach, b) a nearby control site (up or downstream of the project reach) that is
representative of a natural condition, c) the post-project reach, or d) a combination of
some (Before/After or Control/Impact) or all sites (i.e., a full BACI study design). Project
objectives should be measureable and monitoring of pre- and post-project conditions
should be used to evaluate project effectiveness and inform adaptive management. CPW
recommends a minimum monitoring period of two years for baseline and five years
following construction, with an emphasis on documenting baseline conditions, as-built
7

�conditions, and project effectiveness with at least two monitoring events during the postconstruction five-year period.
5) Adaptive Management: AMPs should evaluate proposed and post-project changes to the
river environment including pool-to-pool spacing, hydraulic variability (turbulence,
vorticity, velocity, and surging), changes to the proportion of riffle habitat, changes in
fish population or habitat suitability, and riparian area. These criteria can be used to
inform and develop project objectives and thresholds.
Sediment Deposition
The placement of WWPs in river channels should follow established geomorphic criteria
(Leopold et al. 1964; Dunne and Leopold 1978) for physical relationships of channel width,
pool spacing, and riffle lengths to minimize the potential for channel instability, habitat
impairment, as well as decrease the frequency of structure maintenance and in-channel
disturbance. Sediment characteristics and related processes will vary by valley type, process
domain, stream gradient, and stream hydrology. These factors should be incorporated WWP
designs and inform the placement of structures within the proposed reach.
1) Minimize Sediment Deposition: WWP structures should not disrupt or curtail sediment
transport by inducing sediment deposition upstream or downstream of the structure.
Sediment deposition can eliminate preferred fish and benthic macroinvertebrate habitats,
as well as create favorable conditions (finer substrate) for the spread of whirling disease
in trout. Sediment deposition could also result in reduced channel capacity which could
increase flooding risk to surrounding areas. Sediment deposition will likely be inevitable at
WWP due to the loss of energy over the structure, so maintenance plans to periodically
remove excess sediment are needed.
2) Avoid Rapid Contraction and Expansion in WWP designs: WWPs can create a sequence of
rapidly contracting and expanding riverbank conditions that lead to problems of
sedimentation in pools and a need for more frequent maintenance to remove the excess
sediment. WWP designs should incorporate knowledge of channel maintenance flows
(bankfull conditions) and the potential for contraction/expansion to induce sediment
deposition within WWP reaches.
3) Adaptive Management: AMPs should evaluate proposed, pre-, and post-project changes to
the river environment in the longitudinal profile and representative cross sections
including changes to streambed substrate characteristics. Monitoring should document
areas of sediment deposition, fine sediment deposition areas, and bank erosion. Projects
should consider seeking Colorado 401 Water Quality Certification from the Colorado Water
Quality Control Division and conduct pebble counts at critical cross sections before and
during the post-project monitoring period. Pre-project monitoring data should be used to
inform project objectives and establish thresholds.

8

�Sediment deposition in whitewater park pools
Site Selection
Properly locating WWPs within river systems is one of the best ways of minimizing physical,
ecological, and social impacts to rivers and streams including channel stability, sediment
deposition, fish passage, fish habitat, and recreational angling. Site locations that have been
identified as existing barriers to fish movement (e.g. diversion structures or dams) and that
been heavily modified by past human activities are preferred locations for WWPs.
1) Step-Wise Hierarchical Decision Making Framework for Site Selection: CPW proposes the
following steps be carried out in a step-wise fashion according to the following
hierarchical framework to ensure that the LEDPA is selected during the site selection and
design process. The level of risk with respect to impairment of fish passage and habitat
increases with each successive step. Therefore, more intensive monitoring evaluations
should be commensurate with increasing levels of risk. WWP designs must provide
justification for the following:
a. High Priority Habitat and Special Management Reaches: WWP projects proposed in the
following designated river reaches will result in categorical opposition from CPW
including the following: 1) Designated Cutthroat Trout Waters, 2) Critical Habitat for
Threatened and Endangered Species as well as reaches identified as sensitive habitat
for State Species of Concern, and 3) Gold Medal Waters.
b. Geomorphic Setting: Provide justification for a WWP design alternative that is located
in an unconfined valley setting (unconfined valleys have an Entrenchment Ratio (ER)
&gt;2.2) and stream channel slopes that are 2 % or less. Geomorphic settings consisting of
artificially or naturally confined valleys and Rosgen A, B (step-pool), F, and G stream
types have hydraulic characteristics more similar to those produced by WWP structures
and therefore contain fish species and assemblages that are adapted to living in similar
hydraulic conditions as those commonly associated with WWPs.
c. Partially Channel-Spanning Structure: If a site location cannot be identified within the
appropriate geomorphic setting, provide justification for a WWP design alternative
9

