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                    <text>C O L O R A D O

P A R K S

A N D

W I L D L I F E

2012 Colorado Angler Survey
Fact Sheet

Fishing Regulations
Review
Purpose of the study: To determine the spatial distribution of angling pressure for various species, across
Colorado and collect data to help predict demand for sport fish species around the state over the next 30 years.
Methods: Between March and August 2012, we mailed a survey to 3,000 individuals who had purchased
Colorado fishing licenses in 2011. Half were residents (5% senior, 12% combination, 50% annual and 33%
1-day licenses) and half were non-residents (76% annual and 24% 1- or 5-day licenses). We had a valid
sample of 2,577 anglers and received a total of 1,404 responses to the survey (54% adjusted response rate;
56% of residents and 44% of non-residents).
Highlights of results: Key findings of this survey were:
 Among anglers of all types, the 3 most commonly targeted species were rainbow trout, brown trout
and cutthroat trout. All anglers desired an increase in opportunity for these species over other species.
 Frequent and moderate anglers were more satisfied with their fishing experience in Colorado than
infrequent anglers. Moderate and frequent anglers were very satisfied with the variety of fish and
waters they could fish in Colorado. Infrequent anglers were not satisfied with the number and size of
fish they caught and the complexity of the fishing regulations in Colorado.
 Annual, resident license purchasers fished for an average of 24 days in 2011; annual, non-resident
license purchasers fished for an average of 17 days. Senior license purchasers fished for an average of
13 days and combination license purchasers fished for an average of 15 days.
 Resident anglers were very satisfied with the variety of fish and water available to them to fish, and
their ability to eat the fish they caught. Residents reported the highest level of dissatisfaction with the
number and size of fish they caught and the availability of wild fish.
 Non-residents were satisfied with the variety of fish and water available to them, and their ability to
eat fish they caught. Non-residents were more satisfied with all aspects of fishing than residents.
 Colorado anglers spent the highest mean number of days fishing for carp, wiper and striped bass, and
bluegill, but these were the least commonly targeted species. The three most commonly targeted
species (brown trout, cutthroat trout and rainbow trout) averaged the fewest days fished.
 The Colorado population between 51 and 70 years old will decrease over the next 30 years and the
population between 71 and 90 years old will increase. This change will not likely affect demand for
species or types of access, but may increase participating in fly angling and demand for trout species.
 Demographers predict that the white, non-Hispanic population in Colorado will decrease, while the
Hispanic population will increase in the next 30 years. White and Hispanic anglers prefer similar
species to target and locations in which to fish, but Hispanic anglers are much more likely to keep
their catch, which may lead to increased harvest pressure on trout stocks in Colorado in the future.
 Colorado is predicted to attract immigrants from California, Texas and Arizona in the next 10 to 20
years. This type of in-migration is unlikely to change demand for species or access in an appreciable
way. There may be a slight increase in demand for warm-water game fish such as walleye, but this is
likely to be overshadowed by desire for cold-water species found more commonly in Colorado.
For more information: For more information about this and other Human Dimensions surveys, please
contact Stacy Lischka at stacy.lischka@state.co.us or (303) 291-7279.
COLORADO PARKS AND WILDLIFE ▪ 6060 Broadway, Denver, CO, 80216 ▪ (303) 297-1192 ▪ www.wildlife.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

ANS Boat Interdictions

Zebra and Quagga Mussels

Invasive mussels are being transported on watercraft!
Colorado Parks and Wildlife coordinates a broad multijurisdictional watercraft inspection and decontamination
network to protect waters from invasive zebra and quagga
mussels and other invasive species. Recreational watercraft
is the main vector of introduction for this harmful invader.
Colorado is a headwater state and there are no mussels
upstream. State certified inspectors repeatedly intercept
watercraft infested with mussels.
A total of 281 boats with attached adult zebra or quagga
mussels have been intercepted coming into Colorado’s
waters from out of state at watercraft inspection and

decontamination stations. The infested watercraft were
intercepted at Barr Lake, Blue Mesa, Boulder Marine, Boulder
Reservoir, Boyd Lake, Canon Marine, Carter, Cherry Creek,
Chatfield, Clear Creek, Crawford, Denver CPW Office,
Dillon, Electra Lake, Eleven Mile, Frisco Bay Marina, Granby,
Grand Lake, Grand Junction CPW Office, Great Lakes
Marine, Green Mountain, Highline, Horsetooth, Jackson,
John Martin, Lathrop, McPhee, Navajo, North Sterling,
Pueblo, Ridgway, Rifle Gap, Roadside (SW Colorado),
Ruedi, Shadow Mountain, Spinney Mountain, Stagecoach,
Steamboat Lake, Strontia Springs, Sweitzer, Taylor Park,
Trinidad, Turquoise, Vallecito and Williams Fork.

Zebra and Quagga Mussel Positive Interdiction Sites in Colorado

cpw.state.co.us

�ZEBRA MUSSELS

Where are the infested mussel boats coming from?
It is often difficult to determine the exact location of infestation
due to the frequency of boating use. Also, numerous
interceptions are used boat purchases in which the previous
boating history is not known. For these reasons, the source
of infestation for more than twenty interdicted watercraft is

unknown. The remainder of the infested vessels were coming
from Arizona, Arkansas, California, Illinois, Indiana, Iowa,
Kansas, Michigan, Minnesota, Mississippi, Missouri, New York,
Nevada (Lake Mead), Oklahoma, Ohio, Pennsylvania, Texas,
Utah (Lake Powell), Wisconsin, and the Great Lakes.

Boat Origin for Zebra and Quagga Mussel Positive Interdictions in Colorado

COLORADO PARKS &amp; WILDLIFE

6060 Broadway, Denver, CO 80216

(303) 291-7295 • (303) 297-1192

cpw.state.co.us
1/2020

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

ANS Fact Sheet

AQUATIC NUISANCE SPECIES (ANS) Program OVERVIEW

Background

The State Aquatic Nuisance Species (ANS) Act was signed into law
May 2008. The Act defines ANS as exotic or nonnative aquatic wildlife
or any plant species that have been determined to pose a significant
threat to the aquatic resources or water infrastructure of the state.
The Parks Board passed regulations required by the Act on February
20, 2009 and updated them since that time. The regulations require
mandatory watercraft inspection, and if necessary, decontamination
of all boats coming from out of state, leaving waters with known ANS
and boats entering high-risk waters where inspections are required
by the managing entity. The focus of the program is to prevent zebra
and quagga mussels and other ANS from infesting Colorado’s water
resources and threatening our water storage and distribution systems
for municipal, industrial and agricultural use. The Colorado ANS
Program is highly effective and a model which other states across the
nation are learning from.

Program Goal and Successes

The goal of the program is to protect the state’s natural resources,
outdoor recreation and water supply infrastructure through the
prevention of new introductions of costly invasive species, such as
zebra or quagga mussels, in Colorado. Western states such as Arizona,
Kansas, Nebraska, Oklahoma, South Dakota, and Texas, do not
have mandatory ANS prevention programs and continue to become
infested with zebra or quagga
mussels. Colorado has prevented
the introduction of these awful
invasive species due to the diligent
efforts of watercraft inspection
and decontamination staff, as well
as monitoring, education and
enforcement actions.
There has never been an adult
zebra or quagga mussel found in
Colorado. Pueblo Reservoir was
de-listed in January 2017 following
5 years of negative detections. All
other reservoirs that initially tested
positive in 2008 were de-listed
in 2014 after 5 years of negative
testing. CPW’s ANS Program
has worked to stop the continued
inoculation of our waters to
invasive mussels being introduced
by recreational watercraft.

Sampling and Monitoring

Sampling and monitoring is a key component to the success of the ANS
Program. CPW has sampled 584 “at-risk” waters over the last decade and
it was through this sampling program that invasive mussel larvae were first
detected in Colorado. While CPW ANS staff monitors the state’s public
waters for numerous invasive plant and animal species, the focus of sampling
is on early detection of zebra and quagga mussels. There are three sampling
protocols that target the three life cycles of mussels. CPW also documents
native aquatic plants, mollusks and crayfish while performing monitoring
activities for invasive species.

Colora

d o Pa r
ks &amp; Wi
ldli

Protect Colorad
o’s
Lands and Wate
rs
Learn How You

Can Stop InvaSIve

fe

SpeCIeS

Information and Outreach

CPW and its partner agencies have
implemented a comprehensive, multifaceted invasive species public education
campaign. Accomplishments include
distribution of tens of thousands of
printed rack cards, brochures, handouts,
DVDs and posted signage at offices,
boat ramps and other public access
points. Additionally, a media relations campaign has been
launched using web, radio, print and television.

www.cpw.state.co.u

s

Working Together: Watercraft Inspection
and Decontamination (WID)
Watercraft inspection and decontamination is a requirement of the
ANS Act and continues to be a key component in preventing the spread
of ANS into and within
Colorado. CPW coordinates the
vast network of WID stations
that are operated by CPW, the
National Park Service, Larimer
County, various municipalities
and private industry locations.
In total, the state has collectively
performed over 4.9 million
inspections and 119,814
decontaminations since 2008.

cpw.state.co.us

�2018 Water Resources Development Act

The 2018 Water Resources Development Act (S. 3021) was passed by the 115th
Congress and signed into law last October. Section 1170 includes a provision
which directs the U.S. Army Corps of Engineers to establish, operate, and
maintain new or existing watercraft inspection stations to prevent the spread
of aquatic invasive species in the Columbia, Upper Missouri, Upper Colorado,
South Platte and Arizona (should be Arkansas) River Basins. The provision also
authorizes the Army Corps to assist states with monitoring and rapid response
efforts in the case of an infestation of quagga or zebra mussels.

Green Mountain Reservoir

In August 2017, the U.S. Bureau of Reclamation (BOR) detected quagga mussels
in Green Mountain Reservoir. The specimen were genetically confirmed to be
quaggas by an independent lab for CPW. Rapid Response was initiated by CPW
and began immediately among the multi-jurisdictional partnership. Green
Mountain is now listed as “suspect” for quagga mussels and containment is
ongoing. Resources are needed for infrastructure, security and staffing to prevent
the spread through watercraft. Green Mountain Reservoir within Summit
County is owned by the BOR as part of the Upper Colorado River Collection
System, and is managed for recreation by the White River National Forest.
There were no detections in either 2018 or 2019. If there are no detections in
2020, the reservoir will be de-listed in 2021 per regional standards.

Mussel Boat Interceptions

Infested mussel boat interceptions continue to skyrocket each year. In total
there have been 281 watercraft intercepted with adult zebra or quagga mussels
attached since the ANS program began. In 2019, CPW intercepted 86 watercraft
infested with zebra or quagga mussels coming in from out of state. In 2018, the
state intercepted 51 and in 2017 the state intercepted 26 infested watercraft.
This exponential growth in infested boat interceptions is directly related to the
growing threat invasive species pose to water infrastructure, natural resources
and outdoor recreation. The majority of the intercepted vessels were coming
from Lake Powell, the State of Arizona or the Great Lakes. All boats were
decontaminated to ensure all mussels were dead, and no mussels were visibly
attached.

An administrative solution in both the House and Senate is needed to correct
the typo is Section 1170 from the nonexistent Arizona River Basin to the Senate’s
intent of providing resources to the Arizona River Basin with its headwaters in
Colorado. Urge the Army Corp to not delay appropriations or planning in the
Upper Colorado or South Platte Basins while the Arizona/Arkansas Basin typo
is corrected.
Support is needed to gain the fiscal appropriations as authorized in the bill.

The Stop the Spread of Invasive Mussels Act of 2019

Senator Bennet, along with Senator Daines and Senator Tester from
Montana, introduced the Stop the Spread of Invasive Mussels Act of 2019.
The bill authorizes the Secretary of Interior through the Commissioner of
Reclamation to provide financial assistance to states or local governments
for WID. The bill adds several federal agencies, such as the National Park
Service, as a member of the ANS Task Force and provides authorization for
all members of the ANS Task Force to conduct WID into and out of federal
waters. The bill requires the ANS Task Force to report back on any regulatory
changes necessary to implement WID prevention or containment. Finally,
the bill corrects the typo in Section 1170 of S.3021 to authorize the Arkansas
River Basin. Sponsorship and support is needed to pass this bill.

The threat of invasion from
zebra and quagga mussels
is greater than ever due to
numerous new infestations
in surrounding states such as
Arizona, Kansas, Nebraska,
North Dakota, Oklahoma,
South Dakota, Texas and Utah.

Operating and Financials

The Supreme Court ruling in case 13SC996 significantly reduced the source fund
for the ANS Program (Tier II Severance Tax) as appropriated in the ANS Act.
A broad stakeholder effort was called upon to raise funds and determine long
term solutions.
CPW redirected agency cash and a USFWS Motorboat Colorado Grant to pay
for the 2017 boating season, along with over $1M of partnership funds. The
Colorado General Assembly provided funding to the ANS Program through
SB17-259 and HB18-1338 to be leveraged with federal dollars and other sources
to maintain the program over the 2018–2019 boating seasons.
In 2018, the Colorado General Assembly passed the Mussel Free Colorado Act
(HB18-1008) to provide a new stable funding source through the creation of an
ANS stamp for motorized boats and sailboats—$25 for in state residents and
$50 for out of state visitors. The ANS Stamp will provide approximately $2.4M
(50% of the need) annual funding to the ANS Program. The bill also increases
fines for violations and allows CPW to recoup costs for decontamination of
quarantined and impounded vessels.
Similar to the 2017 joint resolution on ANS which passed the State Legislature
unanimously, the Mussel Free Colorado Act continues to encourage federal
governments, specifically the Bureau of Reclamation, U.S. Forest Service and
Army Corp of Engineers, to provide the other half of the needed funds for the
ANS Program since many of the highest risk waters are federally owned or
managed reservoirs.

How You Can Help

• Contact U.S. Senators and Representatives, and the U.S. Army Corp
of Engineers, and encourage them to secure an administrative solution to
correct the typo in Section 1170 of S. 3021 (WRDA-2018) changing Arizona
River Basin, which does not exist, to Arkansas River Basin, which was the
intent of the Senate.
• Encourage Congressional delegates and the Army Corp of Engineers to
provide fiscal appropriations under Section 1170 of S. 3021 for the Upper
Colorado, South Platte and Arkansas River Basins. This funding will be used
to provide cost share for WID staff, operations, infrastructure improvements,
monitoring and rapid response.
• Support the passage of The Stop the Spread of Invasive Mussels Act of 2019.
• Urge the U.S. Bureau of Reclamation and the U.S. Forest Service to provide
funding to CPW for WID staff, operations and improved infrastructure on
waters under their ownership or recreational management.
• Inform municipal government, county commissioners, state legislators and
federal congressional delegates of
the potential economic and social
impacts that could occur without
ANS preventative measures.

COLORADO PARKS &amp; WILDLIFE       6060 Broadway, Denver, CO 80216       (303) 291-7295  •  (303) 297-1192       cpw.state.co.us
1/2020

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Aquatic Database
AQUATIC DATA MANAGEMENT SYSTEM (ADAMAS)

Colorado Parks and Wildlife’s Aquatic Database
Colorado Parks and Wildlife (CPW) developed a single, centralized
computer database - the Aquatic Data Management System
(ADAMAS) - to provide current and historic scientific data on the
status and trends of the state’s fisheries. ADAMAS, which is updated
throughout the year, contains information on over 13,000 lakes and
stream segments, and all fish managed by CPW for anglers. The
objective was to create a single, quick, reliable and easy-to-access
database to help CPW managers enhance angling opportunities and
provide information for aquatic scientists and other government
agencies researching topics that could benefit Colorado anglers
across the state.
The actual database is housed on a server in Denver. However the
majority of work on the database is conducted by the data analyst
within the Aquatic Research Unit at the CPW Fort Collins Research
Center.
CPW fish biologists and other fishery professionals - including
federal agencies, academics and consultants – continually
contribute data to ADAMAS. The database holds all the raw data
collected annually from surveys across the state and allows CPW
biologists, fishery managers and researchers to monitor changes to
effectively manage Colorado’s fish resources for anglers.
ADAMAS History
In the 1980’s, CPW began to standardize survey and data collection
methods, allowing for data from all over the state to be stored
together. A database would replace paper files, which typically were
lost when biologists retired. CPW leadership decided that a
statewide, centralized depository for the entire database would
benefit everyone.
The first rendition of the database, called the Stream and lake
Databank, was comprised of only related tables. It has continued to
evolve and develop into its present form. The database, now residing
on a centralized installation of Microsoft SQL DBMS, is made up of
hundreds of related tables. While new data is constantly being added
and older data is corrected, the actual database design and structure
are also constantly evolving to accommodate new data and
assessment techniques.
The database currently houses information on over 36,000 individual survey records and over 6 million individual fish, going
back all the way to 1875.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.u

�ADAMAS Users
 CPW biologists use it to monitor fish populations and manage angling opportunities.
 Consulting companies use it to complete federally required environmental assessments.
 Aquatic research scientists use it on a wide range of projects, from fish health, and water quality, to the effects of
climate change and other environmental issues related to fish.
 Water managers use it to inform water management plans and decisions
 The general public utilizes the Colorado Fishing Atlas, which links to the ADAMAS database for information on
stocking, species distributions and regulations.
 Scientists who make a formal data request to the data analyst
(who coordinates the review and approval of all requests and
compiles the data necessary.
ADAMAS Contributors
Eighteen area biologists, four senior biologists, four Aquatic Species
Conservation biologists and seven aquatic research scientists
contribute data to the database annually. Scientific Collection
Permit holders must also submit their data for inclusion into the
database in order for any subsequent permits to be processed. This
includes federal agencies, academic institutions and private
consultants.
ADAMAS Support
A data analyst within CPW's Aquatic Research Section manages
ADAMAS, updates and organizes the information, provides technical support for using the data, and coordinates the data
request process. The analyst also coordinates the development of new software and provides information to professional
fishery scientists focusing on fish health, adaption to climate change and other environmental factors.
ADAMAS Funding: A Sportfish Restoration Program Grant from the U.S. Fish and Wildlife Service and CPW.