�that requires full channel-spanning structures. Partially channel-spanning structure
designs should be used unless project goals or site constraints dictate that a fully
channel-spanning structure is required.
d. Natural Channel Split: If a partial channel-spanning structure is not possible, provide
justification for a WWP site location on a single-thread channel site. Channel splits can
serve dual functions with one split providing WWP recreation while the other channel
split is left as natural and unmodified. A split branch of an existing channel (side
channel or one side of an existing island) should be used unless project goals or site
constraints dictate that a single-thread channel site is required.
e. Artificial Channel Split: If a suitable site location cannot be found on a natural split
branch of an existing channel, provide justification for a WWP site location on an
artificially-constructed split branch channel (constructed side channel or one side of
an artificially-constructed island). Artificial channel splits can serve dual functions
with one split providing WWP recreation while the other channel split is designed to
mimic natural, reference-like conditions. An artificially-constructed split flow channel
(constructed side channel or one side of an artificially-constructed island) should be
used unless project goals or site constraints dictate that a single-thread channel site is
required.
f. Technical Fishway: If the site location is constrained such that an artificiallyconstructed split flow channel is not possible, technical fishway concepts (such as a
constructed bypass channels or riffles, rock ramps, or vertical slots) must be
incorporated into WWP structure designs.
g. No Fish Passage Elements: WWP designs that do not incorporate fish passage elements
into structures are not acceptable and will result in categorical opposition from CPW.
2) Minimize WWP Recreation Conflicts with Anglers and Non-Whitewater Recreation Boaters:
WWP sites should be located to avoid recreational conflicts with anglers and nonwhitewater recreational boaters (i.e., drift boats and canoes). Hydraulic conditions
formed by WWP structures can impede safe boat travel for non-whitewater recreational
boaters. Within WWPs there is an increased potential for whitewater recreational boaters
to displace stream anglers, especially during the summer months. Incompatibilities
between stream anglers and recreational boaters exist. Creel survey data from Colorado
and Wyoming suggest that stream anglers prefer to fish in locations that are uncrowded,
provide pleasant conditions close to nature, and are relaxing. The conditions commonly
encountered at and in the vicinity of WWPs include artificially armored and terraced
banks with minimal vegetation and encourage spectating crowds.
3) Mitigation for Lost Angler Opportunity and Access: Mitigation should be considered to
replace lost angler access, infrastructure, and fishing opportunity. There is a history of
new WWP construction within or replacing existing Fishing Is Fun (FIF) habitat projects
funded through Federal sportfish dollars at sites in Colorado including Pagosa Springs,
Basalt, Ridgway, and Montrose. When new WWPs are proposed within heavily used urban
fishing areas, intensively managed fisheries (those with special harvest restriction
regulations), Gold Medal designated fisheries, FIF-funded habitat projects, or locations
with existing amenities (such as parking access, trails, boat launches, picnic areas, etc...)
funded by Federal sportfish dollars or other fishing interest groups (i.e., Trout Unlimited),
10

�reasonable mitigation is necessary. Mitigation may consist of replacing lost or degraded
infrastructure and amenities, increasing infrastructure to accommodate the new users,
and providing reasonable additional access points or developing alternative locations for
anglers nearby.
4) Off-Site Mitigation: Mitigation locations for offsetting loss of fish habitat from WWP
development should not occur within WWP project reaches, but should occur in separate
locations up- or downstream within the same river watershed if possible. Mitigation
possibilities include developing new areas open to fishing access or enhancing fish habitat
in an area that is not heavily impacted by recreational boating. Fish passage cannot be
mitigated off-site and must be accommodated through a WWP project.
Whitewater Park Project Applications
CPW will only provide technical design review for projects that submit complete applications.
Requests for design reviews must include complete permit applications with all pertinent
information, including project goals and objectives, a design report and plan set, assessment
of existing conditions, list of river stakeholders, revegetation plan, monitoring plans, and a
description of how the project will be maintained over time.
1) Early Consultation with CPW: Contact the local CPW Area Aquatic Biologist as early as
possible in the design process to obtain information regarding the species presence, fish
populations and fisheries management objectives for a proposed project site. CPW
conducts hundreds of fish population surveys on streams and rivers throughout Colorado
annually and uses survey results to inform fisheries population management. Instructions
for submitting formal data requests are available at the CPW Aquatics Data Management
webpage and contact information for CPW Aquatic Biologists is included in Appendix A.
2) Project Goals and Objectives: Applications must clearly identify project goals and
objectives, and describe the context and analysis leading up to the established LEDPA (if
already developed). The applicant should address the potential for fishery and ecological
impacts from the project and how they relate to the project goals and objectives.
3) Design Report and Plan Set: The design report should include a comparison of WWP project
alternatives that were used to inform the selection of the proposed WWP design. The
design report and plan set should clearly detail existing conditions of the proposed WWP
reach, description and layout of the proposed WWP reach, detailed description of fish
passage design elements proposed for each structure, description of existing hydrology and
design flows as they relate to fish passage design elements, and modeled hydraulic
conditions through each WWP structure and fish passage design elements.
4) Assessment of Existing Conditions: An assessment of existing conditions within the
proposed project reach should be conducted in order to determine the level of
anthropogenic impacts at the proposed site. What is the level of departure from a
reference or historic condition with respect to existing hydrology, hydraulics (floodplain
connectivity), geomorphology (sediment supply), physicochemical, and biological
condition? Consider application of the Colorado Stream Quantification Tool (SQT) (CSQT SC
2019) as a means to assign proposed project reach as Functioning, Not Functioning, or
Functioning at Risk. CPW advocates installation of WWPs in reaches that rate as Not
Functioning or Functioning at Risk to minimize impacts to “natural, unmodified” river
11

�channels. Ultimately, is the proposed site located in a natural or already significantly
modified site?
5) Other River Users and Stakeholders: A complete list of river user groups (stakeholders)
should be provided with the project application materials, or at least made aware of the
proposed WWP project.
6) CPW Consultation and Supervision: Early consultation with CPW area staff including a
description of the longitudinal extent of the project, whether or not water rights are a
part of the project (i.e., RCID), and a complete list of project goals. WWPs almost always
involve significant instream structures; both CPW and the CWCB believe that these
structures should be designed and their construction supervised by a Colorado registered
professional engineer and/or a professional hydrologist in consultation with both agencies.
7) Grout: Minimize the use of grout for construction of WWP structures except as needed to
maintain recreation function, human safety, and fish passage elements associated with
the WWP and as part of the criteria for selecting the LEDPA. If grout is used, recess the
grout elevation so it is not flush with the top of the structure elements, leaving spaces
between boulders or blocks for increased roughness and cover for small aquatic organisms.
Recessing grout and spacing boulders to create continuous pathways in the wing walls may
improve conditions for upstream fish passage at WWP structures.
8) Revegetation Plans: Riparian vegetation composed of native species is the primary control
for bank stability in many stream types and should be used to improve long-term stability
of the project. Revegetation plans should be included with the plan set for the project,
including success criteria, planting protocols, irrigation needs, weed control, and postconstruction stewardship. Designs should utilize biostabilization techniques to stabilize
disturbed streambanks as outlined in Living Streambanks: a Manual for Bioengineering
Treatments for Colorado Streams (Giordanengo et al. 2016).
9) Grade Control other than WWP Structures: If grade control is needed for proper function
of the WWP structure, hardened riffles comprised of boulder sills buried in native
substrate is suggested as an alternative to in-channel grout. Hardened riffles can also be
used to protect downstream riffle heads. All proposed structures (recreational or
otherwise) within a project that are necessary to the successful function of the proposed
WWP project are expected to meet LEDPA criteria in the project reach. We expect USACE
to consider all structures associated with a WWP project to be regulated as part of the
project and not subject to exemption.
10) RICDs: Recreational In-channel Diversion (RICD) water rights can be acquired for WWPs in
Colorado to provide recreational experiences in and on the water. RICDs should be
designed, constructed, and managed to minimize or avoid impacts to native and sport
fish. Flows deviating from the natural flow regime, such as water calls during spawning
periods or when young-of-the-year fish are emerging from spawning gravels, could have
adverse impacts on stream ecology (Poff et al. 1997). Meeting with CPW should be
conducted prior to applying for a water right tied to a specified location in the river
system (i.e., RICD). RICD proponents are strongly encouraged to contact USACE to
conducting a feasibility assessment for WWP development at a specific site location prior
to pursuing a water right. Federal regulations do not account for or prioritize permitting
of RICD water rights. This will improve efficiency of State resources and time for a project
12

�that is not federally permittable due to either location or design that does not meet
LEDPA criteria.
11) Post-Construction Site Visit: Require a post-construction site visit prior to rewatering of
structures when possible to verify as-built design and include CPW to identify any aquatic
resource concerns prior to rewatering.
12) Monitoring and Evaluation: Develop monitoring objectives that are measureable and use
objective monitoring criteria for pre- and post-project conditions to evaluate project
effectiveness and inform adaptive management. Comparisons between the proposed and
as-built conditions should be made to define allowable impacts and without exceeding
thresholds for requiring mitigation action.
13) Maintenance and Stewardship: WWP design plans should address structure maintenance,
sedimentation, and debris removal as part of stewardship considerations for at least five
years following project completion.

Whitewater park kayakers on a popular Colorado mountain river
Best Management Practices
1) Spawning Periods: Construction activities that cause streambed disturbance should not be
scheduled during periods when adult spawning migrations, egg incubation, or fry swim-up
are occurring. Fish eggs and fry may die if construction activities mobilize fine sediment
that smothers the streambed in which they reside. Repetitive and cumulative streambed
disturbances during critical reproductive periods can significantly affect population
dynamics and resiliency of local fisheries. In general, instream construction should be
13

�targeted for the months of August and September when flows are lower and impacts to
spawning fish and incubating eggs are less likely. Early communication with CPW is
encouraged as this suggested window could vary based on local considerations such as
elevation, environmental variability, and fish species present.
2) Invasive and Nuisance Species: To prevent the spread of invasive and/or nuisance species
(e.g., Asian Clam, Green River Mud Snail, New Zealand Mud Snail), we strongly encourage
that heavy equipment be cleaned prior to and after construction if the equipment was
previously used in another stream, river, lake, pond, or wetland within ten days of
initiating work. The following methods are recommended for preventing the spread of
invasive aquatic organisms:
a. Disinfection with QAC: Remove all mud and debris from equipment (tracks, turrets,
buckets, drags, teeth, etc…) and spray/soak equipment with a disinfection solution
containing quaternary ammonia compound (QAC). Treated equipment must be kept
moist for at least 10 minutes. The recommended concentration for any commercially
available QAC product used to disinfect equipment is 6 ounces of QAC solution per
gallon of clean water. The following QAC products have been tested by CPW and are
listed in order from highest to lowest concentration of active QAC: Green Solutions
High Dilution Disinfectant 256, Super HDQ Neutral, Quat 4, Vedco 128, and Quat 128.
b. Disposal of QAC: Wastewater treatment plants are capable of processing water
containing small amounts of QAC. Therefore, rinsing used QAC solutions down a
sanitary sewer is a safe method of disposal. However, QACs should be kept out of
storm sewers and other waterways. Always dilute old product before rinsing down
sanitary sewers directly from the container, and follow MSDS and label
recommendations regarding rinsing and disposal of empty containers. Small amounts of
QAC from spray disinfection may come in contact with the environment with few
negative effects. However, it is not recommended to dump large amounts of QAC
solutions directly on the ground. More detailed instructions for disinfection with QAC
products can be provided upon request.
c. Disinfection with Hot Water: Spray/soak equipment with water heated to a
temperature greater than 140 degrees Fahrenheit for at least 10 minutes.
3) Turbidity: Instream construction should be conducted in a manner that will minimize
turbidity of the water in the work area.
4) Petroleum Products and Chemicals: No petroleum products, chemicals, or other
deleterious materials should be allowed to enter or be disposed of in such a manner in
which they could enter the waterway or adjacent wetlands. Accordingly, we recommend
that oil absorbent “booms” be installed downstream of the project site during
construction activities.
References
American Whitewater, 2007. Whitewater parks-considerations and case studies.
https://www.americanwhitewater.org/content/Wiki/stewardship:whitewater_parks
Bouwes, N., S. Bennett, and Joe Wheaton. 2016. Adapting adaptive management for testing
the effectiveness of stream restoration: An intensively monitored watershed example.
Fisheries, 41:2, 84-91, DOI: 10.1080/03632415.2015.1127806.
14