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Blue Mesa Lake Trout
ACHIEVING LAKE TROUT AND KOKANEE FISHERY OBJECTIVES THROUGH LAKE TROUT HARVEST

Can trophy lake trout and kokanee be maintained
through increased harvest of small lake trout?
Kokanee fry survival has declined since lake trout began naturally reproducing within Blue Mesa Reservoir
beginning in the early 1990's. This decline was due to higher predation from the expanding and now larger
population of predatory-sized lake trout (Figure 1). Anglers value opportunities to fish for both kokanee and
lake trout. Kokanee provide the greatest draw for overall catch and harvest while lake trout provide the greatest
draw for their trophy potential. Lake trout over 50 pounds and 44 inches in length have been caught. However,
lake trout require plentiful kokanee as prey to achieve such large size (Figure 2). Improving kokanee fry
survival by reducing predation from small lake trout is necessary for maintaining abundant kokanee and
associated benefits to anglers seeking to catch and eat kokanee or seeking trophy lake trout.

Recent study: managing for coexistence of kokanee and
trophy lake trout
Unsustainable levels of predation by lake trout can lead to rapid declines in kokanee
abundance and in lake trout growth and body condition. Immediately following a 90%
decline in kokanee abundance during the 2000’s, CPW initiated fall netting efforts for
lake trout in 2009 to recover kokanee while still providing for a trophy lake trout
fishery. Additional harvest of lake trout is needed to achieve these management
objectives. Recent research demonstrated that increased removal of primarily small,
young lake trout could improve kokanee fry survival, the overall abundance of
kokanee, and produce more trophy lake trout. It is possible that this increased harvest
could be achieved through a lake trout harvest incentive program, but may require
additional netting if angler harvest is not sufficient.

Figure 1. Reconstructed population trajectory of predatorysized lake trout over time (years) based on abundance
estimates from netting (SPIN) and mark-recapture surveys.

A harvest incentive tournament can lead to improved angler
involvement and efficient fisheries management:
Fall netting by CPW supplemented angler harvest by removing ~1,200 lake trout
annually from 2009 through 2017 while returning trophy-sized lake trout to the water.
Angler harvest averaged ~6,000 lake trout annually over this period (Figure 3). As a
result of these joint efforts, the abundance of predatory-sized lake trout has decreased
recently (Figure 1) and kokanee survival and abundance has improved. However,
recent lake trout monitoring surveys indicate that numerous small lake trout are now
growing larger and expected to switch to preying on fish. CPW feels there is an
opportunity to further encourage angler harvest through a lake trout harvest incentive
program which could more efficiently remove enough small lake trout such that netting
would not be required. Ninety-six percent of lake trout harvested by anglers and netted
by CPW were smaller than 24 inches in length.

Figure 2. Average number of kokanee (KOK) per
predatory-sized lake trout (MAC) present in Blue Mesa
Reservoir over time (years) in relation to when state-record
fish were caught.

Lake trout harvest incentive tournament details:

 Only heads from lake trout less than 24 inches will be accepted.
 Tournament dates: February 1 st through July 31st, 2020.
 Prizes for most heads turned in: $1,000 for most, $500 second most, $250 third most.
 Prize for each tagged fish turned in: $250 (tags are not visible).
 Twenty $200 prizes randomly selected with one chance for each head turned in.
 Head drop off locations: Iola, Elk Creek, and Lake Fork Marinas; Gunnison and
Montrose CPW offices.
 Call (970) 641-7070 or email dan.brauch@state.co.us for more information.

Figure 3. Estimated total harvest of lake
trout (MAC) during different periods by
anglers over time (years) in relation to
CPW fall netting.

References:
Hansen, A.G., and D. Brauch. In review. Long-term population dynamics
of lake trout in Blue Mesa Reservoir, Colorado: guidance for an angler
harvest incentive program. CPW report.
Pate, W.M., B.M. Johnson, J.M. Lepak, and D. Brauch. 2014. Managing
for coexistence of kokanee and trophy lake trout in a montane reservoir.
North American Journal of Fisheries Management 34:908–922.

COLORADO PARKS &amp; WILDLIFE • 300 W. New York Ave., Gunnison, CO 81230 • (970) 641-7070 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Brown Trout Life History
EVALUATING BROWN TROUT POPULATION STRUCTURE AND LIFE HISTORY IN THE SOUTH PLATTE RIVER

Brown Trout Ecology
Brown Trout are native to Europe, Western Asia, and Northern
Africa, and have been introduced to every continent except
Antarctica. Across their native and introduced ranges, Brown
Trout show a large diversity in life-history traits. For example,
some Brown Trout stay within a small section of stream their
entire life. Some populations consist of fish that spawn only
once, while in other populations, fish spawn multiple times
over their lifetimes. There are also populations that are
anadromous, where fish move up freshwater rivers from the
ocean to spawn, as well as adfluvial populations, where fish
migrate between rivers and lakes. Along with these
differences in life history, the physical appearance of Brown
Trout varies greatly, which could be related to genetics,
habitat use, diet, and fish age, among other factors. However,
the connection between life-history traits, factors affecting
physical appearance, and management in North America is not
well studied.
Black (top), intermediate (middle), and red (bottom)
spotting patterns of Brown Trout from the Middle fork
of the South Platte River.

Brown Trout were first introduced to the United States in
1883, and subsequently introduced into Colorado in 1890.
Different source populations were used to establish these wild
Brown Trout fisheries, including lake populations from Scotland and river populations from Germany. Brown
trout populations in Colorado are generally considered a mix of these two, and exhibit physical and behavioral
characteristics of both as part of their life histories. Understanding Brown Trout life history is an important
aspect of managing these populations throughout the state.

Brown Trout in the South Platte River
The Brown Trout population in the Middle
Fork of the South Platte River near Hartsel is
of interest to both anglers and mangers due
to the unique angling opportunities provided
by both Spinney Mountain Reservoir and the
state wildlife areas along the South Platte
River. Between 2006 and 2011, a stream
restoration project was completed in a small
section of the Middle Fork of the South Platte
to create deeper pools for overwinter habitat
and to hold big fish throughout the rest of the
year. Between 2013 and 2016, CPW biologists
and researchers conducted a tagging study to
investigate habitat use by Brown Trout
throughout the river. Fish were tagged with
passive integrated transponder (PIT) tags containing a unique identification number that can be used to track
the location of fish through physical recaptures and scanning during population estimate, and by river-spanning
COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

�antennas placed in the river that can detect the tags when the fish move past their location. These antennas
record the date and time when movements are made year-round. Over the course of the study, three Brown
Trout spotting patterns were identified: those with only black spots, those with red spots, and an intermediate
spotting pattern between the two. To determine if spotting pattern explained observed movement patterns
or the life history of the Brown Trout population in the Middle Fork of the South Platte River, additional data
was collected in 2016, including stable isotopes, a technique used to identify chemical signatures in the fish
tissue and characterize their diets and where feeding occurred, scales for aging fish, and genetic samples.

South Platte River Brown Trout Population Structure and Life History
A total of 1,259 Brown Trout were PIT-tagged over the three year study. Brown Trout with typically black
spots and larger sizes moved longer distances, crossing all four of the stationary antennae at least once during
the study. Although some PIT-tagged fish made long-distance movements, most moved less than 9 miles, with
the majority of the fish moving 1 mile or less, suggesting many were river residents. Larger movements were
made in the fall (September through October) as well as the spring (March through May). Fall movements
occurred during the Brown Trout spawning period, whereas spring movements occurred when the ice was
melting. Brown Trout primarily used the section of river at the upstream end of the study area, and Spinney
Mountain Reservoir. When Brown Trout chose to move, they generally moved between these two locations
fairly quickly, bypassing the habitat in between, including the deep pools in the restoration section, to reach
favorable spawning areas or overwinter habitat in the reservoir.
Brown Trout with black spots were older than Brown
Trout with red spots, and fish with an intermediate
spotting pattern fell between the two, suggesting
spotting pattern changed as fish got older. Isotope
data indicated that black spotted fish and red spotted
fish each eat different foods and reside in different
parts of the system. Black spotted fish spend most of
their time within the reservoir, entering the river to
spawn in the fall, and the red spotted fish live
primarily within the river. Fish with an intermediate
spotting pattern are likely transitioning from life in
the river to life in the reservoir as they get older.
Genetic testing indicated that the Brown Trout with
black spots, red spots, and the intermediate spotting
Average age (95% confidence intervals) of Brown Trout with
pattern were related and form one adfluvial Brown
black, intermediate, and red spotting patterns from the Middle
Fork of the South Platte River.
Trout population. Brown Trout in the Middle Fork of
the South Platte River rely on both the river and the
reservoir to complete parts of their life history (e.g., spawning and juvenile rearing versus overwinter habitat
and growth), and change spotting patterns as the fish age and use these different habitats.

Management of Brown Trout
This study allowed biologists to better understand the life history of Brown Trout in the South Platte River.
Based on the suite of data collected, these Brown Trout represent one population with an adfluvial life history
incorporating characteristics of lake and river source populations historically used to establish Brown Trout in
Colorado and North America. The connection between the reservoir and the upstream spawning habitat is
important to the life history of the Brown Trout in this system, providing habitats needed to complete different
parts of their life cycle. This information will help inform management of the fish in both the Middle Fork of
the South Platte River and Spinney Mountain Reservoir, promoting their persistence, and ultimately providing
more angling opportunities in both systems. Additionally, knowledge of the life history can be used to guide
future river restoration activities in this and other rivers around the state to maximize their success.
Associated Literature

Avila, B. W., E. R. Fetherman, M. C. Kondratieff, E. E. Richer, D. A. Kowalski, M. R. Baerwald, A. Goodbla, and J. Spohn. In preparation.
Brown Trout population structure and life history in the South Platte River, Colorado.

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Dual Disease Resistance
in Rainbow Trout
EVAUATING RESISTANCE TO BACTERIAL COLDWATER DISEASE AND WHIRLING DISEASE IN COLORADO’S TROUT

Bacterial Coldwater Disease (BCWD)
Bacterial coldwater disease (BCWD) is caused by the
bacterium Flavobacterium psychrophilum. Found worldwide, BCWD
causes significant complications and death in hatchery trout
populations. Outbreaks typically occur at temperatures between 39
and 50F. Infected fish show a broad range of clinical disease signs
including lesions, spiral swimming, “black tail”, spinal deformities,
and pale or necrotic gills. Mortality can be high if left untreated, and
antibiotics are commonly used to treat BCWD. As an alternative, the
USDA National Center for Cool and Cold Water Aquaculture (NCCWA) developed a Rainbow Trout strain that is
resistant to F. psychrophilum. With the help of Utah Division of Wildlife Resources, psychrophilum-resistant
Rainbow Trout (PRR) were incorporated into the CPW hatchery system to help manage BCWD outbreaks.
Lesion caused by F. psychrophilum infection

Whirling Disease (WD)
Whirling disease (WD) is caused by the parasite Myxobolus cerebralis.
Spinal deformity, M. cerebralis infection
Signs of infection include skeletal deformities, “black tail”, and
“whirling” or spiral swimming. WD cannot be treated, and susceptible
fish typically die within their first year. M. cerebralis has a complex
multi-stage life cycle, making it extremely difficult to remove from
aquatic environments. One option for management is to use M.
cerebralis-resistant fish. The Hofer strain is genetically resistant to M.
cerebralis, however, it is domesticated and shows reduced survival in
the wild. To increase survival, CPW crossed the Hofer with wild
Rainbow Trout strains. The resulting crosses (HxC, Hofer by Colorado
River Rainbow; HxH, Hofer by Harrison Lake Rainbow) are resistant to M. cerebralis, and survive and reproduce
in the wild. Stocking M. cerebralis-resistant Rainbow Trout has helped reduce WD in aquatic systems
throughout Colorado.

Evaluating Dual Resistance via Dual Exposure to BCWD and WD
CPW uses the PRR to reduce mortality in the hatchery due to F. psychrophilum outbreaks. However, it is
unknown if the PRR are resistant to M. cerebralis. Stocking PRRs with no resistance to M. cerebralis could
result in high losses from WD, as well as increased M. cerebralis prevalence. Conversely, although resistant
to M. cerebralis, the HxH shows increased mortality in the hatchery during BCWD outbreaks. This study
examined if crossing the PRR with the HxH resulted in fish that are genetically resistant to both F.
psychrophilum and M. cerebralis.

Dual exposure laboratory, Colorado State University

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

�Injecting fish with F. psychrophilum

The PRR, HxH, and HHP, the first generation cross between the PRR and
HxH, were used for the dual exposure experiment. To test genetic
resistance to both pathogens, fish were exposed to F. psychrophilum, M.
cerebralis, or both. Fish were exposed to F. psychrophilum using injections
under the skin, and to M. cerebralis using bath exposure to
triactinomyxons, the waterborne infectious stage of the parasite. Mortality
from F. psychrophilum occurs within 28 days of exposure, and was the first
measureable endpoint of the experiment. Fish were then reared for six
months to allow development of myxospores, the countable form of M.
cerebralis in fish, and disease signs and myxospore counts were obtained
from the fish remaining at the end of the experiment.

The PRR experienced the lowest cumulative mortality when exposed to F. psychrophilum, showing that it was
more resistant to F. psychrophilum than either the HxH or HHP. However, with higher myxospore counts than
either the HxH or HHP, the PRR did not show any resistance to M. cerebralis. The HxH had much lower
myxospore counts and was more resistant to M. cerebralis than the PRR. However, as had been observed in
the hatchery, the HxH did not exhibit any resistance to F. psychrophilum. The myxospore counts for the HHP
were intermediate to those of the HxH and PRR, and the HHP experienced high cumulative mortality when
exposed to F. psychrophilum, showing that it had not gained resistance to F. psychrophilum from the PRR.
Coinfection with F. psychrophilum and M. cerebralis increased mortality in the PRR, HxH, and HHP compared
to single-pathogen exposure.

Left: Cumulative percent mortality (CPM) for the HHP, HxH, and PRR across six treatments, 1) control (no pathogen exposure), 2)
F. psychrohilum only (Fp), 3) exposure to F. psychrohilum followed by exposure to M. cerebralis (Fp Mc), 4) M. cerebralis only
(Mc), 5) exposure to M. cerebralis followed by exposure to F. psychrophilum (Mc Fp), and 6) mock injection with TYES media
(TYES). Right: Myxospores per fish head as a measure of M. cerebralis infection for the HHP, HxH, and PRR in each of the six
treatments at the end of the experiment.

Management Implications
The results of this experiment suggest that it was not possible to create fish that are resistant to both F.
psychrophilum and M. cerebralis using the HxH and PRR. In a follow up experiment, we exposed pure strains
and their crosses to F. psychrophilum and found that the first generation cross between the Harrison Lake
Rainbow Trout and the PRR showed reduced mortality and resistance to F. psychrophilum. Therefore, it may
be possible to produce Rainbow Trout that are resistant to both pathogens, though their resistance to M.
cerebralis still needs to be evaluated. More research is needed to determine if other strains not included in
these experiments can be used to create fish resistant to both pathogens. Until then, hatchery outbreaks of
BCWD in susceptible fish such as the HxH can be reduced by maintaining high water quality, flows, and reduced
densities to prevent stressful rearing conditions. Using PRRs in hatcheries where F. psychrophilum outbreaks
are common will help reduce mortality from BCWD on the unit, but due to their susceptibility to WD, these
fish should not be stocked in aquatic systems in which M. cerebralis is established.
Associated Literature
Fetherman, E. R., B. Neuschwanger, B. W. Avila, and T. B. Riepe. 2020. Sport Fish Research Studies. Annual Report. Colorado Parks and
Wildlife, Aquatic Research Section. Fort Collins, Colorado.

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

�</text>
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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Electric Fish Barrier
Research
Background
There are over 105,000 irrigation structures on rivers and streams across Colorado,
most in fish bearing waters. Fish loss in irrigation canals is known to be a large problem in the
western U.S. but the impact on fish populations in Colorado is unknown. The South Canal is an
irrigation ditch near Montrose, Colorado that diverts an average of 857 cubic feet per second from
March to November from the Gunnison River for agriculture. The construction of a hydropower
plant was expected to increase mortality of fish in the canal so an electric fish barrier was installed
at the diversion structure in 2012.

Research Objectives
The objective of this work is to evaluate if electric fish barrier technology can reduce the loss of
sport fish in irrigation canals in Colorado by monitoring fish populations in the South Canal and
documenting tagged fish that cross the barrier.

Electric Fish Barrier
The fish barrier was operational before the 2013
irrigation season. It consists of a series of vertically
suspended electrodes across the east portal of the
Gunnison Tunnel. The system uses pulsed direct
current (DC) to deter fish. DC is the safest type of
electrical current for fish and has been shown to repel
fish without injuring them. The barrier was designed to
exclude broodstock rainbow and brown trout with a
field strength of 1 volt per inch, a relatively low power
setting for electric fish barrier designs in the United
States. Pulsed DC current is size-selective; it affects
larger fish more than smaller fish.

Approach
A total of 23,031 fish from 3 to 29 inches in length were
tagged and released in the Gunnison River above the
barrier. Fish were removed from the canal before the
barrier was operational and annual electrofishing
surveys were used to estimate fish populations and look
for tagged fish.
COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO, 80216 • (303) 297-1192 • www.cpw.state.co.us

�Results and Conclusions


The electric barrier successfully prevents
large fish from entering the canal and being
lost to the Gunnison River population.



No fish larger than 16 inches in length have
passed the barrier, and only four fish larger
than 12 inches have passed through the
barrier.



Smaller age 1, age 2, and some age 3 fish can
pass through the barrier.



The barrier prevents 26-71% of all of the
spawning sized fish from entering the canal
and being lost to the Gunnison River
population.



The number of brown trout in the canal
declined after the barrier, but growth and
survival of smaller fish that pass the barrier
maintain a stable fish population.



The electric barrier successfully protects
larger brood fish in the Gunnison River but
more work is needed to see if it can be
adjusted to better exclude smaller fish.