�Brubaker, A., E. E. Richer, D.A. Kowalski, and M.C. Kondratieff. 2018. Making waves: The
effects of whitewater parks on fish passage. 43rd Annual Meeting of the Western Division
of the American Fisheries Society. Anchorage, Alaska. May 22, 2018.
Colorado Stream Quantification Tool Steering Committee (CSQT SC). 2019. Colorado Stream
Quantification Tool and Debit Calculator (CSQT) User Manual, Beta Version. U.S.
Environmental Protection Agency, Office of Wetlands, Oceans and Watersheds (Contract #
EPC-17-001), Washington, D.C.
CPW (Colorado Parks and Wildlife). 2015. State Wildlife Action Plan. Denver, Colorado.
Dunne, T., &amp; Leopold, L. (1978). Water in environmental planning. San Francisco, California:
W.H. Freeman and Company. 818 pp.
Forty, M., J. Spees, and M. C. Lucas. 2016. Not just for adults! Evaluating the performance of
multiple fish passage designs at low-head barriers for the upstream movement of juvenile
and adult trout Salmo trutta. Ecological Engineering 94:214-224.
Fox, B.D., B.P. Bledsoe, E. Kolden, M.C. Kondratieff and C.A. Myrick. 2016. Ecohydraulic
evaluation of whitewater parks as a fish passage barrier. Journal of the American Water
Resources Association. DOI: 10.1111/1752-1688.12397.
Giordanengo, J. H., R. H. Mandel, W. J. Spitz, M. C. Bossler, M. J. Blazewicz, S. E. Yochum,
K. R. Jagt, W. J. LaBarre, G. E. Gurnee, R. Humphries, and K. T. Uhing. 2016. Living
streambanks: A manual of bioengineering treatments for Colorado streams. Colorado
Water Conservation Board, Denver.
Hagenstad, M., J. Henderson. R. S. Rauncher, J. Whitcomb. 2000. Preliminary evaluation of
the beneficial value of waters diverted in the Clear Creek whitewater park in the city of
Golden, Stratus Consulting.
Hardee, T.L. 2017. Evaluating fish passage at whitewater parks using a spatially explicit 2D
hydraulic modeling approach. M.S. Thesis, Department of Civil and Environmental
Engineering, Colorado State University. 107 pp.
Harman, W., R. Starr, M. Carter, K. Tweedy, M. Clemmons, K. Suggs, C. Miller. 2012. A
function-based framework for stream assessment and restoration projects. US
Environmental Protection Agency, Office of Wetlands, Oceans, and Watersheds,
Washington, DC EPA 843-K-12-006.
Kolden, E., B.D. Fox, B.P. Bledsoe, and M.C. Kondratieff. 2015. Modelling whitewater park
hydraulics and fish habitat in Colorado. River Research and Applications. DOI:
10.1002/rra.2931.
Kondratieff, M. C. and E. E. Richer. 2017. Stream Habitat Investigations and Assistance.
Federal Aid Project F-161-R23. Colorado Parks and Wildlife, Aquatic Research Section.
Fort Collins, Colorado.

15

�Kowalski, D.A. 2019. Colorado River aquatic resource investigations. Federal Aid Project F237-R26. Colorado Parks and Wildlife, Aquatic Wildlife Research Section. Fort Collins,
Colorado.
Leopold, L., Wolman, G., &amp; Miller, J. (1964). Fluvial processes in geomorphology. San
Francisco, California: W.H. Freeman and Company. 544 pp.
Loomis, J., and J. McTernan. 2011. Fort Collins whitewater park economic assessment.
Department of Agricultural and Resource Economics, Colorado State University.
NMFS (National Marine Fisheries Service). 2008. Anadromous salmonid passage facility design.
NMFS, Northwest Region, Portland, Oregon.
Poff, N. L., J. D. Allan, M. B. Bain, J. R. Karr, K. L. Prestegaard, B. D. Richter, R. E. Sparks,
and J. C. Stromberg. 1997. The natural flow regime: a paradigm for river conservation and
restoration. BioScience 47(11): 769-784.
Richer, E.E., E.R. Fetherman, M.C. Kondratieff and T.A. Barnes. 2017. Incorporating GPS and
Mobile Radio Frequency Identification to Detect PIT-Tagged Fish and Evaluate Habitat
Utilization in Streams. North American Journal of Fisheries Management. DOI:
10.1080/02755947.2017.1374312.
Richer, E. E., A. B. Brubaker, D. A. Kowalski, and M.C Kondratieff. 2018. Making waves: the
effects of whitewater parks on fisheries. Sustaining Colorado Watersheds Conference,
Avon, Colorado. October 10, 2018.
Schlosser, I. J., and P. L. Angermeier. 1995. Spatial variation in demographic processes for
lotic fishes: conceptual models, empirical evidence, and implications for conservation.
American Fisheries Society Symposium 17:392-401.
Stephens, T. A., B. P. Bledsoe, B. D. Fox, E. Kolden, and M. C. Kondratieff. 2015. Effects of
whitewater parks on fish passage: a spatially explicit hydraulic analysis. Ecological
Engineering 83: 305–318.
Swarr, T.R. 2018. Improving rock ramp fishways for small-bodied Great Plains fishes. M.S.
Thesis, Department of Fish, Wildlife, and Conservation Biology, Colorado State University.
89 pp.
Thompson, K.G., and Z.E. Hooley-Underwood. 2019. Present distribution of three Colorado
River Basin native non-game fishes, and their use of tributary streams. Colorado Parks and
Wildlife Technical Publication 52.