4,000

3,500
Brown Trout
3,000

Population Estimate

Rainbow Trout
2,500

2,000

1,500

1,000

500

0
Oct 2011

Mar 2013

Oct 2013

Jul 2014

Oct 2014

Brown trout numbers have declined since the barrier
was built in 2012 but small fish that pass into the canal
maintain a stable trout population.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO, 80216 • (303) 297-1192 • www.cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Fish Bioenergetics Research
COUNTING CALORIES FOR SPORT FISH IN COLORADO’S COLDWATER RESERVOIRS

What is fish bioenergetics and why is it useful?
Fish bioenergetics is the study of how factors such as metabolism, water temperature, and quality of prey regulate the
ability of fish to convert energy from food into body mass or growth in weight. Much effort has been devoted to
developing models that describe the bioenergetic characteristics of different species of sport fish found in Colorado’s
lakes and reservoirs and elsewhere. These models help biologists address a number of critical fisheries management
questions and guide efforts to improve growth within fish populations and ultimately fishery quality. For example, fish
bioenergetics is useful for answering:

o What limits fish growth the most: temperature, prey availability,
or the quality of prey?

o How much prey do predators eat? Is it too much, or can prey
populations handle higher levels of predation?

o What is the optimal number of fish to stock based on the available
food supply in a lake or reservoir?

Caloric content of fish: the currency of bioenergetics
Knowing the “energy richness” of fish is important because it is an indicator of fish health, prey quality, and because
energy is the common currency used to convert the amount of food eaten by a fish into weight gain or loss in
bioenergetics models; this approach is similar to counting calories in the foods we eat and tracking the fate of those
calories once ingested. Many factors influence the caloric content of fish, and it is possible for the same species to differ
among lakes. In this study, we estimated the caloric content (number of calories in one gram of fish tissue) of key predator
and prey fish found in Colorado’s coldwater reservoirs to improve the applicability of bioenergetics models to CPW’s
management of these species. The average caloric content of different species examined is shown in the figure below:

The most striking observation was the high caloric content of kokanee salmon, which was 2-fold or more greater on
average than the other species of prey fish available to predators. For example, a lake trout in Blue Mesa Reservoir would
need to eat two or three rainbow trout, white suckers, or yellow perch to obtain the same amount of energy gained from
eating a single kokanee of equal size. This observation also indicates that kokanee are more efficient consumers of
zooplankton, a primary food source for prey fish in our fluctuating coldwater reservoirs. Thus, kokanee help maximize
fishery productivity. Overall, this research highlights the irreplaceable role kokanee play as a sport fish for anglers and as
energy rich prey fish for predators in Colorado’s reservoir food webs. Managing for populations of healthy, abundant
kokanee ensures the long-term maintenance of CPW’s kokanee stocking program and benefits therein to anglers seeking
to catch and eat kokanee or seeking to catch predatory sport fish like lake trout that rely on plentiful kokanee for
supporting high feeding and growth.
Associated peer-reviewed publication: Johnson, B.M., W.M. Pate, and A.G. Hansen. In press. Energy density and dry matter content in fish: new observations and an
evaluation of some empirical models. Transactions of the American Fisheries Society.

COLORADO PARKS &amp; WILDLIFE • 317 W. Prospect Rd., Fort Collins, CO 80526 • (970) 472-4432 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Fish Passage at River Structures
RESEARCH AND DESIGN GUIDELINES

Introduction
Instream structures, such as culverts, water diversions and dams, can negatively affect fish by
fragmenting populations, reducing migratory ranges, and limiting access to habitat for spawning, feeding and refugia.
Many rivers in Colorado contain man-made structures that create partial (obstacles) or complete barriers depending on
the fish species and life stage. Habitat fragmentation associated with instream barriers is a serious threat to Colorado’s
Species of Greatest Conservation Need (SGCN) and sport
fisheries. Therefore, it is important that fisheries managers
(A)
identify and evaluate the influence of instream structures on
fish populations.

Fish Passage Research Objectives
The primary goal of fish passage research is to restore
connectivity in fragmented river systems by: (1) evaluating the
effectiveness of existing fishways; (2) evaluating the barrierpotential of common river structures; and (3) establishing fish
swim performance criteria for native and sport fishes.

Current Fish Passage Research Projects
Active fish passage research projects include: (1) evaluation of
native fish passage at existing fishways located on Front Range
transition zone streams; (2) evaluation of fish passage at
instream whitewater park structures; (3) laboratory studies to
develop fish swim and jump performance criteria for Colorado
fishes where data is lacking; and (4) development of new
techniques and technologies for investigating fish movement
and passage in rivers.

(B)

Fishway Design
Fishways, or “fish ladders”, are engineered structures
designed to facilitate passage around an obstacle or barrier.
Fishways attempt to incorporate species- and life stagespecific swimming and jumping abilities into designs. Common
elements of successful fishways include: (1) low velocity
pathways that do not exceed burst speeds or endurance
capabilities for target species (Figure A); (2) water depths that
do not limit swimming performance (Figure B); (3) vertical
drops that do not exceed the jumping ability for target species
- note that many species native to Colorado do not exhibit
jumping behaviors (Figure C); (4) sufficient attraction flow, or
the flow that emanates from a fishway entrance, to ensure
that fish can locate the fishway; and (5) maintenance of the
above design elements over the expected range of
streamflows.

(C)

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

�Fishway Examples
Some examples of successful fishways include engineered rock ramps (Figure D), constructed riffles (Figure E), and
vertical slot fishways (Figure F). Each type of fishway has advantages and disadvantages related to which fish species
and life stages are present and the conditions of the project site.

Engineered Rock Ramp

Constructed Riffle

Vertical Slot

Diversion Crest

Piney Creek,
Wyoming

Fossil Creek Reservoir
Inlet Diversion,
Cache la Poudre River

(D)

Rock Weirs

CCC Ditch,
San Miguel River

(E)

(F)

Aquatic Habitat Types
From the high-gradient, boulder-dominated, step-pool
channels of snowmelt fed mountain streams to the lowgradient, well-vegetated, pool-riffle rivers of the eastern
plains to the majestic, vertically-confined canyons on the
arid Colorado Plateau, aquatic habitats in Colorado are as
diverse as the geographic regions where they are found.
Native Colorado fishes have unique morphological
characteristics that are adapted to the natural conditions
found in each aquatic habitat type. These adaptations affect
the swimming abilities of fish, influencing how they move
through and use diverse habitats. Fisheries managers must
take the diversity of fish species into consideration when
evaluating river structures and designing fishways.

Fish Swimming Performance by Family
Family Name
Percidae (Perches)

SGCN (#)

Fundulidae (Topminnows)
Cottidae (Sculpin)
Ictaluridae (Catfish)
Cyprinidae (Minnows)
Catostomidae (Suckers)
Centrarchidae (Sunfish)

All illustrations of fish © Joseph R. Tomelleri

3

Prolonged Speed (ft/s)
0.4 - 1.2

Burst Speed (ft/s)
NA - 2.4

Jump Height (ft)
0*

Habitat Types
EP

1
0
1
13
5
1

1.3 - 1.6
1.4 - 1.7
1.3 - 2.0
1.3 - 2.4
1.3 - 2.5
1.1 - 2.9

2.6 - 3.4
3.3 - 3.9
2.0 - NA
2.4 - 4.4
2.2 - 3.2
2.6 - NA

0.1 - 0.2
0*
NA - 0.2
0* - 0.5
NA - 0.8
0.4 - NA

EP
CP, MS
EP, TZ
CP, EP, MS, RG, TZ
CP, EP, MS, RG, TZ
EP

Salmonidae (Trout)
3
2.3 - 4.0
4.5 - 7.5
1.0 - 7.0
MS, RG, TZ
SGCN = Species of Greatest Conservation Need, # of species/subspecies; * = fish species does not exhibit jumping behavior; NA =
data were not available; CP = Colorado Plateau, EP = Eastern Plains, MS = Mountain Streams, RG = Rio Grande; TZ = Transition Zone

The values reported above are summarized from multiple species within each family and are intended to support passage
for juvenile life stages. Swim speeds and jumping abilities within species are size dependent. Species-specific performance
criteria should be used whenever possible. The selection of target species for individual projects should be based on the
management objectives for the site in question. Consultation with the local Area Aquatic Biologist at CPW is strongly
encouraged during the early planning stages for any fish passage project in Colorado. The information in this fact sheet is
based on the best available data and knowledge, but is subject to revision as more information becomes available.
COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

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                    <text>C O L O R A D

O

P A R K S

&amp;

W I L D L I F E

Gill Lice in Colorado
NOVEMBER 2018

Gill Lice
Gill lice are an external parasite of trout and salmon found throughout the US including Colorado. These parasites have
the potential to negatively influence salmonid fisheries throughout Colorado because they can impair fish growth,
behavior and survival.
To date, only one species of gill lice has been identified in Colorado,
Salmincola californiensis. This species of gill lice is known to infect
kokanee salmon, cutthroat trout, and rainbow trout throughout the
state. No infection of gill lice has been confirmed on any other
species of fish in Colorado. CPW continues to work on potential
management actions or other approaches to minimize the negative
impacts of gill lice on Colorado’s valuable native and sport fisheries.
Specifically, CPW is attempting to:
• Assess the current infection presence and intensity of gill lice.
• Determine the extent of negative effects on fish
populations throughout the state.
• Identify conditions leading to high levels of gill lice.
• Identify potential management actions and approaches
to reduce negative impacts on fisheries.

Gill Lice Background
Gill lice (of the genus Salmincola), are a parasitic zooplankton that generally attach to the gills, mouths and fins of
salmonid fishes (e.g., salmon, trout). Since the initiation of the gill lice project in Colorado in 2012, gill lice have been
identified on cutthroat trout, rainbow trout and kokanee salmon. It is important to note that gill lice infestation does not
make fish unsafe for consumption.
Heavy gill lice infections can prevent fish from breathing normally, inhibiting oxygen uptake through damaged gill
filaments. Gill lice infestations can also negatively affect fish behavior, immune system function, growth, warm water
tolerance and ultimately survival.
Gill lice were first reported in Colorado in the early 1900’s. Until relatively recently, the negative effects associated with gill
lice on fish were limited to infestations in hatcheries, though the presence of gill lice in some Colorado water bodies
outside of hatcheries was known. However, more recently, anecdotal evidence has linked declines in fish populations
with the occurrence of gill lice, and it is important to understand these more recent observations. Specifically, kokanee
salmon populations in many lakes throughout the west have completely collapsed after becoming infected with gill lice.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman, Suite 618, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us

�Kokanee salmon and native cutthroat trout are ecologically and economically important fish species in Colorado that have
experienced recent population declines. These populations are especially important to conserve, preserve and protect.
Gill lice research is focused on identifying and developing options for limiting the spread of gill lice to these populations as
well as minimizing negative effects from gill lice in native fish populations where gill lice are already present. This research
will help maintain and potentially enhance these valuable fisheries and the ecologically and economically important fish
species that support them.
CPW research on gill lice is especially important because controlling or eliminating the parasite in wild fish populations is
currently diffic ult or impossible, and this project is aimed at gaining an understanding of gill lice to address this
challenge and present viable management options to reduce negative impacts associated with gill lice spread.

Current CPW Gill Lice Research Projects
•
•
•
•

Working with researchers and students at Colorado State University (CSU) to improve the understanding of how gill
lice affect trout and salmon in the state.
Investigating the current and potential consequences of gill lice on kokanee salmon and native cutthroat trout.
Investigating the effects of fish size and stress on infection rates to determine potential management options. Fish
stressed by high population density, low water levels, warm water temperatures or spawning may be more
susceptible to gill lice.
Completing experiments to determine if some fish species are more likely to become infected with gill lice.

Anglers Can Help!
If you would like to report gill lice in Colorado, please e-mail estevan.vigil@state.co.us or george.schisler@state.co.us at CPW.
Please include information about:
Species of fish caught?
Where was the fish caught?
How long was the fish?
What parts of the fish had gill lice?
Any photos.
COLORADO PARKS &amp; WILDLIFE • 1313 Sherman, Suite 618, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Lake Trout Population Monitoring
RAPID ASSESSMENT OF LAKE TROUT POPULATION SIZE IN LAKES AND RESERVOIRS

How are lake trout populations monitored?
Lake trout are top predators, reproduce naturally, and are important
sport and food fish for anglers in Colorado’s lakes and reservoirs.
Keeping close tabs on their abundance and size structure is necessary
for assessing the appropriateness of harvest regulations, ensuring lake
trout remain in balance with prey fish populations, and determining
whether management goals are achieved. However, estimating the
abundance of lake trout in our large coldwater reservoirs at the
frequency needed to inform management using conventional methods
such as mark-recapture is impractical. Thus, CPW researchers and
biologists evaluated the utility of “Summer Profundal Index Netting”
(SPIN) for application to lake trout in Colorado beginning in 2011.
Developed by the Ontario Ministry of Natural Resources, SPIN is a
standardized gill netting method for estimating the density of lake
trout ≥12 inches over a 1-2 week period. Colorado has since adopted
SPIN for monitoring lake trout populations in Blue Mesa Reservoir,
Grand Lake, Lake Granby, and Taylor Park Reservoir.

Survey design, implementation, and results:
Standard gill nets specific to this protocol are set for two hours during
daylight along the bottom in random orientation when surface
temperatures exceed 64°F. You may see these nets being set or pulled
at these lakes by CPW personnel during the summer months. The
number of sets depends on the surface area of the lake or reservoir. Set
locations cover depth intervals of 6-32 ft, 32-64 ft, 64-96 ft, 96-128 ft,
128-192 ft, 192-256 ft, and &gt;256 ft. Sampling is also conducted in
different portions of the lake or reservoir if necessary to account for
differences in lake trout habitat. Catch rates of lake trout in the gill
nets set in Colorado systems are compared to catch rates of lake trout
in the same type of gill nets in other water bodies where estimates of
lake trout density were available. The advantage of this method is the
use of data from numerous other systems as a calibration tool to
quantify lake trout densities and abundances in Colorado. Please see
Lepak (2011) and Hansen (2018) for additional information.

Management implications:
Results from each SPIN survey conducted to date shows that lake
trout density varies 7-fold across study systems. This indicates that
processes governing the productivity of lake trout vary or operate
differently in each lake or reservoir. Thus, management plans and
corresponding harvest regulations must be tailored to each individual
lake trout population in order to achieve desired goals for the fishery.

Map of Blue Mesa Reservoir showing gill net set locations (N = 83
dots) and the associated depth interval sampled (colors) during the
2016 SPIN survey:

Summary data from each SPIN survey conducted to date. Asterisks
indicate the presence of Mysis shrimp in the body of water.
Survey year

Number of
lake trout
caught

Mean
length
(inches)

Density
(fish/acre)

Total area
surveyed
(ha)

Abundance
estimate

Blue Mesa

81

129

17.2

4.51

7,559

34,071

Grand Lake*

36

87

16.5

5.14

477

2,452

Taylor Park*

36

271

16.4

7.94

1,507

11,950

Blue Mesa

81

211

16.7

3.18

8,424

26,753

Lake Granby*

71

501

16.4

23.17

6,870

159,193

Blue Mesa

83

180

17.2

2.89

8,424

24,368

Grand Lake*

36

109

17.2

6.56

477

3,131

Blue Mesa

95

313

16.3

4.60

6,496

29,857

2011

2013

2014

References:
Hansen, A.G. 2018. Summer profundal index netting for tracking trends in the abundance of lake trout in
coldwater lakes and reservoirs of Colorado: results from 2018. Internal CPW report. 6 pages.
Lepak, J.M. 2011. Evaluating summer profundal index netting (SPIN) as a standardized quantitative method
for assessing lake trout populations. Internal CPW report. 10 pages.

Number of
gill net sets

Lake or
reservoir

2016

2018

COLORADO PARKS &amp; WILDLIFE • 317 W. Prospect Rd., Fort Collins, CO 80526 • (970) 472-4432 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Managing Mercury in Sport Fish
TRIPLOID WALLEYE BIOACCUMULATE LESS MERCURY

Mercury and Colorado Sport Fish
According to the U.S. Food and Drug Administration, fish are part of a healthy
diet, providing lean protein, healthy omega-3 fatty acids, iron, and more vitamin
B12 and vitamin D than any other food. Although fish are an excellent source of
protein, integration of mercury into lake and stream food webs results in traces of
mercury being present in fish, sometimes at levels that present health concerns to
people who consume fish regularly. Because mercury levels increase as you move
up the food chain (i.e., ‘bioaccumulate’), concentrations are highest in top
predators such as walleye. Through monitoring and research, both the Colorado
Department of Public Health and Environment and Colorado Parks and Wildlife
(CPW) work to help anglers maximize the benefits of consuming fish while
protecting themselves and their families from mercury in the environment.

How does mercury get here?
Mercury is released into the atmosphere through a variety of natural (e.g., forest
fires) and human-caused (e.g., burning fossil fuels) pathways. Mercury can then
disperse far from its source before being deposited into lakes and streams, making
it difficult to control. While national and global actions to reduce mercury
pollution are essential, strategies at the local scale are needed to protect the health
of Colorado’s anglers. A recent study on Narraguinnep Reservoir (southwest
Colorado) conducted by CPW and researchers from Colorado State University
found that ‘triploid’ walleye (have a third set of chromosomes and cannot
reproduce naturally) stocked by CPW had 22-24% lower mercury concentrations
on average than their naturally-reproducing ‘diploid’ counterparts (have a normal
number of chromosomes). This study revealed that stocking triploid fish alone or
in tandem with other mitigation strategies can lead to safer fish for anglers to eat.

Map of Colorado showing waters with a current
fish consumption advisory (red balloons) versus
those without (green). Interactive map accessible
here:
https://cdphe.colorado.gov/waterquality/clean-water/rivers-lakes-andstreams/fish-consumption

Why do triploid walleye bioaccumulate less mercury?
The answer begins with how and why triploid walleye are produced by CPW.
Every spring CPW collects eggs from wild populations of walleye to raise in the
hatchery and stock fry into reservoirs that support high angling pressure. Some of
these eggs are specially treated to produce triploids. Triploidy is induced by
placing fertilized eggs into a hydrostatic pressure chamber for a defined period
(see images to right). As a result, the fry that hatch retain an extra set of
chromosomes, rendering them incapable of naturally-reproducing over their
lifespan. Because triploid walleye are sterile, they can be stocked to support sport
fisheries in strategic locations where natural reproduction is not wanted. As this
new research discovered, the inability of triploid walleye to reproduce also affects
their mercury levels. Spawning is energetically costly, particularly for female fish
that need to develop eggs. As a result, diploid walleye need to eat more food to
grow and successfully spawn every year when compared to triploid walleye. Since
most (&gt;95%) of the mercury present in predatory fish comes from their prey,
eating less food to grow the same amount means triploid fish are exposed to less
mercury, and have lower mercury concentrations.
Associated Publications:
Farrell, C.J., B.M Johnson, A.G. Hansen, and C.A. Myrick. In press. Induced triploidy reduces mercury
bioaccumulation in a piscivorous fish. Canadian Journal of Fisheries and Aquatic Sciences.