16

�Appendix A
CPW Aquatic Biologist Contact Information

�MOFFAT
a
Ya mp

River

Tory Eyre
Meeker
(970)878-6074

JACKSON

LARIMER

Steamboat
Springs Office

ROUTT

Meeker
Office

GRAND

70

EAGLE

mp
Unco
e River
a hgr

n

el
gu
Mi

Ri
ve

SAN MIGUEL

r

OURAY

TELLER

SW

Jim White
Durango
(970)375-6712

Ri o Grande Ri v

CUSTER

er

RIO Monte
GRANDE Vista

i ve
Conejos R

OTERO

HUERFANO

COSTILLA
i
Purgatoire R

Miles
25

50

LAS ANIMAS

25

AQUATIC BIOLOGISTS
COLORADO PARKS AND WILDLIFE
0

KIT
CARSON

Smoky H
il

CHEYENNE

l River

75

100

NATIVE AQUATIC SPECIES
BIOLOGISTS
NORTHEAST REGION
Boyd Wright
(970)472-4366

BENT

PUEBLO

o
rf an
Hue

r

NORTHWEST REGION
Lori Martin
(970)255-6186

SOUTHEAST REGION
Josh Nehring
(719)227-5224

r
i ve

Lamar Office

PROWERS

Jim Ramsay
Lamar
(719)336-6607

SE

mo s
aR
i ve
r

NORTHEAST REGION
Jeff Spohn
(303)981-3634

KIOWA

Carrie Tucker
Pueblo
(719)561-5312

ALAMOSA

CONEJOS

ARCHULETA

nR

LINCOLN

Office
Ala

Durango
Office
Durango Regional LA PLATA
Administrative Office

Cory Noble
Colorado Springs
(719)227-5222

ub

a
lic

CROWLEY

Estevan Vigil
Monte Vista
(719)587-6908
MI NERAL

t
S ou

-Lake Pueblo
-Pueblo Office

SAGUACHE

SAN
JUAN

DOLORES

EL PASO

iver
ree R

p
Re
ork
hF

SENIOR AQUATIC BIOLOGISTS

SOUTHWEST REGION
John Alves
(970)375-6721

ARA PAHOE

ELBERT

DOUG LAS

Dan Brauch
Gunnison
(970)641-7070
HINSDALE

WASHINGTON

70

r
iv e

i ver
es R
l or

Sa

YUMA

Mike Atwood
Salida
(719)530-5525FREMONT

Salida
Office

Gunnison
Office

G unn
ison Ri ver

NE

Brush
Office

a
Arik

Littleton Administrative Office

as R
ans

Do

MONTROSE

MONTEZUMA

JEFFERSON

CHAFFEE

Montrose
Office

76

Paul Winkle
Denver
(303)291-7232

Colorado Springs
Office

GUNNISON

Eric Gardunio
Montrose
(970)252-6017

er
Ri v

Headquarters

A rk

DELTA

th Pla tt e

ADAMS

PARK

Grand
Junction
Office

S ou

PHILLIPS

Mandi Brandt
Brush
(970)842-6330

Denver Administrative Office
Northeast Regional Office

Tyler Swarr
South Park
(720)576-9782

LAKE

PITKIN

MESA

CLEAR
CREEK

S UMMIT

Kendall Bakich
Glenwood Springs
(970)947-2924

MORGAN

BOULDER
GILPIN

e River
Blu

GARFIELD

Ben Felt
Grand Junction
(970)255-6126

Glenwood
Springs
Office

Fort Collins
Office

Ben Swigle
Fort Collins
(970)472-4364

Hot Sulphur
Springs
Office

r
o Rive
Colorad

LOGAN

WELD

Thompson Riv er
Bi g

Jon Ewert
Hot Sulphur
(970)725-6214

r
e Rive

RIO BLANCO

25

Poudre River

ve
r

Whit

er

Ri

en
River

Bill Atkinson
Steamboat
Springs
(970)870-2868

SEDGWICK

Kyle Battige
Fort Collins
(970)472-4396

ian

Gr e

d
na
Ca

NW

Ri v

NORTHWEST REGION
Jenn Logan
(970)947-2923
SOUTHWEST REGION
Dan Cammack
(970)275-9617
SOUTHEAST REGION
Paul Foutz
(719)227-5217

BACA

r
ve

CPW Region Boundary
Created by CPW GIS 2/12/2020
314 W. Prospect St
Fort Collins, CO 80526
G:\Projects\Publications\Boundaries\AquaticBoundaries\CPW_AquaticBiologists_2020_11x17.mxd

�Appendix B
CPW Whitewater Park Fact Sheet

�C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Whitewater Park Studies
RESEARCH RESULTS AND DESIGN GUIDELINES

Whitewater Park Research
With over 30 whitewater parks (WWPs) either completed or in the
planning phases, Colorado is the epicenter for WWP development in the
United States. Although WWPs provide economic and recreational
benefits for local communities (Hagenstad et al. 2000; Loomis and
McTernan 2011), they may have unintended impacts on instream biota
and stream functions, particularly when the hydraulic conditions formed
by the WWP are different from those naturally found in the surrounding
river. The impact of WWPs on habitat connectivity and instream habitat
quality have been the focus of several recent studies. Although these
studies have primarily focused on fish passage and habitat, impacts to
aquatic insects and sediment transport may also occur at WWPs.

Fish Passage Impacts
The elements that create a desirable surf wave (increased velocity,
decreased depth, a hydraulic jump, and a stable, often grouted stream
channel) create conditions that can impede fish movement. Swimming
speeds and jumping ability vary greatly between fish species.
Suppression of upstream trout movement has been documented at WWP
structures, but the degree of impact varied by fish size and characteristics
of the individual structure (Stephens et al. 2015; Fox et al. 2016). As trout
are among the strongest swimming and jumping fish species in Colorado,
small-bodied and weaker-swimming fish native to Colorado streams are
even more susceptible to habitat fragmentation associated with WWP
development.

Brown Trout

Mottled Sculpin

Fish Habitat Impacts
Although WWPs create deep pools, observed fish densities were significantly higher in natural pools than in WWP pools
(Kolden et al. 2015; Kondratieff et al. in preparation). Habitat degradation in WWPs was associated with the unnatural
hydraulics created by the recreational features and conversion of riffle habitat to drops over the wave structures.

Design Guidelines
CPW recommends that adequate environmental safeguards be included in the design and construction of WWPs to ensure
that stream functions, fisheries, and recreational fishing are not adversely impacted. Each structure must be examined on a
case-by-case basis, and monitoring and adaptive management should be included in the proposed project budget.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

�Site Selection




Design and construction of WWPs should preserve the
natural aesthetic qualities of the river. WWPs should be
located in degraded reaches when possible and should aim
to improve the natural functions of the reach rather than
maintain degraded conditions. WWPs should not be
constructed in natural, un-modified river channels
(American Whitewater 2007).
WWP sites should be selected to minimize recreational
conflicts with anglers. There is increased potential for
boaters to displace anglers at WWP sites, especially during
the summer months. If WWP construction affects a popular
fishing location, mitigation such as new fishing access or
habitat improvements should be considered.