Using gill nets to capture and test triploid and
diploid walleye in Narraguinnep Reservoir for
mercury levels.

Sequence of images depicting the process of
collecting, fertilizing, treating and hatching
triploid walleye eggs for stocking into key
Colorado reservoirs.

COLORADO PARKS &amp; WILDLIFE • 317 W. Prospect Rd., Fort Collins, CO 80526 • (970) 472-4432 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Native Cutthroat Trout
SEPTEMBER 2018

Rediscovery of the San Juan River basin’s native trout

© GRANT WILCOX/CPW

Nine years ago, the Greenback Cutthroat Trout Recovery Team commissioned a collaborative study by researchers
from the University of Colorado, along with state, and federal biologists to examine Cutthroat Trout specimens
collected in the late 1800s and housed in our nations most prestigious museums. The goal of that study was to
establish what the native range of these fish
was prior to European settlement. While
these trout look similar to each other in
outward appearance, inspection of their DNA
revealed that six discrete lineages historically
called Colorado home – essentially one in
each major drainage basin. Four lineages are
still found on the landscape today: two that
comprise Colorado River Cutthroat Trout
west of the Continental Divide, Greenback
Cutthroat Trout in the South Platte River
drainage, and Rio Grande Cutthroat Trout in
their namesake river basin. The authors

©KEVIN ROGERS/CPW

Distinct lineages of Cutthroat Trout evolved in six
major drainage basins in Colorado

Specimen collected from the San Juan River by C. E. Aiken
in 1874 (now at the National Museum of Natural History
in Washington D. C.)

reported that the remaining two lineages were thought
to be extinct: the Yellowfin Cutthroat Trout historically
found in the headwaters of the Arkansas River, and one
native to the San Juan River that was only known from
two specimens collected near Pagosa Springs during
the Wheeler Survey in 1874 while mapping the
territories in what is now the American southwest.

After learning that a previously unknown lineage of Cutthroat Trout was native to the San Juan basin, we launched an
intensive search to determine if any relict populations remained tucked away in the corners of the rugged San Juan
Mountains of southwest Colorado.

COLORADO PARKS &amp; WILDLIFE • 317 W. Prospect Rd., Fort Collins, CO 80526 • (970) 472-4432 • cpw.state.co.us

�Approach

©KEVIN ROGERS/CPW

Armed with new genetic markers with which to identify this San Juan lineage trout, we gathered tissue samples (fin
clips) from all known cutthroat trout populations in southwest Colorado. We isolated DNA and sequenced a specific
mitochondrial gene to identify any candidate fish that might have the same marker as those found in the museum
specimens. We then examined their
nuclear DNA to determine if any
Present day San Juan lineage trout from a small stream in the
hybridization had occurred with
headwaters of the San Juan River
Rainbow Trout or other subspecies of
Cutthroat Trout, or if they appeared
to be pure.

Results

•

•
•

Surveyed the 20 waters thought to
contain pure Cutthroat Trout in the
San Juan River basin
Eight of those waters harbored
populations containing the same
mitochondrial DNA marker found
in the two museum specimens
collected in 1874
Examination of nuclear DNA also
suggests they are a unique lineage
No evidence of hybridization with
Rainbow Trout or other nonnative
Cutthroat Trout lineages was detected

Colorado River (Blue)
Cu9hroat Trout

•

South Pla9e
San Juan
Colorado River (Green)
Rio Grande
Rainbow Trout

DNA tree comparing relatedness of Colorado’s native Cutthroat
Trout lineages (from map figure) to each other and Rainbow
Trout. Longer branches represent more distant relationships

Management implications
This represents an exciting conservation success story, and is testimony to the effectiveness of our recovery strategies
for preserving native trout diversity. Though genetic tools to distinguish these fish were not available until recently,
biologists recognized that hidden diversity was likely still out there, and protected populations accordingly. Our
conservation efforts will now focus on replicating these populations and securing them from invasion by nonnative
trout so that future generations can enjoy this rare piece of Colorado’s legacy.

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Predation on Stocked Fish
ASSESSING THE PREDATORY THREAT OF INTRODUCED SPECIES TO MANAGED SPORT FISH

Does predation limit the survival of sport fish
stocked into Colorado’s lakes and reservoirs?
All fish are vulnerable to predation by larger fish during some phase of life. If predation is high and persistent, stocked
fish may not survive long enough to grow to a size that can be caught and harvested in the numbers desired by anglers. In
these situations, management actions such as stocking larger fish or adjusting harvest regulations on predators could boost
survival. Determining whether different species pose a significant predatory threat to stocked sport fish, and if so, how to
best manage the situation, first requires an examination of the interactions among suspected predators and stocked fish.

Recent study: threat of yellow perch to stocked kokanee salmon fry
Yellow perch were first detected by CPW in Blue Mesa Reservoir, Colorado’s premier kokanee salmon fishery, in 2001.
Yellow perch feed on Daphnia (food for kokanee), other invertebrates, but also small fish. There was growing concern
and perception from the public and some angler groups that yellow perch were having a detrimental effect on the stocked
kokanee fry that maintain this phenomenal fishery. CPW determined that it was important to address these concerns
objectively by conducting a focused study on the subject. During 2016, researchers estimated the seasonal diet and habitat
use of yellow perch and compared these to brown trout and to the habitat use and growth of kokanee fry. Brown trout
have been present since the reservoir was filled in 1965 so they were a good reference species for gauging the predatory
threat of yellow perch.

Key results and management implications:
Contrary to perceptions, yellow perch showed a limited predatory threat to kokanee
fry. Yellow perch behaved similarly to brown trout by occupying nearshore areas
away from offshore kokanee and eating mostly crayfish and insects throughout the
year. Predation by yellow perch and brown trout on kokanee was brief and confined
to the inlet during spring when fry stocked upstream entered the reservoir. The pie
charts below show how quickly kokanee fry disappeared from the diet of each
predator. Kokanee were 50% of stomach contents for each species the morning fry
entered the reservoir, but dropped to 0% one week later. Because brown trout were
more abundant and consumed more individual fry, their predation on kokanee in the
inlet was 39-times greater than yellow perch. Kokanee fry outgrew 58% of adult
yellow perch by June and 95% by October, further limiting predation. Yellow perch
did not measurably affect the amount of Daphnia available to kokanee. This
research highlights the importance of addressing uncertainties with appropriately
designed studies relevant to the species and reservoir in question to guide informed
management decisions.

Peer-reviewed publication: Hansen, A.G.,
J.S. Thompson, L.N. Hargis, D. Brauch,
and B.M. Johnson. Under review.
Predatory threat of illegally introduced
yellow perch in a salmonid dominated
reservoir food web. North American
Journal of Fisheries Management.

COLORADO PARKS &amp; WILDLIFE • 317 W. Prospect Rd., Fort Collins, CO 80526 • (970) 472-4432 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Rainbow Trout Stocking Evaluations
INCREASING POST-STOCKING SURVIVAL OF WHIRLING DISEASE RESISTANT RAINBOW TROUT

Whirling Disease Resistant Rainbow Trout

Whirling disease resistant Rainbow Trout have been stocked by CPW hatcheries over the past decade to
reestablish Rainbow Trout populations in waters where populations experienced significant declines due to the
establishment of whirling disease in the early 1990s. The German Rainbow (GR), or Hofer, a resistant but domesticated
Rainbow Trout strain, has been crossed with other Rainbow Trout strains to maintain both whirling disease resistance and
wild survival characteristics. Crosses of the GR with other wild strains, such as the Colorado River Rainbow and Harrison
Lake Rainbow Trout, are used for stocking various locations, for instance in rivers versus lakes, dependent upon life
history characteristics of the wild strain and the management objectives for each body of water. Since these crosses are a
relatively new addition to Colorado’s stocking program, the effects of fish size at stocking and variety were evaluated to
determine which stocking methods result in increased post-stocking survival and recruitment in newly-established
whirling disease resistant Rainbow Trout populations.

Rainbow Trout Stocking Evaluations in the upper Colorado River
Subcatchable Stocking Evaluations
A stocking strategy that was adopted in the years after whirling disease
became established in Colorado, prior to the advent of resistant strains,
was the use of large (&gt; 6 inches) subcatchable fish. This was done
because larger fish were less susceptible to whirling disease infection and
predation. Whirling disease resistant Rainbow Trout, a cross between
the GR and the Colorado River Rainbow Trout strains (H×C), were first
stocked into the upper Colorado River below Windy Gap Reservoir in
2006, with additional stocking events in 2009 and 2010. These resistantStocking Floy-tagged H×C in Colorado River
strain stocking events were also conducted with large subcatchable fish,
similar to previous stocking events with susceptible strains. Prior to being stocked, all fish were individually tagged using
external Floy tags to identify individuals upon recapture during annual population estimates. Although some natural
reproduction took place in the upper Colorado River during this time, wild fry numbers remained low and little
recruitment to the adult population occurred. As a result of both the low post-stocking survival of the stocked
subcatchable fish and the low numbers of fry produced by these fish, the adult Rainbow Trout population continued to
decline from 91 fish per mile in 2008 to 6 fish per mile in 2011 (Figure A; Fetherman et al. 2014).

RBT per mile

200

200

(A)

100

100

0

0
2008

2009

2010

2011

Figure (A) Decline in the
adult rainbow trout
population following
subcatchable stocking and
(B) increase in the adult
rainbow trout population
following fry stocking in the
upper Colorado River.

(B)

2013

2014

2015

2016

2017

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

�Fry Stocking Evaluations
Because of the continued decline in the adult Rainbow Trout
population following stocking of subcatchables, fry stocking was
proposed as a possible alternative to increase recruitment to the adult
population. Subcatchables are often held in the hatchery for six months
or more to get the fish to larger sizes for stocking, whereas fry are
stocked 1.5 months after hatching, reducing the amount of time spent
in the hatchery and the potential influence of the hatchery environment
on post-stocking survival. Use of larger subcatchables was no longer
necessary due to the inherent resistance of the new strains. H×C were
first stocked as fry in the upper Colorado River in 2013, with H×C fry
continuing to be stocked annually through 2016. Fry were distributed by raft in appropriate fry habitat throughout the
upper Colorado River. Electrofishing estimates conducted throughout the study section showed that the H×C fry
persisted as well as naturally-produced brown trout fry in the same reaches. Additionally, the number of adult Rainbow
Trout per mile increased each year following the commencement of fry stocking (Figure B), suggesting that the H×C fry
were recruiting to the adult population. As such, stocking H×C fry versus subcatchables appeared to be the best stocking
strategy for the Colorado River, and similar rivers across the state.
Stocking H×C fry in the Colorado River

German Rainbow (GR) versus H×C Fry Stocking

Electrofishing for stocked
rainbow trout fry

The observed survival of the H×C fry was promising. However, studies had shown
that eventual dilution of resistance of these stocks could be a concern (Fetherman et al.
2012). Reduced resistance in future generations would be much less likely when
stocking pure GR fish. Therefore, stocking pure GR as fry could be an option for
establishing whirling disease resistant Rainbow Trout populations in certain situations.
Comparative Survival of GR and H×C Fry
In a recent collaborative study with Colorado State University, a laboratory and field
experiment were undertaken to compare the post-stocking survival of GR and H×C
fry. The field experiment was conducted in one mile reaches of nine Colorado streams,
selected for consistency in size and physical habitat characteristics. Each stream was
stocked with 5,000 GR and 5,000 H×C fry, and survival was assessed at two months, six
months, and one year post stocking. After one year in the streams, the strains survived equally well. In the laboratory
experiment, equal numbers of GR and H×C were stocked into tanks with brown trout as predators and held for 24 hours.
Again, survival did not differ between the strains. The results of this study indicated that stocking the GR as fry may be a
viable option in some streams where whirling disease is enzootic (Avila et al. In review). To test this in a larger river
system, GR fry were stocked in place of H×C fry in the Colorado River in 2016, and continue to be evaluated annually for
survival and recruitment to the adult population. Fry stocking and recruitment evaluations will continue in the upper
Colorado River through 2022.
GR fry two months post-stocking

References
Avila, B. A., D. L. Winkelman, and E. R. Fetherman. In review. Survival of whirling
disease resistant Rainbow Trout fry in the wild: a comparison of two strains.
Journal of Aquatic Animal Health.
Fetherman, E. R., D. L. Winkelman, G. J. Schisler, and M. F. Antolin. 2012 Genetic
basis of differences in myxospore count between whirling disease-resistant and
–susceptible strains of Rainbow Trout. Diseases of Aquatic Organisms 102:97-106.
Fetherman, E. R., D. L. Winkelman, M. R. Baerwald, and G. J. Schisler. 2014. Survival
and reproduction of Myxobolus cerebralis resistant Rainbow Trout in the Colorado
River and increased survival of the age-0 progeny. PLoS ONE 9(5):e96954.

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

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                    <text>REBUILDING FLOOD-DAMAGED DIVERSION
STRUCTURES TO BENEFIT MULTIPLE USES
Colorado Parks and Wildlife (CPW), the Colorado Water Conservation Board (CWCB), federal agencies and
other stakeholders can assist municipalities, ditch and reservoir companies and other water users in
reconstructing flood-damaged water diversion structures in a manner that will maintain historical water
diversions while providing for fish and boat passage. CPW stream reconstruction specialists and engineers are
available to share conceptual designs for structures successfully used throughout Colorado on all sizes of
streams. These designs ensure that water users can fully divert under their decreed water rights while allowing
fish access to the entire reach of stream, at little to no additional costs. Designs range from the traditional
concrete diversion structure with associated rock ramps to stand-alone boulder cross-vanes, or a combination of
concrete diversion structures and boulder cross-vanes.

TRADITIONAL DIVERSION WITH ROCK RAMP

BOULDER CROSS-VANE DIVERSION

TRADITIONAL DIVERSION WITH ROCK RAMP

BOULDER CROSS-VANE DIVERSION

�Potential benefits of these types of diversion structures include:
•
•
•
•
•
•

Allowing full diversion of decreed water rights while facilitating fish and boat passage
Increasing habitat connectivity for fish
Improving recreational potential and reducing hazards
Withstanding high flows (boulder cross-vane diversions on Boulder and South Boulder Creek survived
the recent flooding, minus a few boulders)
Serving as a temporary diversion structure during this period of high construction costs, or while the
owner pursues additional financial assistance
Enhancing stream health without impacting ability to fully divert and use water

If you are interested in exploring this approach to rebuilding, please contact:
Ken Kehmeier, Colorado Parks and Wildlife
317 W. Prospect
Ft. Collins, CO 80526
ken.kehmeier@state.co.us

FUNDING SOURCES INCLUDE:
CWCB Emergency Loans: Available to fund repair or replacement of raw water supply structures damaged by
flood, including river diversion dams, ditch rehabilitation, and measuring devices; can incorporate design
elements to facilitate fish passage. Terms: 3 years of no payment and zero interest + 27 yrs of payments at
current CWCB interest rates. (Contact Anna Mauss at 303-866-3441, ext. 3224; anna.mauss@state.co.us).
CWCB Fish and Wildlife Resources Fund: Grants to owners of existing water supply facilities to help
balance between development of the state’s resources and protection of the state’s fish and wildlife resources.
Can be used for construction projects designed to directly mitigate or significantly reduce the environmental
impacts of existing water facilities. (Contact Chris Sturm at 303-866-3441, ext. 3236; chris.sturm@state.co.us).
U.S. Department of Agriculture: A list of USDA funding sources is attached to this fact sheet.
For information on CWCB’s loan and grant programs, see
http://cwcb.state.co.us/LoansGrants/Pages/LoansGrantsHome.aspx

OTHER RESOURCES:
Trout Unlimited can provide seed money to help with fish-friendly rebuilding efforts; assist in
seeking grants for restoration projects, and provide volunteer support on projects (eligible as match for FEMA).
(Contact Stephanie Scott at sscott@tu.org).
U.S. Fish and Wildlife Service can provide assistance in project design, hydrologic analysis and some funding
towards improving or establishing “fish friendly” passage designs, irrigation diversions, road culverts and
bridges and the removal of obsolete dams and instream structures. Projects must have a fish and wildlife tie
and focus on state or federal protected species. Funding decisions are based on region-wide needs, USFWS
priorities and needs identified by USFWS’s various aquatic resource partners. (Contact Scott Roth at
scott_roth@fws.gov or Pam Sponholtz at Pamela_sponholtz@fws.gov).

�U.S. Department of Agriculture Programs – Potential uses for Colorado Flood Recovery
NRCS Program
Emergency Watershed
Protection Program
(EWPP)

Emergency Watershed
Protection Flood Plain
Easements (EWP-FPE)

Environmental Quality
Incentive Program (EQIP)

Wildlife Habitat Incentive
Program (WHIP)

Wetland Reserve Program
(WRP)

Relevance to Flood Recovery
Financial and technical assistance for
sediment and debris removal, stream
bank and channel stabilization, or
dike/levee repairs. Some limited
potential to assist with stream
channel repairs associated with
irrigation structures.
Financial and technical assistance to
purchase a permanent easement and
restore the floodplain to natural
conditions where it is a more
economical and prudent approach to
reducing a threat to life or property.

Time Frame
Predominantly short
term.

Financial and technical assistance for
Repair replacement of on farm
irrigation systems. Some limited
potential for off farm irrigation
systems. Potential for land
reclamation of ag land damaged by
overland flood flows.
Potential use for bank stabilization,
channel modification, and
vegetation establishment to
develop and improve wildlife
habitat.

Long Term.
Competitive grant
process.

Potential use in flood plains to
convert damaged property from
agricultural use to permanent
wetlands.

Long term.

Long Term.
Competitive grant
process

Long Term.
Competitive grant
process requiring
submittal of
application for EQIP
assistance through
local NRCS office.