Ecological Design Considerations







WWP structures must be designed to allow upstream fish passage for all life stages of native and sport fishes present
throughout the annual hydrologic cycle. Fish passage is dependent on water velocity, water depth, vertical height of
structures, linear distance of the passage corridor, surface roughness, and attraction flow.
Hydraulic characteristics at WWP features generally conflict with ideal conditions for fish passage. Therefore, a fish
passage channel separate from the WWP structure may be necessary. The passage channel should meet hydraulic design
criteria for target species across a range of flows.
Hydraulic modeling of the proposed structure should be conducted during the initial design phase to evaluate potential
impacts to fish passage and habitat.
Streambed and bank disturbance due to construction activities should be scheduled for a time of year when egg
incubation is not occurring. An increase in fine sediment to the stream during incubation can suffocate developing
embryos. Erosion control and revegetation plans utilizing native riparian species should be required for each project.
WWP structures should not cause sediment deposition upstream or downstream of the structure. Sediment deposition
can eliminate fish and benthic macroinvertebrate habitats, create favorable conditions for the spread of whirling disease
in trout, and increase flooding risk if sediment deposition decreases channel capacity.
Recreational In-channel Diversion (RICD) water rights can be acquired for WWPs to provide recreational experiences
in and on the water. These protected flows should be managed to benefit boating recreation as well as conservation and
management of native and sport fish. Flows deviating from the natural flow regime, such as water calls during spawning
periods, could have adverse impacts on stream ecology (Poff et al. 1997).

References
American Whitewater, 2007. Whitewater Parks – Considerations and Case Studies.
https://www.americanwhitewater.org/content/Wiki/stewardship:whitewater_parks
Fox, B. D., B. P. Bledsoe, E. Kolden, M. C. Kondratieff, and C. A. Myrick. 2016. Ecohydraulic evaluation of whitewater parks as a fish passage barrier. Journal of the American
Water Resources Association. DOI: 10.1111/1752-1688.12397.
Hagenstad, M., J. Henderson, R. S. Raucher, J. Whitcomb. 2000. Preliminary evaluation of
the beneficial value of waters diverted in the Clear Creek Whitewater Park in the City of
Golden. Stratus Consulting.
Kolden, E., B. D. Fox, B. P. Bledsoe, and M. C. Kondratieff. 2016. Modelling whitewater
park hydraulics and fish habitat in Colorado. River Research and Applications. DOI:
10.1002/rra.2931.
Kondratieff, M. C., K. Kinzli, and E. R. Fetherman. In preparation. Eco-hydraulic evaluation
of whitewater parks as fish habitat in Colorado.
Loomis, J., and J. McTernan. 2011. Fort Collins Whitewater Park economic assessment.
Department of Agricultural and Resource Economics, Colorado State University.
Poff, N. L., J. D. Allan, M. B. Bain, J. R. Karr, K. L. Prestegaard, B. D. Richter, R. E. Sparks,
and J. C. Stromberg. 1997. The natural flow regime: a paradigm for river conservation and
restoration. BioScience 47(11): 769-784.
Stephens, T. A., B. P. Bledsoe, B. D. Fox, E. Kolden, and M. C. Kondratieff. 2016. Effects
of whitewater parks on fish passage: a spatially explicit hydraulic analysis. Ecological
Engineering 83: 305–318.

�Appendix C
CPW Fish Passage at River Structures Fact Sheet

�C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Fish Passage at River Structures
RESEARCH AND DESIGN GUIDELINES

Introduction
Instream structures, such as culverts, water diversions and dams, can negatively affect fish by
fragmenting populations, reducing migratory ranges, and limiting access to habitat for spawning, feeding and refugia.
Many rivers in Colorado contain man-made structures that create partial (obstacles) or complete barriers depending on
the fish species and life stage. Habitat fragmentation associated with instream barriers is a serious threat to Colorado’s
Species of Greatest Conservation Need (SGCN) and sport
fisheries. Therefore, it is important that fisheries managers
(A)
identify and evaluate the influence of instream structures on
fish populations.

Fish Passage Research Objectives
The primary goal of fish passage research is to restore
connectivity in fragmented river systems by: (1) evaluating the
effectiveness of existing fishways; (2) evaluating the barrierpotential of common river structures; and (3) establishing fish
swim performance criteria for native and sport fishes.

Current Fish Passage Research Projects
Active fish passage research projects include: (1) evaluation of
native fish passage at existing fishways located on Front Range
transition zone streams; (2) evaluation of fish passage at
instream whitewater park structures; (3) laboratory studies to
develop fish swim and jump performance criteria for Colorado
fishes where data is lacking; and (4) development of new
techniques and technologies for investigating fish movement
and passage in rivers.

(B)

Fishway Design
Fishways, or “fish ladders”, are engineered structures
designed to facilitate passage around an obstacle or barrier.
Fishways attempt to incorporate species- and life stagespecific swimming and jumping abilities into designs. Common
elements of successful fishways include: (1) low velocity
pathways that do not exceed burst speeds or endurance
capabilities for target species (Figure A); (2) water depths that
do not limit swimming performance (Figure B); (3) vertical
drops that do not exceed the jumping ability for target species
- note that many species native to Colorado do not exhibit
jumping behaviors (Figure C); (4) sufficient attraction flow, or
the flow that emanates from a fishway entrance, to ensure
that fish can locate the fishway; and (5) maintenance of the
above design elements over the expected range of
streamflows.

(C)

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

�Fishway Examples
Some examples of successful fishways include engineered rock ramps (Figure D), constructed riffles (Figure E), and
vertical slot fishways (Figure F). Each type of fishway has advantages and disadvantages related to which fish species
and life stages are present and the conditions of the project site.

Engineered Rock Ramp

Constructed Riffle

Vertical Slot

Diversion Crest

Piney Creek,
Wyoming

Fossil Creek Reservoir
Inlet Diversion,
Cache la Poudre River

(D)

Rock Weirs

CCC Ditch,
San Miguel River

(E)

(F)

Aquatic Habitat Types
From the high-gradient, boulder-dominated, step-pool
channels of snowmelt fed mountain streams to the lowgradient, well-vegetated, pool-riffle rivers of the eastern
plains to the majestic, vertically-confined canyons on the
arid Colorado Plateau, aquatic habitats in Colorado are as
diverse as the geographic regions where they are found.
Native Colorado fishes have unique morphological
characteristics that are adapted to the natural conditions
found in each aquatic habitat type. These adaptations affect
the swimming abilities of fish, influencing how they move
through and use diverse habitats. Fisheries managers must
take the diversity of fish species into consideration when
evaluating river structures and designing fishways.

Fish Swimming Performance by Family
Family Name
Percidae (Perches)

SGCN (#)

Fundulidae (Topminnows)
Cottidae (Sculpin)
Ictaluridae (Catfish)
Cyprinidae (Minnows)
Catostomidae (Suckers)
Centrarchidae (Sunfish)

All illustrations of fish © Joseph R. Tomelleri

3

Prolonged Speed (ft/s)
0.4 - 1.2

Burst Speed (ft/s)
NA - 2.4

Jump Height (ft)
0*

Habitat Types
EP

1
0
1
13
5
1

1.3 - 1.6
1.4 - 1.7
1.3 - 2.0
1.3 - 2.4
1.3 - 2.5
1.1 - 2.9

2.6 - 3.4
3.3 - 3.9
2.0 - NA
2.4 - 4.4
2.2 - 3.2
2.6 - NA

0.1 - 0.2
0*
NA - 0.2
0* - 0.5
NA - 0.8
0.4 - NA

EP
CP, MS
EP, TZ
CP, EP, MS, RG, TZ
CP, EP, MS, RG, TZ
EP

Salmonidae (Trout)
3
2.3 - 4.0
4.5 - 7.5
1.0 - 7.0
MS, RG, TZ
SGCN = Species of Greatest Conservation Need, # of species/subspecies; * = fish species does not exhibit jumping behavior; NA =
data were not available; CP = Colorado Plateau, EP = Eastern Plains, MS = Mountain Streams, RG = Rio Grande; TZ = Transition Zone

The values reported above are summarized from multiple species within each family and are intended to support passage
for juvenile life stages. Swim speeds and jumping abilities within species are size dependent. Species-specific performance
criteria should be used whenever possible. The selection of target species for individual projects should be based on the
management objectives for the site in question. Consultation with the local Area Aquatic Biologist at CPW is strongly
encouraged during the early planning stages for any fish passage project in Colorado. The information in this fact sheet is
based on the best available data and knowledge, but is subject to revision as more information becomes available.
COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

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                <text>Colorado Parks and Wildlife’s (CPW) statutory mission is to perpetuate the wildlife resources of the State, to provide a quality State Parks system, and to provide enjoyable and sustainable outdoor recreation opportunities that educate and inspire current and future generations to serve as strategic stewards of Colorado’s natural resources (C.R.S. § 33-9-101 (12) (b)). As CPW is responsible for the management and conservation of aquatic resources within the State, we are asked to review projects that may affect aquatic habitats or populations. Specifically, CPW staff is often engaged by the Army Corps of Engineers (USACE) to review permit applications related to the design, construction, and monitoring of whitewater parks (WWPs) regulated under Section 404 of the Clean Water Act. WWP projects typically fall under the following permits: &lt;br /&gt;
&lt;ul&gt;
&lt;li&gt;NWP 27 - Aquatic Habitat Restoration, Establishment, and Enhancement Activities&lt;/li&gt;
&lt;li&gt;IP - An individual, or standard permit, is issued when projects have more than minimal individual or cumulative impacts, are evaluated using additional environmental criteria, and involve a more comprehensive public interest review.&lt;/li&gt;
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                <text>Kondratieff, M. C., K. R. Bakich, E. E. Richer, D. A. Kowalski, and B. F. Atkinson. 2020. Whitewater Park Projects: Guidance for Reviewing 404 Projects. Colorado Parks and Wildlife Aquatic Research Section, Fort Collins, CO. 26 pp.</text>
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                    <text>American Dippers
American dippers are dark gray and have long legs.
They live by clear flowing streams. They are the only
songbirds that catch their food underwater! They eat insects

Illustration by Helen Zane Jensen ©2008

that live in streams. These birds earned their name because
they are always "dipping" their bodies up and down.

Wildlife Discovery Page-American Dipper/Elementary

http://cpw.state.co.us/learn/Pages/TR-WildlifeDiscovery.aspx

�</text>
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                    <text>American Dippers
American dippers are medium sized, gray birds that have long
legs and constantly bob their bodies up and down. Dippers are
America 's only true aquatic songbirds. They live along clear moving
waters. Dippers eat insects that live in these waters. They are yearround residents in Colorado and migrate only as far as the closest
unfrozen stream or river.

American dippers breed twice a year. Males and females often
rejoin and reuse the same nest. Chicks stay in the nest for about 24
days. When they leave, they can already swim and dive to catch food.
Dippers can hold their breath underwater for up to 30 seconds! These
birds have a heavy layer of waterproof feathers and can withstand
even the coldest waters. In the late summer, American dippers molt
theirs wing and tail feathers together. This leaves the birds flightless
for a short time.

e,o

"ou._,oO

Illustration by Helen Zane Jensen ©2008

----

Wildlife Discovery Page-American Dipper/Middle School

http://cpw.state.co.us/learn/Pages/TR-WildlifeDiscovery.aspx

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                    <text>Illustration by Helen Zane Jensen ©2008

Wildlife Discovery Page-American Dipper/Primary

http://cpw.state.co.us/learn/Pages/TR-WildlifeDiscovery.aspx

�</text>
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                    <text>C O L O R A D O

P A R K S

Adventures as a
Young Naturalist

Come on an outdoor journey
discovering Colorado’s
state parks.