Who Is Eligible
Public and private landowners are
eligible for assistance but must be
represented by a project sponsor.
Sponsors are a local unit of
government.
Private landowners within the
floodplain.

How To Apply
The Sponsor applies for NRCS
assistance via a letter to the NRCS
State Conservationist within 60
days of the flood. Sponsor and
NRCS work with affected land
owners to identify and prioritize
recovery funding needs.
Agricultural landowners apply
directly to NRCS local offices.

A project sponsor is required for
projects involving the conversion
of residential or other developed
land uses from the floodplain.
The Sponsor is required to
purchase the remaining property
after structures are removed.

For properties with residential or
other land use requiring removal
of structures the landowners work
with a local EWP Sponsor to make
an application to NRCS.

Assistance provided to individuals
for use on agricultural land.

Individual submit an application
for EQIP assistance through their
local NRCS office.

Assistance provided to individuals
for use on agricultural and
nonindustrial private forest land.

Assistance provided to individuals
for use on agricultural* land.

Individual submit an application
for WHIP assistance through their
local NRCS office.

What Is Provided
EWPP provides varying amounts
of technical assistance and 75% of
cost for installing eligible recovery
measures. Recovery measures
may be temporary or permanent
measures.

Comments
Project eligibility limits. Funding
availability is subject to
congressional appropriation.

NRCS will pay up to 100% of the
easement value and up to 100% of
the costs for easement
restoration.

Project eligibility limits. Funding
availability is subject to
congressional appropriation.

For residential housing, NRCS will
pay up to 100% of the easement
value and up to 100% of the
structure's value if the landowner
chooses to have it demolished or
relocated.
EQIP provides 100% of needed
technical assistance and an
incentive payment of typically
50%-75% of cost to construct
required conservation practices.

NRCS EWP-FP Program Contact:
John Andrews. 720-544-2834

WHIP provides 100% of technical
assistance and an incentive
payment typically in range of 50%75% of cost to construct habitat
improvement measures.

NRCS purchases 30 year or
perpetual easement and shares
cost of constructing measures
necessary to restore wetland
functions.

NRCS EWPP Program Contact:
John Andrews 720-544-2834

Program availability for new
contracts may be altered by
pending legislation to reauthorize
the Farm Bill.
NRCS EQIP Program Contact:
Dawn Jackson. 720-544-2805
Applications for new projects are
on hold pending legislation to
reauthorize the Farm Bill.
WHIP fund availability is aligned
to National &amp; State priorities.
Preble’s Mouse habitat is a
priority issue in the flooded areas.
NRCS WHIP Program Contact:
Dawn Jackson. 720-544-2805
Program availability for new
contracts may be subject to
pending legislation.
NRCS WRP Program Contact:
Dawn Jackson. 720-544-2805

�FSA Program
Emergency Conservation
Program (ECP)

Relevance to Flood Recovery
Time Frame
Financial assistance for debris removal, Predominantly short
term.
fence restoration, grading, shaping
and releveling ag land, and restoring
conservation structures.

Who Is Eligible
Assistance provided to
individuals for use on agricultural
lands.

How To Apply
Individuals submit an application
for assistance through their local
FSA Office.

What Is Provided
ECP provides financial assistance
up to 75% of cost for installing
eligible recovery measures, or up
to 90% for Limited Resource
producers.

Emergency Loan
Program

Provides emergency loans to help
producers recover from production
and physical losses due to drought,
flooding, other natural disasters or
quarantine.

Owners or operators of land
located in a county declared by
the President or designated by
the Secretary of Agriculture as a
primary disaster area or
quarantine area who suffered at
least a 30% loss.

Individuals submit a loan
application through their local FSA
Office.

Producers can borrow up to 100%
of actual production or physical
losses, to a maximum amount of
$500,000. Interest rates vary
with a loan term of one to twenty
years.

Must apply within 8
months of the county’s
disaster or quarantine
designation.

Comments
FSA ECP Program Contact:
Jenny Peterson 720-544-2895

FSA Emergency Loan Program
Contact: Gary Wall 720-5442892

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Renibacterium salmoninarum
Transmission Research
TRANSMISSION OF RENIBACTERIUM SALMONINARUM (BACTERIAL KIDNEY DISEASE) IN HATCHERY-REARED FISH

Prevalence of Renibacterium salmoninarum in the US. Highest
detections occur in the Pacific Northwest and Midwest.

Renibacterium salmoninarum, the
bacterium that causes bacterial kidney
disease (BKD), was first described in Scotland in 1930,
and first detected in the US in 1934. BKD is now found
in trout and salmon throughout the US and worldwide,
and can cause significant mortality in infected
populations. In the US, the bacterium is most
commonly detected in Coho and Chinook Salmon in the
Pacific Northwest, and Brook Trout and Brown Trout in
the Midwest. The transfer of infected eggs and the
practice of pasteurizing fish by-products for fish feed
(now discontinued) are suspected to have contributed
to the worldwide spread of the pathogen.

R. salmoninarum can be transmitted in two ways, presenting
challenges for prevention and management. Horizontal
transmission of the bacteria is not completely understood.
However, fish may become infected through direct contact
with another infected fish, ingestion of contaminated fecal
matter, or contact with a contaminated water source from
which the bacteria enters through the gills, eyes, gut, and
external injuries of the fish. Vertical transmission occurs
during the spawn. Bacteria in the female find their way into
the eggs during egg development. There is no method for
safely disinfecting the inside of the eggs, and using iodine to
disinfect the egg surface during hatchery spawns is not
effective for preventing vertical transmission.

Horizontal and vertical transmission of R.
salmoninarum.

R. salmoninarum is a regulated pathogen in Colorado. Current regulations prevent the transfer or stocking of
infected fish. While infrequently found in Colorado since the 1960’s, it has recently been detected in six
state and federal hatcheries. In 2015, R. salmoninarum was detected in the CPW Glenwood Springs
Hatchery. To eradicate the pathogen and prevent stocking infected fish, the hatchery was temporarily
depopulated. R. salmoninarum was also detected in other CPW hatcheries rearing valuable brood stocks of
whirling disease-resistant Rainbow Trout and Native Greenback Cutthroat Trout, and depopulation of these
hatcheries was not an option. Recent management options have focused on methods to prevent vertical
transmission during fish spawning, although other approaches are also being examined.

Management Options for Controlling Bacteria Transmission
Lethal spawning has been evaluated in Rainbow Trout and Cutthroat Trout brood stocks at the CPW Poudre
Rearing Unit (PRU) and Fish Research Hatchery (FRH). Eggs are fertilized and collected as normal from brood
fish during the spawn. After being spawned, adult fish are euthanized and tested for R. salmoninarum. Eggs
from positive parents are then culled so that only eggs from negative parents are retained on the unit for
stocking and management purposes. This technique has shown promise in preventing vertical transmission.
However, more research was needed to understand the transmission rates of R. salmoninarum, develop
management and regulatory protocols to decrease the prevalence of the pathogen within hatchery brood
stocks, and limit the dissemination of positive progeny to other waters and hatcheries in Colorado.
COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

�Treatment

Adults

Progeny

Antibiotics administered orally or by injection
have been shown to reduce bacterial
Control
10 Tanks
67 Families
concentrations in fish. Erythromycin (Erymin
79 Families
334 Fish
200), an antibiotic approved for use in fish,
3 Tanks
was injected into Rainbow Trout at the CPW
12 Families
FRH to determine if its use could reduce R.
Erymin 200
12 Tanks
salmoninarum levels in spawning fish and
control vertical transmission to progeny. Male
64 Families
0 Tanks
200 Fish
and female brood fish were injected with the
64 Families
antibiotic three times prior to spawning. The
25 Tanks
injections reduced bacteria levels in the
Key
injected fish to below detectable levels.
0 Families
2 Tanks
However, progeny that tested positive for R.
Negative
Positive
salmoninarum were produced by both treated
and control fish that had tested negative for the bacteria. Additionally, positive control adults produced
negative progeny. Results also indicated that the male may play a role in vertical transmission in inland
trout, a significant finding as only females were previously known to contribute to vertical transmission of
the bacteria. This study revealed several unknowns regarding pathogen transmission that needed to be
answered for effective management in Cutthroat Trout. CPW and Colorado State University are currently
conducting two studies to better understand transmission rates in Native Greenback Cutthroat Trout.

Transmission Studies in Native Greenback Cutthroat Trout
Horizontal Transmission
Horizontal transmission of R. salmoninarum within a hatchery may pose a risk to fish throughout the unit, as
well as feral fish in the river below a unit if the bacteria are present in the hatchery effluent. In summer
2019, two sentinel cages were deployed in six locations at PRU and at three time periods encompassing a
range of optimal temperatures for bacterial growth (10-18°C). Each cage housed 10 Cutthroat Trout. After
30 days, fish were collected and tested for R. salmoninarum. Only one fish out of a total of 360 tested
positive. The cage containing the positive fish had been deployed near highly-positive Rainbow Trout. These
results suggest the rate of horizontal transmission to fish both on, and in the river below a positive unit may
be insignificant, even when conditions are optimal for the bacteria to persist.
Vertical Transmission
Eggs from unique male-female pairs of Cutthroat Trout
were collected during the spawn at PRU in May 2019.
Adult fish were tested for R. salmoninarum after being
spawned to determine the potential infection status of
the progeny created from these spawns. Thirty-two tanks
of progeny are being reared at the FRH: one containing
progeny from a male negative × female negative pair, six
from a male positive × female negative pair, four from a
male negative × female positive pair, and 21 from a
Progeny collected from positive adults
male positive × female positive pair. Progeny from each
tank will be tested for R. salmoninarum and associated antibodies, indicating that the progeny had mounted
an immune response to the bacteria at some point prior to the testing, at six months and one year of age.
The results will be used to understand vertical transmission rates, male and female contributions to vertical
transmission, and the bacterial load at which transmission occurs from parent to offspring. Overall, the
results from the transmission studies will be used to inform future management options. Results from these
transmission studies will be available in 2021.
Associated Literature
Fetherman, E. R., G. J. Schisler, and B. W. Avila. 2018. Sport Fish Research Studies. Federal Aid Project F-394-R18. Federal Aid in Fish and
Wildlife Restoration, Job Progress Report. Colorado Parks and Wildlife, Aquatic Wildlife Research Section. Fort Collins, Colorado.
Fetherman, E. R., B. Neuschwanger, T. Davis, C. L. Wells, and A. Kraft. In review. Use of Erymin 200 injections for reducing vertical
transmission of Renibacterium salmoninarum in a Rainbow Trout brood stock. Diseases of Aquatic Organisms.

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

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                    <text>The "Three Species" Background
H istorically, the bluehead sucker, the flannelmouth sucker and the roundtail chub - the so-called three species inhabited much of the upper Colorado River Basin below 8,500 feet in western Colorado. Today, the three species occupy only 50 percent of
their native range in the state. Colorado wildlife managers are concerned about the range contraction and the invasion of non-native suckers,
which hybridize with the native suckers.
Dam construction that obstructed fish movement, increased water diversions and reservoirs replacing free flowing streams as habitat may
have influenced the range contraction. The remaining free-flowing water habitat below reservoirs has also changed - colder temperatures,
less fluctuation in water levels and reduced sediment.
Four related native fish - the bonytail chub, the razorback sucker, the humpback chub and the Colorado pikeminnow - experienced greater
range reductions and are federally listed endangered species. Colorado wildlife researchers want to prevent additional federal Listings.
Three Species Research Objectives
To prevent a federal endangered species listing for these three native fish species and the resulting restrictions on state management
options, Colorado Parks and Wildlife (CPW) will create a conservation plan to protect and enhance the remaining populations. Under
that plan, CPW could expand and repatriate tJ1e three species, using
hatchery produced offspring from captive broodstock. CPW has
several research projects underway.
CurrentThree Species Research Projects
Field work to address the various objectives was initiated in 2012.
Scores of randomly chosen sites witJ1in tJ1e historic range have been
visited in 2012 and 2013 lo determine whether three species fishes
are present. H undreds of genetic tissue samples an d hundreds of fin ray bones have been collected and are undergoing analysis. O ngoing
research projects include:

Bluehead sucker

Determine current distribution of the three species in comparison
to historic distribution.
Assess genetic purity and diversity in sucker populations and
make recommendations for numbers and sources of native sucker
broodstocks housed at CPW's Native Aquatic Species Restoration
Facility (NASRF) in Alamosa.
Evaluate the extent of sucker hybridization occurring in several
major drainages and assess differing outcomes compared to drainages experiencing lesser rates of hybridization.

Flannelmouth sucker

Evaluate spawning locations in tributaries, fidelity to spawning
locations, and rearing habitats in the White River basin above
Kenney Reservoir (a relatively intact native fish community), in
collaboration with Colorado State University (CSU).
Exanline population age structure and growth patterns using
thin-sectioned fin ray bones to determine flow and temperature
conditions leading lo successful spawning an d recruitment of native suckers in flow-impaired streams compared to streams with
better flows.

Roundtail chub

Help develop field keys for sucker and hybrid sucker identification.
Collaborators in the present research include Colorado State University, University of Wyoming, and Colorado Parks and Wildlife.
COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Highlights of Recent Three Species Research and Management

Bluehead sucker genetic analyses suggest that there is
considerable genetic exchange among populations, leading to the
acquisition of two broodstocks representing CPW's Northwest
and Southwest Regions.
Flannel.mouth sucker genetic analyses also revealed populations
that exchange genetic material and tl1at exhibit considerable
genetic diversity at the individual population level. One
broodstock from tl1e Northwest Region has been acquired.
Another will likely be obtained for tl1e Soutl1west Region in tl1e
future. Additional genetic analyses are underway using a second
method to confirm the results of prior research.
Roundtail chub broodstocks have been held at CPW's
NASRF facility for a number of years preceding the current
research effort. Offspring from tl1e broodstocks are stocked
in soutl1western Colorado and on tl1e Jicarilla Apache Indian
Reservation in New Mexico.

Cutting fin ray bone

All of the three species are considered to be relatively longlived, witl1 specimens exceeding 20 years of age known for
each species. Large, mature flrumelrnouth suckers tagged using
passive integrated transponders (PIT) in the Colorado River in
2003 have been detected eight to ten years later as tl1ey passed
over stationary PIT tag antenna between 2011 and 20 13.
Fin rays are known to be suitable structures for aging these longlived fish, comparable to using otolitl1s - the bone from the
inner ear - which require lethal srunpling.
Models to predict native sucker biomass from two-dimensional
strerun hydraulic modeling were developed for western
Colorado rivers in the 2000s.
In the srune study, bluehead sucker abundance was found to be
a reliable indicator ofbaseflows adequate to sustain the entire
native fish assemblage.

Sample points in northwest Colorado

tn

Magnified fin ray bone section

Sample points in southwest Colorado

1M

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Tiger Trout Research
TRANSFORMING UNDESIRABLE FISH INTO QUALITY FISHING OPPORTUNITIES FOR ANGLERS

What are tiger trout and how
can Colorado’s lakes benefit?
Many of Colorado’s coldwater lakes contain fish species, such as suckers and minnows, which provide little value to
anglers, compete with managed sport fish, and have the potential to reduce water quality. However, these undesirable fish
may be valuable prey items. Sterile fish that have the ability to grow to predatory size quickly and eat the unwanted fish
could act as a biological control agent and provide a unique fishing opportunity. Being unable to reproduce, sterile fish
can be closely managed through stocking and harvest regulations. Tiger trout, a sterile hybrid between male brook trout
and female brown trout, have the potential to fill this role. Tiger trout have already been stocked in some Colorado lakes.
Our understanding of what tiger trout eat and how well they grow and survive in lakes with different species of
undesirable fish remains limited, and should be fully investigated as a management tool.

Fine-mesh net full of fathead minnows.

Flushing the stomach of a young tiger
trout to evaluate diet (this fish was
eating insect larvae).

Three-year-old, 5 ½ lb tiger trout from
a lake containing fathead minnows.

Study objectives
Factors such as number of tiger trout stocked per acre of lake and the presence of small-bodied minnows versus largerbodied suckers may affect the ability of tiger trout to grow quickly enough and survive long enough to eat and suppress
the population of undesirable fish. The objective of this research is to quantify the feeding, growth and survival of tiger
trout stocked into lakes with different species of undesirable fish. Results will help CPW prioritize which lakes receive
tiger trout, inform appropriate numbers to stock, and calibrate expectations on the effectiveness of tiger trout as a
biological control agent. Informed stocking translates into (1) efficient use of the limited number of tiger trout produced
by our hatchery system, (2) a greater chance tiger trout will perform well at the onset of stocking, and (3) more rapid
development of a quality fishing opportunity for anglers.

Early results
Preliminary observations suggest that tiger trout grow well and
can suppress populations of undesirable fish in lakes containing
small-bodied minnows. For example, based on catch rates in nets
(number of fish captured per hour of soak time), fathead
minnows were much more numerous in three lakes where tiger
trout were either absent or only present for one year (blue bar in
figure to right) compared with two lakes where tiger trout were
present for 2-3 years (red bar). In addition, tiger trout grew from
an average of 4.0 inches at time of stocking to 19.2 inches after
just 3 years in one of the lakes. Continued study and monitoring
is needed to validate this apparent strong influence of tiger trout.
COLORADO PARKS &amp; WILDLIFE • 317 W. Prospect Rd., Fort Collins, CO 80526 • (970) 472-4432 • cpw.state.co.us

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                    <text>What is Toe Wood?
“Toe wood” is a term that describes the use of large woody materials such as log stems, root wads and
limbs for stabilizing stream banks and enhancing fish habitat (Figure A). The “toe” is the region at the base of the bank
most prone to erosion where it transitions from a vertical to horizontal plane. Using large wood to restore streams is not
a new concept. Toe wood evolved from the use of root wads to stabilize streambanks and was originally called “root
rap,” derived from the term “riprap,” with the earliest applications from the 1960s (Rosgen 2019). Toe wood is useful for
temporarily stabilizing banks until riparian vegetation becomes established, creating a “living,” vegetated streambank
for long-term stability. CPW research has documented the direct benefits of toe wood functioning as fish habitat by
improving habitat complexity, increasing fish population abundance and biomass, and enhancing pool depths.
Specifically, toe wood-treated pools were deeper (Figure B; average difference = 1.5 ft), held more quality (&gt;14“) adult
Brown Trout (× 19), higher Rainbow Trout biomass (Figure C; × 8.1), and higher total Brown Trout abundance (× 2) than
pools without large wood. In addition, total fish population abundance (× 1.5) and biomass (Figure D) were highest in
toe wood pools (Kondratieff and Richer 2022). Our research has demonstrated the effectiveness of toe wood as a bank
stabilizing technique and for riparian vegetation establishment, providing increased shading, overhead cover, and
terrestrial insect habitat, thereby indirectly benefitting fish populations (Figure E).

(A)

(B)

(C)

(D)

�Why Toe Wood?
Human activities have led to an overall reduction in the amount, size, and sources of wood found in natural channels.
Many studies have demonstrated the value of large wood for fish habitat in streams. Wood deposited on outside
channel bends or within pools can induce scour, enhancing pool depths in streams. In contrast, wood deposited in the
middle of stream channels in riffles or glides can induce deposition leading to island formation and development of
complex side channels. Wood deposited on floodplains creates roughness that traps and promotes deposition of rich,
nutrient-laden fine soils that benefit riparian plant species. Toe wood can be used to improve physical habitat conditions
and increase carrying capacity for fish if it addresses limiting factors such as a lack of low flow refugia (i.e., deep pools),
undercut banks, or habitat complexity. A major benefit of using toe wood as an alternative to other toe-stabilizing
techniques like rock or boulder materials is that wood is a natural, biodegradable material. Over time, wood
decomposes providing food, nutrients, carbon inputs, and other ecological benefits back to the stream and riparian
ecosystem that boulders, rock, or concrete riprap materials do not provide. Toe wood installations can be done using
completely natural and native materials without the use of steel bolts, cables, anchors, or large ballast boulders. This
serves as an economic advantage of toe wood for use in trout streams found within the Rocky Mountain region as
sources of woody materials are readily available in close in proximity to project sites, reducing processing, transporting,
and hauling costs.

Limitations
Toe wood is not recommended for use in every river setting in the same way that naturally occurring large wood is not
found in every stream. Some places where toe wood is not recommended include deep canyons or other places where
floodplains are non-existent, including highly-confined river channels. Toe wood should not be installed immediately
upstream of bridges, culverts, and other locations where dislodged wood pieces can hang up on structures downstream
causing blockages or obstructing river flows. River locations where there are multiple user groups such as kayakers and
rafters might not be good candidates for installing toe wood for human safely reasons since wood in streams can form
dangerous hydraulic conditions that can increase drowning risk. Other design considerations that might limit the use of
toe wood in certain river settings include consideration of the buoyancy, quality, and type of wood materials available.
Toe wood installed in locations that are subject to high levels of angler or cattle foot traffic should include fencing or
other exclusionary devices to limit disturbance to erosion-prone outside banks and protect riparian plantings.
(E)

Kondratieff, M. C., and E. E. Richer. 2022. Stream Habitat Investigations and Assistance Annual Report. Colorado Parks and Wildlife, Aquatic Wildlife
Research Section. Fort Collins, Colorado.
Rosgen, D. 2019. Natural channel design for river restoration. In P. Maurice (Ed.), Encyclopedia of water: science, technology, and society, pp. 11431155. Hoboken, New Jersey: John Wiley &amp; Sons, Inc. DOIhttps://doi.org/10.1002/9781119300762.wsts0100

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                    <text>,

C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Whirling Disease Research
September 2013

Whirling Disease Research Objectives

Colorado Parks and Wildlife (CPW) has made whirling disease research a high priority for more than 20 years
as part of the agency’s mission to conserve and perpetuate wildlife resources. When whirling disease first appeared in the state in the
1980s, CPW researchers started exhaustive projects to control the spread of whirling disease, limit the damage to fisheries and develop
other solutions. The CPW work also includes cooperative investigations with researchers in other states to coordinate resources.
Whirling Disease Background
Whirling disease is caused by a microscopic parasite that infects trout and salmon,
resulting in severe deformities and death in young fish. The disease gets its name
from the abnormal whirling or erratic tail chasing movements of some infected fish.
Symptoms also may include a black tail in young fish and a deformed skull or spinal
column in adult fish. There is no known cure for fish infected with the parasite.
Impacts of Whirling Disease in Colorado
A shipment of trout from a private, out-of-state hatchery accidently brought the
parasite to Colorado in the 1980s. By the mid-1990s, whirling disease had spread
throughout most of the state’s major river drainages.
Whirling disease has impacted rainbow, brook and cutthroat trout populations
throughout Colorado, and is known to occur in high mountain lakes above 12,000
feet. The parasite also infected most of the state’s coldwater hatcheries, reducing
production and fishing opportunities for anglers. Whirling disease doesn’t infect
humans and eating an infected fish isn’t harmful.

CPW Whirling Disease Studies

The cooperative investigations initiated by CPW have led to the development of several effective control strategies in CPW hatcheries,
which stopped the spread of whirling disease through stocking operations. Research continues into protecting uninfected waterways
and providing excellent fishing opportunities. These current CPW research projects have shown a great deal of promise:
Trout Studies
CPW researchers developed rainbow trout strains that are resistant to whirling disease and are evaluating their uses in stocking
operations. These strains of fish are able to survive exposure to the parasite. Some of these strains are now produced in CPW
hatcheries for stocking. Current evaluations are focused on determining which varieties have the best growth, survival and value for
anglers. There is ongoing work to optimize the production of these rainbow trout strains in our state hatcheries.
Other disease resistant varieties of rainbow trout are being tested for their ability to reproduce and survive in the wild to re-establish
and maintain natural self-sustaining populations. Several locations that were stocked with resistant strains already show evidence of
natural reproduction and survival in rivers, including the Gunnison, Arkansas and Colorado. Additional sites are being stocked in
coordination with further research and management activities.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Resistant Worm Studies
The complex life cycle of whirling disease poses challenges to controlling the spread. The parasite, Myxobolus cerebralis, starts life
as a water-borne spore on the bottom of waterways. Small worms, Tubifex tubifex, eat the parasite spores and become infected. The
parasite multiplies within the worms. the offspring are released into the water in a microscopic free-floating stage that attaches to fish,
infecting them.
Recent findings:
• Various lineages of worms have differences in susceptibility to whirling disease. Some of these worms are able to eat the whirling
disease parasite without becoming infected. Transplanting resistant strains of worms to areas where the susceptible varieties exist
may help reduce the level of infection in those areas.
• Studies in the Upper Colorado River Basin and Windy Gap Reservoir dating back to 1997 found four lineages of T. tubifex,
including three with lesser or no susceptibility to the disease. Ongoing research found that those three lineages have become
the dominant varieties in the reservoir while the most susceptible worm lineage has declined to less than 15 percent of the
population. There’s no known cause for the shift, but CPW researchers are continuing their work.
Whirling Disease Life Cycle

Myxobolus cerebralis

Young rainbow trout

Infected adult brown trout

Ongoing Prevention Strategies
CPW researchers worked with wildlife managers and other agencies to develop prevention strategies that are used successfully
throughout Colorado to control the spread of whirling disease into additional waterways:
• Protocols to keep hatcheries disease free, including periodic testing and installation of ultraviolet lights at one hatchery to kill
whirling disease spores.
• Policies to protect uninfected streams and rivers from receiving stocking fish from infected hatcheries.
Whirling Disease In Colorado

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Whitewater Park Studies
RESEARCH RESULTS AND DESIGN GUIDELINES

Whitewater Park Research
With over 30 whitewater parks (WWPs) either completed or in the
planning phases, Colorado is the epicenter for WWP development in the
United States. Although WWPs provide economic and recreational
benefits for local communities (Hagenstad et al. 2000; Loomis and
McTernan 2011), they may have unintended impacts on instream biota
and stream functions, particularly when the hydraulic conditions formed
by the WWP are different from those naturally found in the surrounding
river. The impact of WWPs on habitat connectivity and instream habitat
quality have been the focus of several recent studies. Although these
studies have primarily focused on fish passage and habitat, impacts to
aquatic insects and sediment transport may also occur at WWPs.

Fish Passage Impacts
The elements that create a desirable surf wave (increased velocity,
decreased depth, a hydraulic jump, and a stable, often grouted stream
channel) create conditions that can impede fish movement. Swimming
speeds and jumping ability vary greatly between fish species.
Suppression of upstream trout movement has been documented at WWP
structures, but the degree of impact varied by fish size and characteristics
of the individual structure (Stephens et al. 2015; Fox et al. 2016). As trout
are among the strongest swimming and jumping fish species in Colorado,
small-bodied and weaker-swimming fish native to Colorado streams are
even more susceptible to habitat fragmentation associated with WWP
development.

Brown Trout

Mottled Sculpin

Fish Habitat Impacts
Although WWPs create deep pools, observed fish densities were significantly higher in natural pools than in WWP pools
(Kolden et al. 2015; Kondratieff et al. in preparation). Habitat degradation in WWPs was associated with the unnatural
hydraulics created by the recreational features and conversion of riffle habitat to drops over the wave structures.

Design Guidelines
CPW recommends that adequate environmental safeguards be included in the design and construction of WWPs to ensure
that stream functions, fisheries, and recreational fishing are not adversely impacted. Each structure must be examined on a
case-by-case basis, and monitoring and adaptive management should be included in the proposed project budget.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St., Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

�Site Selection




Design and construction of WWPs should preserve the
natural aesthetic qualities of the river. WWPs should be
located in degraded reaches when possible and should aim
to improve the natural functions of the reach rather than
maintain degraded conditions. WWPs should not be
constructed in natural, un-modified river channels
(American Whitewater 2007).
WWP sites should be selected to minimize recreational
conflicts with anglers. There is increased potential for
boaters to displace anglers at WWP sites, especially during
the summer months. If WWP construction affects a popular
fishing location, mitigation such as new fishing access or
habitat improvements should be considered.

Ecological Design Considerations







WWP structures must be designed to allow upstream fish passage for all life stages of native and sport fishes present
throughout the annual hydrologic cycle. Fish passage is dependent on water velocity, water depth, vertical height of
structures, linear distance of the passage corridor, surface roughness, and attraction flow.
Hydraulic characteristics at WWP features generally conflict with ideal conditions for fish passage. Therefore, a fish
passage channel separate from the WWP structure may be necessary. The passage channel should meet hydraulic design
criteria for target species across a range of flows.
Hydraulic modeling of the proposed structure should be conducted during the initial design phase to evaluate potential
impacts to fish passage and habitat.
Streambed and bank disturbance due to construction activities should be scheduled for a time of year when egg
incubation is not occurring. An increase in fine sediment to the stream during incubation can suffocate developing
embryos. Erosion control and revegetation plans utilizing native riparian species should be required for each project.
WWP structures should not cause sediment deposition upstream or downstream of the structure. Sediment deposition
can eliminate fish and benthic macroinvertebrate habitats, create favorable conditions for the spread of whirling disease
in trout, and increase flooding risk if sediment deposition decreases channel capacity.
Recreational In-channel Diversion (RICD) water rights can be acquired for WWPs to provide recreational experiences
in and on the water. These protected flows should be managed to benefit boating recreation as well as conservation and
management of native and sport fish. Flows deviating from the natural flow regime, such as water calls during spawning
periods, could have adverse impacts on stream ecology (Poff et al. 1997).

References
American Whitewater, 2007. Whitewater Parks – Considerations and Case Studies.
https://www.americanwhitewater.org/content/Wiki/stewardship:whitewater_parks
Fox, B. D., B. P. Bledsoe, E. Kolden, M. C. Kondratieff, and C. A. Myrick. 2016. Ecohydraulic evaluation of whitewater parks as a fish passage barrier. Journal of the American
Water Resources Association. DOI: 10.1111/1752-1688.12397.
Hagenstad, M., J. Henderson, R. S. Raucher, J. Whitcomb. 2000. Preliminary evaluation of
the beneficial value of waters diverted in the Clear Creek Whitewater Park in the City of
Golden. Stratus Consulting.
Kolden, E., B. D. Fox, B. P. Bledsoe, and M. C. Kondratieff. 2016. Modelling whitewater
park hydraulics and fish habitat in Colorado. River Research and Applications. DOI:
10.1002/rra.2931.
Kondratieff, M. C., K. Kinzli, and E. R. Fetherman. In preparation. Eco-hydraulic evaluation
of whitewater parks as fish habitat in Colorado.
Loomis, J., and J. McTernan. 2011. Fort Collins Whitewater Park economic assessment.
Department of Agricultural and Resource Economics, Colorado State University.
Poff, N. L., J. D. Allan, M. B. Bain, J. R. Karr, K. L. Prestegaard, B. D. Richter, R. E. Sparks,
and J. C. Stromberg. 1997. The natural flow regime: a paradigm for river conservation and
restoration. BioScience 47(11): 769-784.
Stephens, T. A., B. P. Bledsoe, B. D. Fox, E. Kolden, and M. C. Kondratieff. 2016. Effects
of whitewater parks on fish passage: a spatially explicit hydraulic analysis. Ecological
Engineering 83: 305–318.

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Windy Gap Fish Movement Study
EVALUATING MOVEMENT IN THE COLORADO AND FRASER RIVERS PRIOR TO CONSTRUCTION OF THE CRCC

Windy Gap Fish Movement Study Objectives
The Windy Gap fish movement
study is being conducted in Grand
County in conjunction with the
construction of the Colorado River
Connectivity
Channel
(CRCC),
which will reconnect the Colorado
and Fraser rivers above Windy Gap
Reservoir to the Colorado River
below Windy Gap Reservoir for the
first time since the early 1980s.
Construction of the CRCC around
Windy Gap Reservoir will allow fish
movement and provide access to favorable habitats, such as optimal spawning locations, that have been
previously unavailable to populations upstream and downstream of the reservoir. The primary objectives of
the study are to evaluate fish movement patterns prior to the construction of the CRCC, and that the CRCC is
being used for fish passage after construction. Evaluations are taking place on Northern Water, City of Granby,
and private properties above and below Windy Gap Reservoir. To accomplish this, the fish movement study
involves collaboration between these entities, as well as CPW aquatic researchers and biologists, Trout
Unlimited, and Grand County Learning by Doing.

Population Estimates and PIT Tagging
Population estimates conducted in 2020-2022 provide baseline data on fish populations, and allow researchers
to monitor and compare changes in these populations before and after construction of the CRCC. These
estimates also allow opportunities to tag fish with Passive Integrated Transponder (PIT) tags used to evaluate
fish movements. PIT tags use the same technology as those commonly used to identify pets, providing a unique
ID for every fish that is detected by an antenna. PIT-tagged fish were released annually in the fall at two sites
in the Colorado River and two sites in the Fraser River above Windy Gap Reservoir. Additionally, PIT-tagged
fish were released in the spring throughout a four-mile stretch of the Colorado River below Windy Gap
Reservoir. Since the study began in 2020, 4,234 fish have been PIT tagged in the Colorado and Fraser rivers,
including 3,138 Brown Trout, 611 Rainbow Trout, 6 Cutbows (hybrids of Cutthroat Trout and Rainbow Trout),
and 479 Mottled Sculpin. Mottled Sculpin are currently absent downstream of Windy Gap Reservoir. The CRCC
is intended to facilitate sculpin distribution, increasing the diversity of the Colorado River downstream of the
reservoir.

Left to right: 32-mm PIT tag for evaluating Brown Trout and Rainbow Trout movements in the Colorado and Fraser rivers; 12-mm PIT tag being
inserted into a Mottled Sculpin; paired stationary antennas installed on the bottom of the river used to evaluate directionality of movement;
antenna station containing readers for recording tags as they pass stationary antennas; mobile antennas deployed in the Fraser River.

COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

�Monitoring Movement with Stationary and Mobile Antennas
Movements of PIT-tagged fish are passively monitored using two stationary antenna sites in the Colorado River
below Windy Gap Reservoir and one stationary antenna site at the confluence of the Colorado and Fraser
rivers above Windy Gap Reservoir. Paired antennas were installed on the bottom of the river to allow
researchers to determine direction of movement, depending upon which of the two antennas a fish passes
first. The antennas are connected to readers that store the date and time for every tag detected and are
operated continuously 24 hours a day, 365 days a year. Antennas constructed in the bottom of rafts use GPS
sensors to actively locate PIT-tagged fish in the Colorado and Fraser rivers in April, July, and October. These
mobile antennas allow researchers to determine the location of sedentary fish such as Mottled Sculpin or
certain age classes of Brown Trout and Rainbow Trout that move shorter distances, making it less likely that
they would be detected as a stationary antenna site.
Data collected by the stationary
antennas have provided some
valuable insights into fish
movement within the study area.
Generally, fish move more often
at night than during the day,
especially Brown Trout, likely to
avoid predators. Brown Trout
and Rainbow Trout exhibit
seasonal movements concurrent
with their spawning seasons,
Brown Trout in the fall and
Rainbow Trout in the spring.
Movement distance and timing is
also dependent on fish size.
Mottled sculpin do not exhibit
seasonal movements, but rather
move sporadically throughout
the year.

Visualizing the Data
Maps such as this one depicting tagged fish release sites in colored stars and the location of fish from those
release sites in the same colored circles allow researchers to draw conclusions about the distances fish have
moved and overall movement patterns of the populations as a whole. Using data obtained from the mobile
antenna surveys, we have seen
that, upstream of Windy Gap
Reservoir, some fish released in
the Fraser River moved into the
Colorado River, and vice versa,
and that long distance movements
are made in the Colorado River
below Windy Gap Reservoir during
certain times of year. Data
collected from the stationary
antenna sites, mobile antenna
surveys, and recaptures of tagged
fish
during
the
population
estimates will be used to obtain
estimates of detection, survival,
and movement probabilities, by
species and fish size, before and
after construction of the CRCC.
COLORADO PARKS &amp; WILDLIFE • 317 West Prospect Road, Fort Collins, CO 80526 • (970) 472-4436 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Public Access
HUNTING, FISHING AND OUTDOOR RECREATIONAL OPPORTUNITIES

Introduction

Colorado Parks and Wildlife (CPW) offers hunters, anglers, wildlife watchers and other outdoor recreationists a variety of locations to
access the outdoors. Through cooperation with private landowners and the Colorado State Land Board as well as through properties and
conservation easements owned or managed by CPW, over 2.5 million acres of land are made available to Colorado’s outdoor recreationists.