&amp;

W I L D L I F E

�Welcome to Colorado’s state parks. There are over 40 state parks across Colorado
and each one is a special place to explore. Complete these fun activities, taking
your first steps to become a naturalist—a person who loves and cares about the
outdoors. Just remember to explore with an adult, have fun, and enjoy nature—it
is full of surprises.

We live in a world of
many colors. You can see
rainbows of color when
light bounces off a drop of
water or a wet summer sky.

Find this feather to
experience more
exciting adventures!

�There is a magical
mini-world of
flowers, leaves,
pebbles, and critters
under your feet.

�grub

great horned owl

fox squirrel

raccoon

black bear

These animals are living in their
natural home. Draw a line from the
picture of the larger animal to its
matching animal. Then color
the animals.

�bee

red-tailed
hawk

woodpecker

grasshopper

Trees make great homes for all kinds of animals. They provide food and cover. Walk
around a tree. Look up in the leaves. Feel and smell the bark. Explore the ground
underneath. Can you spot any signs that animals live in the tree? Here is a clue—look for
holes, scratches, or nests.

�Use the number guide to
color this mallard duck.
1 – blue 4 – yellow
2 – red
5 – white
3 – green

1
5
3

4

1

2

5

4

5

4

5

1
2

15

3
4

14
5
10

13

6
7

9
11

12

8

Follow the numbers to trace the
outline of the fish. Count all the spots
and write the number here
.
Mallard ducks have webbed feet for swimming. Their bills
strain plants and insects from the water. Colors and spots on
fish help them hide. Find an animal in the park and watch it
quietly. What is it doing?

�Use this guide to color these
shapes found in the butterfly’s wing.
red
green

blue
yellow

�.
.
.
e
r
a
es
i
r
e
v
o
sc
i
d
e
t
i
or
v
a
f
y
M

!

Funded in part through
Great Outdoors Colorado with Colorado Lottery proceeds.

Acknowledgements:

1313 Sherman Street, #618
Denver, CO 80203
303.866.3437
cpw.state.co.us

EDU1408_4/18

ming
o
c
e
b
s,
ps in over more
e
n
t
o
s
i
t
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s
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r
isc
r fi
atu
Congr e taken you continue to d fe, have fun
v
a
You ha alist! As you ber to stay s !
s
r
a natu ature, remem e more park
r
n
about rs, and explo
outdoo

Design Team: Sandy Berg, Roxanne Brickell-Reardon, Deb Duke, Linda Hamilton, Matt Huerter, Faye Koeltzow, Debbie Matlock, Mary McCormac, Sara Melena, Sheryl
Radovich, Angel Tobin • Author: Sheryl Radovich • Artist: Paul Gray • Design: Deb Duke • Reviewers: Roxanne Brickell-Reardon, Deb Duke, Linda Hamilton, Matt Huerter, Faye
Koeltzow, Debbie Matlock, Mary McCormac, Sara Melena, Sheryl Radovich, Kathy Seiple, Angel Tobin, Mark Young • Technical Reviewers: Nicole Bickford, Mary Ann Bonnell •
Project Managers: Roxanne Brickell-Reardon, Debbie Matlock
• Editor: Lynn Almer • Original Idea: Todd Farrow • Funding: Great Outdoors Colorado

�</text>
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                    <text>The Tassel-Eared Squirrel
The Abert's squirrel has long tufts or tassels of fur
on its ears. The tassels are longest in winter. In summer,
the squirrel's ear tassels
may be smaller or
they may disappear.

r
-

.

~--

:.--

How can you tell if a squirrel is an Abert's squirrel in
summer if it loses its ear tufts? The Abert's squirrel lives
in only one place-the ponderosa pine forest. It builds its
nest in the ponderosa pine tree and eats seeds from the
pine cones!

;-

::;-

"'

,. ,:

,

'~
_,,
\\ I·~';&gt;

Illustration by Helen Zane Jensen ©2008

Wildlife Discovery Page-Abert’s Squirrel/Elementary

http://cpw.state.co.us/learn/Pages/TR-WildlifeDiscovery.aspx

�</text>
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                    <text>The Squirrel and the Tree
Abert's squirrels live in only one habitat-the ponderosa pine
forest. These squirrels depend on ponderosa pine for the essentials
of life. Abert's squirrels build nests high up in these trees. The nests
look similar to a large, messy bird nest. They either construct a
ball-like mass of twigs from pine or build their nests within "witches'brooms," growths of small pine twigs infected by dwarf mistletoe.
Since Abert's squirrels do not hibernate, they use their nests yearround. They sleep in the nest at night and use it to hide from
predators during the day.

Most of the Abert's squirrels' diet is made up of parts of the
ponderosa pine. In warm months, it eats the tree's buds and cones.
They turn the pine cone slowly, like the way people eat corn on the
cob, peeling away the cone scales to reach the meaty seeds. Often,
they eat their cones in a favorite spot on a branch in the tree. A pile
of cone scales under a ponderosa pine is a sure sign that Abert's
squirrels live there. In the winter the squirrels eat
the inner bark of the tree. They also eat mistletoe
and fungi growing on the tree.
Abert's squirrels clearly benefit from
the trees. What do the ponderosa pines
get from this relationship? When Abert's
squirrels dig up and eat a certain fungus
growing around the roots of the treeEctomychorrhizal fungus-they disperse
spores. This helps the fungus reproduce.
The fungus benefits Ponderosa pines by
growing around the roots of the trees
and helping the trees to maintain
moisture in a dry environment.

Illustration by Helen Zane Jensen ©2008

Wildlife Discovery Page-Abert’s Squirrel/Middle School

http://cpw.state.co.us/learn/Pages/TR-WildlifeDiscovery.aspx

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