Quick Facts:
Program

Number of Properties

Number of Acres

State Wildlife Areas

348

769,000

State Trust Land

241

490,000

Walk-in Access

900

200,000

State Parks

431

200,000

Ranching for Wildlife2

29

1,200,000

State wildlife areas (SWAs) are state-owned lands that provide habitat for Colorado’s 950
species of wildlife as well as wildlife-related recreation to the public. While most activities
focus on hunting, fishing and wildlife viewing, each SWA has different permitted activities,
based on location and available resources. These parcels of SWA land are paid for primarily
by sportsmen and managed for the benefit of wildlife. CPW manages about 769,000 acres
of land within 348 SWA lands around the state. SWAs range in size from just a few acres to
several thousand. In general, there is no access fee to utilize a SWA. Since many SWAs are
managed for wildlife and wildlife habitat, seasonal closures may limit public access. CPW
encourages people to check online or check the Colorado State Recreational Lands brochure
for the specific regulations of SWAs.
(cpw.state.co.us/Documents/RulesRegs/Brochure/lands.pdf)

State Fish Units

There are 18 state fish units (SFUs) around Colorado that offer limited public access. SFUs include fish hatcheries like Durango and rearing
operations like Chalk Cliffs. Although the primary purpose of a SFU is the hatching and rearing of stockable fish, SFUs do allow self-guided
tours and a few offer limited wildlife recreation.

© WAYNE D. LEWIS, CPW

State Trust Lands

The State Land Board (SLB), an agency within the Colorado Department of Natural Resources,
owns and manages approximately 3 million acres of state-owned land commonly referred to
as State Trust Land. These properties were set aside at statehood for the specific purpose of
generating revenue and providing on going support for Colorado public schools and other
public institutions. Because these are working lands, State Trust Land is generally closed to the
public. In 1993, CPW entered into a long-term lease agreement with the SLB to provide limited
sportsmen access on approximately 500,000 acres of State Trust Land. Through the program,
sportsmen may enjoy a variety of wildlife-related uses including hunting, fishing and wildlife
viewing. Opportunities vary among properties, as do rules and regulations for authorized
use. Currently, there are 241 STL parcels providing almost 490,000 acres of wildlife-related
recreational access.
(cpw.state.co.us/Documents/RulesRegs/Brochure/lands.pdf)
1 This includes Lone Mesa State Park which is currently open for limited big-game hunting opportunities through a special permit
2 Access permitted only with a limited number of big-game hunting licenses acquired through the limited draw process

© WAYNE D. LEWIS, CPW

State Wildlife Areas

�Walk-In Access Program

The Walk-In Access (WIA) program is designed to open privately owned
land to public small-game hunting by entering into public access agreements
with landowners willing to allow public hunting. Each year, CPW holds an
enrollment period for landowners to offer their land into the WIA Program.
CPW then inspects the land and either accepts or rejects it based on its value
for hunting. All properties that are accepted into the program are published in
the regular Walk-In Atlas (late August) or the Late Cropland Walk-In Atlas (late
October) brochures. Enrolled lands are posted with WIA boundary signs so that
hunters can clearly identify properties that are open to WIA hunting. In 2015,
CPW enrolled almost 200,000 acres in the WIA program. A small-game license is
required to utilize a WIA property as small-game hunting is the only allowable use
of WIA properties. Locations of WIA properties can be found in the Small Game
Walk-in Atlas or the Late Croplands brochure.
(cpw.state.co.us/Documents/RulesRegs/Brochure/WalkInAtlas.pdf)
(cpw.state.co.us/Documents/RulesRegs/Brochure/LateCropland.pdf)

C O L O R A
D O

A D O
C O L O R

P A R K S

&amp;

NOVICE HUNTER PROGRA
PROPERT Y MAPS |

&amp;

W I L D L I
F E

I F E
W I L D L

d
2014 Late Croplan
las
At
-In
Walk
INSIDE: UPDATED WALK-IN

P A R K S

2015 Small Game
Walk-In Atlas
ACCESS TO PROPERTIES STARTS

SEPT. 1, 2015

M

online brochure

cpw.state.co.us

cpw.state.co.us
online brochure

State Parks

Colorado manages 433 state parks around the state. State parks offer a variety of
activities such as boating, hiking, fishing, hunting, camping and more. Colorado’s
state parks are open every day of the year. All state parks require an entrance
fee. Daily passes range from $7 - $9 and an annual park pass is $70. Thirty-four
parks offer camping at over 4,000 campsites and 58 cabins and yurts. Many state
parks are built around a lake or waterway, which means boating, fishing and
other water sports are among the headliners. In addition to water sports, over 700
miles of trails offer recreational activities for many park users. Limited hunting
opportunities are available on many state parks.
(cpw.state.co.us/placestogo/parks)

3 This includes Lone Mesa State Park which is currently open for limited big-game hunting opportunities through a special permit

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St, #618, Denver, CO 80203 • (303) 297-1192 • cpw.state.co.us

IE0002_PDF_12/09/15

Ranching for Wildlife (RFW) started as a pilot program in 1986 with the intent to
improve public hunting access to private land and to form a wildlife management
partnership with participating landowners. The program provides Colorado
residents a unique opportunity to hunt on private ranchland that would normally
be closed to the public. Participating ranches provide public hunting recreation
access to their land free of charge to those who draw licenses. Public licenses on
each ranch are available through CPW’s draw process. The number of licenses
on each ranch is determined by negotiations between the landowner and
CPW. Currently, 29 ranches provide over 1.2 million acres of hunting access to
Colorado residents who draw one of these limited licenses.
(cpw.state.co.us/thingstodo/Pages/RFW.aspx)

© CPW

Ranching for Wildlife

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Invasive Species Fact Sheet:
Waterfleas
Bythotrephes longimanus (Spiny), Ceropagis pengoi (Fishhook), Daphnia lumholtzi
Waterfleas are zooplankton aquatic crustaceans that have a
jumpy or jerky mode of swimming. The Daphnia waterflea was
introduced from the aquarium trade and fish stocking. They are
native to Africa, Asia and Australia. Like invasive mussels, the
Bythotrephes and Ceropagis were introduced into the Great Lakes
from ships' ballast water coming from Eurasia. Unlike the fleas
dogs are known to carry, waterfleas are very different. They do not
live outside the water, and do not bite or harm people or pets.
Locations in Colorado:
• Bythotrephes longimanus (spiny): no known
• Ceropagis pengoi (fishhook): no known
• Daphnia lumholtzi: Chatfield State Park, Douglas Reservoir,
John Martin State Park, Navajo State Park, Pueblo Lake State
Park and Pueblo State Hatchery
Identification:
• Unique body shape: distinguished from other zooplankton by
its long tail (70% of body is tail)
• Depending on age – the spine may contain 1-4 barbs (the
older the water flea the more barbs)
• Head is mostly a single large black eye
• Swimming antennae propels flea through the water, allowing
travel between shallow and deeper waters.
• Range in length from 0.4 to 1.6 mm, depending on sex (females are larger) and age
Biology:
• Found mostly in temperate freshwater lakes, can tolerate
brackish water
• Most abundant in the summer and fall
• Can tolerate temps between 4º-30º C (39º-86ºF) and .04 to
8% salinity
• Both sexual and asexual life cycles, depending on environmental conditions
• Less than two weeks per generation with asexual life cycle
• Resting eggs may lie dormant for long periods of time, allowing the spread of these exotics into different bodies of water
• Resting eggs may also pass, unharmed, through the digestive
tracts of fish
Bythotrephes longimanus (Spiny)

Pathway of introduction and
spread:
• Eggs and adults are easily transported
in: ballast tanks, bilge water, bait buckets,
live wells, and on fishing lines, anchor lines, and nets
• It only takes one adult or egg to start an infestation
• If female waterfleas die out of water, under certain conditions
they produce eggs that resist drying and freezing, which can
establish a new infestation later
Negative impacts:
• Outcompete native juvenile fish for food, causing low survival rates, and because barbs stick in throat, are unpalatable
to juvenile fish
• Avoid predation by larger fish by retreating to deeper waters
during the day (10-20m) where they are less visible and
ascending (0-10m) at night where food is abundant and
temperatures higher, increasing metabolism and growth rates
• Their long spines can cause them to become entangled on
fishing lines and can clog eyelets of fishing rods
Management:
Once waterfleas are established, they are almost impossible to eradicate. Educating the public and following state watercraft inspection
and decontamination procedures are the best methods to contain
current infestations and prevent further spread to new waters.
Prevention:
Preventing the spread of invasive waterfleas is the best strategy;
here is what you can do:
• CLEAN all plants, animals and mud from your fishing gear,
boat, trailer, and equipment before leaving the water. Clean
your boat, tackle, downriggers, trailer, waders, etc. with hot
water (above 120ºF) when you get home
• DRAIN live wells, ballast and bilge water before you leave the
water body
• DRY boats and equipment to dry completely before launching in other waters
• DISPOSE of unused bait in the trash, not in the water

Ceropagis pengoi (Fishhook)

Images courtesy of Michigan Dept. of Environmental Quality

Daphnia lumholtzi

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-7295 • cpw.state.co.us

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                <text>Colorado Parks and Wildlife</text>
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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

ANS Boat Interdictions

Zebra and Quagga Mussels

Invasive mussels are being transported on watercraft!
Colorado Parks and Wildlife coordinates a broad multijurisdictional watercraft inspection and decontamination
network to protect waters from invasive zebra and quagga
mussels and other invasive species. Recreational watercraft
is the main vector of introduction for this harmful invader.
Colorado is a headwater state and there are no mussels
upstream. State certified inspectors repeatedly intercept
watercraft infested with mussels.
A total of 281 boats with attached adult zebra or quagga
mussels have been intercepted coming into Colorado’s
waters from out of state at watercraft inspection and

decontamination stations. The infested watercraft were
intercepted at Barr Lake, Blue Mesa, Boulder Marine, Boulder
Reservoir, Boyd Lake, Canon Marine, Carter, Cherry Creek,
Chatfield, Clear Creek, Crawford, Denver CPW Office,
Dillon, Electra Lake, Eleven Mile, Frisco Bay Marina, Granby,
Grand Lake, Grand Junction CPW Office, Great Lakes
Marine, Green Mountain, Highline, Horsetooth, Jackson,
John Martin, Lathrop, McPhee, Navajo, North Sterling,
Pueblo, Ridgway, Rifle Gap, Roadside (SW Colorado),
Ruedi, Shadow Mountain, Spinney Mountain, Stagecoach,
Steamboat Lake, Strontia Springs, Sweitzer, Taylor Park,
Trinidad, Turquoise, Vallecito and Williams Fork.

Zebra and Quagga Mussel Positive Interdiction Sites in Colorado

cpw.state.co.us

�ZEBRA MUSSELS

Where are the infested mussel boats coming from?
It is often difficult to determine the exact location of infestation
due to the frequency of boating use. Also, numerous
interceptions are used boat purchases in which the previous
boating history is not known. For these reasons, the source
of infestation for more than twenty interdicted watercraft is

unknown. The remainder of the infested vessels were coming
from Arizona, Arkansas, California, Illinois, Indiana, Iowa,
Kansas, Michigan, Minnesota, Mississippi, Missouri, New York,
Nevada (Lake Mead), Oklahoma, Ohio, Pennsylvania, Texas,
Utah (Lake Powell), Wisconsin, and the Great Lakes.

Boat Origin for Zebra and Quagga Mussel Positive Interdictions in Colorado

COLORADO PARKS &amp; WILDLIFE

6060 Broadway, Denver, CO 80216

(303) 291-7295 • (303) 297-1192

cpw.state.co.us
1/2020

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

ANS Fact Sheet

AQUATIC NUISANCE SPECIES (ANS) Program OVERVIEW

Background

The State Aquatic Nuisance Species (ANS) Act was signed into law
May 2008. The Act defines ANS as exotic or nonnative aquatic wildlife
or any plant species that have been determined to pose a significant
threat to the aquatic resources or water infrastructure of the state.
The Parks Board passed regulations required by the Act on February
20, 2009 and updated them since that time. The regulations require
mandatory watercraft inspection, and if necessary, decontamination
of all boats coming from out of state, leaving waters with known ANS
and boats entering high-risk waters where inspections are required
by the managing entity. The focus of the program is to prevent zebra
and quagga mussels and other ANS from infesting Colorado’s water
resources and threatening our water storage and distribution systems
for municipal, industrial and agricultural use. The Colorado ANS
Program is highly effective and a model which other states across the
nation are learning from.

Program Goal and Successes

The goal of the program is to protect the state’s natural resources,
outdoor recreation and water supply infrastructure through the
prevention of new introductions of costly invasive species, such as
zebra or quagga mussels, in Colorado. Western states such as Arizona,
Kansas, Nebraska, Oklahoma, South Dakota, and Texas, do not
have mandatory ANS prevention programs and continue to become
infested with zebra or quagga
mussels. Colorado has prevented
the introduction of these awful
invasive species due to the diligent
efforts of watercraft inspection
and decontamination staff, as well
as monitoring, education and
enforcement actions.
There has never been an adult
zebra or quagga mussel found in
Colorado. Pueblo Reservoir was
de-listed in January 2017 following
5 years of negative detections. All
other reservoirs that initially tested
positive in 2008 were de-listed
in 2014 after 5 years of negative
testing. CPW’s ANS Program
has worked to stop the continued
inoculation of our waters to
invasive mussels being introduced
by recreational watercraft.

Sampling and Monitoring

Sampling and monitoring is a key component to the success of the ANS
Program. CPW has sampled 584 “at-risk” waters over the last decade and
it was through this sampling program that invasive mussel larvae were first
detected in Colorado. While CPW ANS staff monitors the state’s public
waters for numerous invasive plant and animal species, the focus of sampling
is on early detection of zebra and quagga mussels. There are three sampling
protocols that target the three life cycles of mussels. CPW also documents
native aquatic plants, mollusks and crayfish while performing monitoring
activities for invasive species.

Colora
d o Pa r
ks &amp; Wi
ldlife

Protect Colorad
o’s
Lands and Wate
rs
Learn How You

Can Stop InvaSIve

SpeCIeS

Information and Outreach

CPW and its partner agencies have
implemented a comprehensive, multifaceted invasive species public education
campaign. Accomplishments include
distribution of tens of thousands of
printed rack cards, brochures, handouts,
DVDs and posted signage at offices,
boat ramps and other public access
points. Additionally, a media relations campaign has been
launched using web, radio, print and television.

www.cpw.state.co.u

s

Working Together: Watercraft Inspection
and Decontamination (WID)
Watercraft inspection and decontamination is a requirement of the
ANS Act and continues to be a key component in preventing the spread
of ANS into and within
Colorado. CPW coordinates the
vast network of WID stations
that are operated by CPW, the
National Park Service, Larimer
County, various municipalities
and private industry locations.
In total, the state has collectively
performed over 4.9 million
inspections and 119,814
decontaminations since 2008.

cpw.state.co.us

�2018 Water Resources Development Act

The 2018 Water Resources Development Act (S. 3021) was passed by the 115th
Congress and signed into law last October. Section 1170 includes a provision
which directs the U.S. Army Corps of Engineers to establish, operate, and
maintain new or existing watercraft inspection stations to prevent the spread
of aquatic invasive species in the Columbia, Upper Missouri, Upper Colorado,
South Platte and Arizona (should be Arkansas) River Basins. The provision also
authorizes the Army Corps to assist states with monitoring and rapid response
efforts in the case of an infestation of quagga or zebra mussels.

Green Mountain Reservoir

In August 2017, the U.S. Bureau of Reclamation (BOR) detected quagga mussels
in Green Mountain Reservoir. The specimen were genetically confirmed to be
quaggas by an independent lab for CPW. Rapid Response was initiated by CPW
and began immediately among the multi-jurisdictional partnership. Green
Mountain is now listed as “suspect” for quagga mussels and containment is
ongoing. Resources are needed for infrastructure, security and staffing to prevent
the spread through watercraft. Green Mountain Reservoir within Summit
County is owned by the BOR as part of the Upper Colorado River Collection
System, and is managed for recreation by the White River National Forest.
There were no detections in either 2018 or 2019. If there are no detections in
2020, the reservoir will be de-listed in 2021 per regional standards.

Mussel Boat Interceptions

Infested mussel boat interceptions continue to skyrocket each year. In total
there have been 281 watercraft intercepted with adult zebra or quagga mussels
attached since the ANS program began. In 2019, CPW intercepted 86 watercraft
infested with zebra or quagga mussels coming in from out of state. In 2018, the
state intercepted 51 and in 2017 the state intercepted 26 infested watercraft.
This exponential growth in infested boat interceptions is directly related to the
growing threat invasive species pose to water infrastructure, natural resources
and outdoor recreation. The majority of the intercepted vessels were coming
from Lake Powell, the State of Arizona or the Great Lakes. All boats were
decontaminated to ensure all mussels were dead, and no mussels were visibly
attached.

An administrative solution in both the House and Senate is needed to correct
the typo is Section 1170 from the nonexistent Arizona River Basin to the Senate’s
intent of providing resources to the Arizona River Basin with its headwaters in
Colorado. Urge the Army Corp to not delay appropriations or planning in the
Upper Colorado or South Platte Basins while the Arizona/Arkansas Basin typo
is corrected.
Support is needed to gain the fiscal appropriations as authorized in the bill.

The Stop the Spread of Invasive Mussels Act of 2019

Senator Bennet, along with Senator Daines and Senator Tester from
Montana, introduced the Stop the Spread of Invasive Mussels Act of 2019.
The bill authorizes the Secretary of Interior through the Commissioner of
Reclamation to provide financial assistance to states or local governments
for WID. The bill adds several federal agencies, such as the National Park
Service, as a member of the ANS Task Force and provides authorization for
all members of the ANS Task Force to conduct WID into and out of federal
waters. The bill requires the ANS Task Force to report back on any regulatory
changes necessary to implement WID prevention or containment. Finally,
the bill corrects the typo in Section 1170 of S.3021 to authorize the Arkansas
River Basin. Sponsorship and support is needed to pass this bill.

The threat of invasion from
zebra and quagga mussels
is greater than ever due to
numerous new infestations
in surrounding states such as
Arizona, Kansas, Nebraska,
North Dakota, Oklahoma,
South Dakota, Texas and Utah.

Operating and Financials

The Supreme Court ruling in case 13SC996 significantly reduced the source fund
for the ANS Program (Tier II Severance Tax) as appropriated in the ANS Act.
A broad stakeholder effort was called upon to raise funds and determine long
term solutions.
CPW redirected agency cash and a USFWS Motorboat Colorado Grant to pay
for the 2017 boating season, along with over $1M of partnership funds. The
Colorado General Assembly provided funding to the ANS Program through
SB17-259 and HB18-1338 to be leveraged with federal dollars and other sources
to maintain the program over the 2018–2019 boating seasons.
In 2018, the Colorado General Assembly passed the Mussel Free Colorado Act
(HB18-1008) to provide a new stable funding source through the creation of an
ANS stamp for motorized boats and sailboats—$25 for in state residents and
$50 for out of state visitors. The ANS Stamp will provide approximately $2.4M
(50% of the need) annual funding to the ANS Program. The bill also increases
fines for violations and allows CPW to recoup costs for decontamination of
quarantined and impounded vessels.
Similar to the 2017 joint resolution on ANS which passed the State Legislature
unanimously, the Mussel Free Colorado Act continues to encourage federal
governments, specifically the Bureau of Reclamation, U.S. Forest Service and
Army Corp of Engineers, to provide the other half of the needed funds for the
ANS Program since many of the highest risk waters are federally owned or
managed reservoirs.

How You Can Help

• Contact U.S. Senators and Representatives, and the U.S. Army Corp
of Engineers, and encourage them to secure an administrative solution to
correct the typo in Section 1170 of S. 3021 (WRDA-2018) changing Arizona
River Basin, which does not exist, to Arkansas River Basin, which was the
intent of the Senate.
• Encourage Congressional delegates and the Army Corp of Engineers to
provide fiscal appropriations under Section 1170 of S. 3021 for the Upper
Colorado, South Platte and Arkansas River Basins. This funding will be used
to provide cost share for WID staff, operations, infrastructure improvements,
monitoring and rapid response.
• Support the passage of The Stop the Spread of Invasive Mussels Act of 2019.
• Urge the U.S. Bureau of Reclamation and the U.S. Forest Service to provide
funding to CPW for WID staff, operations and improved infrastructure on
waters under their ownership or recreational management.
• Inform municipal government, county commissioners, state legislators and
federal congressional delegates of
the potential economic and social
impacts that could occur without
ANS preventative measures.

COLORADO PARKS &amp; WILDLIFE       6060 Broadway, Denver, CO 80216       (303) 291-7295  •  (303) 297-1192       cpw.state.co.us
1/2020

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Boreal Toad (S. Rocky Mtn. Population)
ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

appears to be the
main cause of
declines, especially in the Southern
Rocky Mountain population.

© BRAD LAMBERT, COLORADO NATURAL HERITAGE PROGRAM

Species Distribution

Range
Historically, the Southern Rocky
Mountain population of boreal toads
occurred in Colorado, New Mexico,
and Wyoming. Once considered
common, boreal toad populations
declined sharply in Colorado and
Wyoming and have not been found in
New Mexico since the early 1980s.

Boreal toads (Anaxyrus boreas, Family Bufonidae) lay strands of several thousand eggs in
shallow ponds. The eggs hatch after two weeks, and tadpoles remain for 60–90 days.

Species Description

Identification
With bumpy green-to-brown skin
(occasionally rust or gray) and a thin
white stripe along the back, boreal
toads look similar to Woodhouse’s
toads. Boreal toads occur mostly above
8,000 feet in elevation, whereas Woodhouse’s toads occur mostly below 8,000
feet. Females grow up to 4.3 inches,
and males grow up to 3.7 inches.
Boreal toads do not have a vocal sac;
males emit only a very soft chirp.
Preferred Habitats
Boreal toads occupy numerous wetland types (emergent marshes, kettle
ponds, beaver ponds, streams, lakes,
reservoirs, and wet meadows), often
surrounded by coniferous forests. Boreal toads travel up to 1.5 miles from
their breeding and summer ranges to
hibernate in microhabitats that do not
freeze, including under logs and rocks,

in rodent burrows, chambers beneath
large boulders, and beaver lodges.
Diet
Adult boreal toads consume a wide
variety of insects, spiders, and snails.
Tadpoles eat organic substances
suspended in the water column or on
bottom.
Conservation Status
Federal: Not listed.
Colorado: Listed as Endangered,
designated Tier 1 Species of Greatest
Conservation Need.
BLM: Listed as Sensitive Species.
USFS: Listed as Sensitive Species.
Boreal toad populations have declined
for numerous reasons, including
diseases, environmental stress to the
immune system, ultraviolet radiation,
environmental toxins, altered
hydrology, and habitat disturbance.
Disease caused by chytrid fungus

Distribution of the Southern Rocky Mountain population of the boreal toad by watershed throughout the
range and by county within Colorado. Based on data
from Colorado Parks and Wildlife and Colorado Natural
Heritage Program.
Version Date: November 2020

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Distance to upland foraging and
hibernacula

within 1.5 miles

Elevation

8,000–11,500 feet

Shoreline

gradually sloping shallows with temperature
gradient for both egg and tadpole development

Sunlight availability

open and sunny along water edges

Vegetation in water

open or sparse along shore

Water depth

6–12 inches

Water persistence

May through late September best, but growth
and development is temperature-dependent

Water temperature

59–70 °F

© P. S. CORN, USGS N. ROCKY MTN. SCIENCE CENTER

Preferred Habitat Conditions

Acknowledgements
Bradley Lambert (Colorado Natural Heritage
Program, Fort Collins, CO) and Harry Crockett
(Colorado Parks and Wildlife, Fort Collins, CO)
reviewed earlier versions and provided input on
preferred habitat conditions.

Management Recommendations

This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. Biologists with
expertise in boreal toads have suggested numerous practical steps that can be taken to
improve habitat quality for this species.

Suggested Reading and Citations
Campbell, J. B. 1970. Hibernacula of a population
of Bufo boreas in the Colorado Front Range.
Herpetologica 26:278-282.
CPW (Colorado Parks and Wildlife). 2015. State
Wildlife Action Plan. Colorado Parks and
Wildlife, Denver, CO.

Hydrology
• Maintain natural hydrologic conditions that support boreal toad habitat.
• Maintain water level at one foot or more with gentle sloping banks that also
provide water six inches deep.
• Avoid diverting water from occupied sites.
• Construct shallow shorelines in new impoundments.

Gerber, B. D., S. J. Converse, E. Muths, H. J.
Crockett, B. A. Mosher, and L. L. Bailey. 2017.
Identifying species conservation strategies
to reduce disease-associated declines.
Conservation Letters July:1-10.
Hammerson, G. A. 1999. Amphibians and Reptiles
in Colorado. University Press of Colorado and
Colorado Division of Wildlife.

Vegetation
• Keep open along at least partial shoreline, allowing sunlight to penetrate.

Jackson, T. 2008. Report on the status and
conservation of the boreal toad Bufo boreas
boreas in the Southern Rocky Mountains
2006-2007. Colorado Division of Wildlife,
Denver, CO.

Contamination
• Do not apply pesticides or herbicides near breeding ponds.
• Reduce heavy metals and other toxins.
• Avoid spreading contamination of the chytrid fungus by not introducing water or
mud from one infected wetland to another.
• Decontaminate boots, boats, and other equipment.

Keinath, D., and M. McGee. 2005. Boreal Toad
(Bufo boreas boreas): a technical conservation
assessment. [Online]. USDA Forest Service,
Rocky Mountain Region. Available: http://
www.fs.fed.us/r2/projects/scp/assessments/
borealtoad.pdf.
Loeffler, C. (Ed.). 2001. Conservation plan and
agreement for the management and recovery
of the southern Rocky Mountain population
of the boreal toad (Bufo boreas boreas), Boreal
Toad Recovery Team. 76 pp. + appendices.

Land Use
• If timbering in adjacent forests, create diverse age structure.
• Restrict livestock in breeding ponds.
• Consider timbering activities that preserve boreal toad habitat.
• Avoid building new roads adjacent to boreal toad habitat.

U. S. Fish and Wildlife Service. Date unspecified.
Species Status Assessment Report for the
Eastern Population of the Boreal Toad,
Anaxyrus boreas. Available: https://www.fws.
gov/mountain-prairie/es/species/amphibians/
borealtoad/Boreal%20toad%20SSA%20
Report%20(1).pdf

© BRAD LAMBERT, CNHP

© BRAD LAMBERT, CNHP

© BILL BATTAGLIN, USGS

Conservation
• Protect suitable habitat.
• Obtain required permits from CPW before stocking any fish, and consider
stocking only native species.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Boreal Toad (v. Nov 2020)

Assessment of habitat before and after restoration or management actions
Project Name: ______________________________________ Project Area (acres): __________ Habitat Area (acres): __________
Size of Contiguous Habitat outside Project Area (acres): ________ Ownership (circle): Same / Different / Conservation Easement
Scorecard Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the
numbers in the value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If
condition is outside range or is not described, enter a zero.
Project Area and Habitat Area: The project area includes the entire area affected by the project. The habitat is the area that will
provide (in case of pre-project) or does provide (post-project) habitat for each potential target species within the project area. The
habitat area may be the same size as the project area or it might be smaller and it may be defined differently for different target
species. If there is contiguous habitat area outside the project area, note the size and whether the ownership of the contiguous areas
is the same or different and whether it is under conservation easement or other habitat protection. If the habitat area within your
project area is noncontiguous and/or if sections are in very different conditions, consider using multiple scorecards so that each
scorecard represents the general conditions. If you use multiple scorecards, identify each habitat area on a map.

Key habitat variable and conditions

PreProject

Value

Date of assessment
Edge of water and water depth
Gradually sloping shallows with water 6–12 inches deep
&gt;3–6 inches or &gt;12–18 inches
1–3 inches

26.0
17.3
8.7

Sunlight availability
Open, sunny along water edges
Some shade, but more sun than shade
More shade than sun

26.0
17.3
8.7

Water persistence
Present from beginning of May through end of September
Present end of May through end of August
Present mid-June through mid-August

26.0
17.3
8.7

Vegetation in water
Open or sparse along shore
Thick enough to produce some shade
Thick enough to allow only some sunlight
Total (of 100 possible): add all numbers in before or after columns

22.1
14.7
7.4

Expected
PostProject

Actual
PostProject

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

2014
Fact Sheet
A REVIEW OF STATEWIDE RECREATION AND CONSERVATION PROGRAMS

Colorado Parks and Wildlife:
Motion and Momentum

© DENNIS MCKINNEY

© CPW

© CPW

Bob Broscheid, Director
Colorado Parks and Wildlife

© CPW

Colorado Parks and Wildlife is a national leader in outdoor recreation and
wildlife management. In 2013, we hosted millions of visitor days at 42 amazing
state parks, including the brand new Staunton State Park which opened in May.
In 2013, we put millions of hunters and anglers into the field across the state. We
registered thousands of boats, snowmobiles and off-highway vehicles. We put
tracking collars on sage-grouse chicks, researched black bear use of residential
areas, followed radio-collared mountain lions on the Front Range, stocked
millions of fish statewide, and managed the more than 900 species of wildlife
everyone likes to see as part of this great state. We did it all as an enterprise
agency that receives no annual funding from the state’s general tax fund.
Colorado Parks and Wildlife enters 2014 with a feeling of pride for our past
successes. We appreciate the support of Coloradans, the legislature, the
outdoorsmen and the outdoorswomen, the adventurers, the visitors and the
fans of our amazing natural resources. We say “thank you” to the people who
supported the renewal of the Colorado Habitat Stamp in 2013. We extend our
deepest appreciation for the thousands of volunteers who have given hundreds
of thousands of hours to support our mission. We say “thanks” to everyone who
bought a parks pass, a hunting license, a fishing license or registered their boat,
snowmobile or OHV. We say “congratulations” to the thousands of people who
completed a hunter education class, a boater safety class, learned to camp in
the “Let’s Camp” program, or participated in one of the hundreds of outdoor
recreation seminars offered last year. To those who didn’t, we say “come join us”
and see all that makes Colorado amazing.
We are optimistic, yet we are realistic. There are many things on the horizon in
2014 that will require continued support from those who care about what we
do at Colorado Parks and Wildlife. We recently wrapped up our “Path Forward”
merger report to the legislature and we already have our sites set on a new, more
comprehensive strategic plan to guide our agency forward. We’ll keep working
to assure that Colorado’s native species avoid expensive federal regulatory
intervention. We’ll embark on the process of setting the big game hunting season
structure for 2015-2019. We will work to make sure that recreation and wildlife
are rightly considered as Colorado undertakes water strategy planning. We will
work to implement the strategies of the 2014 Statewide Comprehensive Outdoor
Recreation Plan (SCORP). There is work to be done and Colorado Parks and
Wildlife is in motion. Come join us.

cpw.state.co.us

�Colorado Parks &amp; Wildlife Facts and Figures
Visitation and Participation, FY 12-13
Percent of Colorado residents who visited State Parks		
Most popular activities when visiting a state park		
			

Total Hunting and Fishing License Sales		
Resident
Non-Resident

Hunting Licenses
361,387
106,032

Fishing Licenses
634,126
330,803

11,501,520

CPW Employees (FY 12-13)

1,506, 686

Volunteers (Calendar Year 2012)

46%¹
Hiking, Fishing,
Camping, Boating
Combination
74,338
–

1 Source: 2008 State Parks Market Assessment Study (Corona Research)

Authorized FTE
Temporary Employees

© CPW

State Park Visitation			

Employees and Volunteers

Total volunteer hours
Overall number of volunteers
Average hours per volunteer
Overall Value

(based on 2012 Independent Sector average hourly rate)

909.5
1,077
337,235
6,511
52
$7,466,382

2013 Statewide Outdoor Recreation Activity Days2

2 Source: 2014 Statewide Comprehensive Outdoor Recreation Plan (SCORP)

222,589,592
37,328,022
26,411,408
14,712,057
14,456,827
14,158,319
6,474,549
6,091,660
2,964,683

© MATT INDEN/MILES

Trail recreation – Non-motorized: hiking, walking, biking
Boating and water-based recreation – excluding fishing
Fishing
Motorized recreation
Wildlife Viewing, including bird watching
Camping – Tent camping (not backpacking)
RV Camping (responses for “developed and RV camping”)
Big Game Hunting
Small Game/Upland Bird Hunting

© CPW

Activity Days (by Colorado Residents)

Colorado Parks &amp; Wildlife Revenue Sources
Total Revenue $198,712,410 (FY 12-13)

40%
License and other
wildlife fees:
$80,203,643

16%
Federal and State
Grants:
$32,863,135

14%

22%

Park Entrance, camping
fees, reservations and
registrations:
$27,420,973

Lottery and Great
Outdoors Colorado:
$43,964,394

5%
Severance Tax and
General Fund (no GF in
FY 11-12 or FY 12-13):
$7,814,588

3%
Sales of goods, donations,
interest and other revenues:
$6,445,679

�Where the outdoors begins

Colorado Parks and Wildlife offers fishing clinics for all ages.
We’ve added archery ranges in Cheyenne Mountain and Barr Lake
State Parks as well as new archery ranges in Basalt and Glenwood
Springs. Our “Let’s Camp” program teaches urban families how
to take the first steps to a lifetime of camping memories. Hunter
education courses lead to mentored first hunts for hundreds of
Coloradans each year. We can teach you to canoe, about the night
sky full of stars, how the bear makes it through the winter and we
can help your kids catch their first fish. But we warn you, once you
really experience Colorado, you’ll be addicted. It’s a mile-highplus rush of WOW that is as close as your nearest state park or
the local CPW office. We’re the healthiest, most active state in the
nation. Come join us on an adventure and find out why.

Acres
220,437

Ownership and Management Partnerships
Colorado State Land Board
Colorado Parks and Wildlife (Fee Title)
U.S. Army Corps of Engineers
U.S. Bureau of Reclamation
Local Governments
Irrigation Companies
U.S. Forest Service/Bureau of Land Management
Other

36.2%
23.5%
11.5%
11.3%
8.0%
6.4%
2.6%
0.5%

Conservation Easements in State Parks (Acres)

4,551

Wildlife Properties, 3rd Party Conservation
Easements and Public Access Areas
345 State Wildlife Areas
76 State Habitat Areas
19 State Fish Hatcheries
State Land Board Public Access
State Land Board Total Surface Control (CPW)

Acres
672,044
155,455
484,471
12,361

Ownership and Management Partnerships
Colorado Parks and Wildlife (Fee Title)
Leasehold Interests (excluding SLB leases)
Conservation Easement
3rd Party Conservation Easement
Public Access/ROW Easement

Conservation Easements in Wildlife Areas
(Held by 3rd parties; Funded by CPW)

Walk-in Hunting Access Programs
Small Game Walk-In Access (WIA) acres
Big Game Walk-In Access (BGAP) acres
Ranching for Wildlife

37%
18%
18%
18%
9%

Acres
189,432

Acres
250 ,000
147,013
1,200,000

Some cuts will come to important programs. Funding for habitat
acquisition programs is being reduced by $1 million. Seventeen
positions in the agency have been eliminated including engineering,
education, watchable wildlife, and research of species such as mule
deer, native fish and boreal toads.
While no one likes to tighten their belt when tough budget
decisions have to be made, the Colorado Parks and Wildlife
staff made the difficult decisions to recommend a package of
reductions that will make sure that Colorado Parks and Wildlife
can face a brighter future and continue the positive work that is
laid before us.

Partnering for habitat
and recreation

Did you know that much of the most
important wildlife habitat in the state is
on private land? In fact, about 95 percent
of the critical winter range for mule deer
is in private ownership. From pronghorn
to prairie chicken and burrowing owls to badgers, there are many
species that are almost fully dependent on private lands. With the
continued support and cooperation of private landowners - from
farmers and ranchers to energy companies - Colorado Parks and
Wildlife is making positive strides to protect important habitat.

© CPW

42 S� tate Parks
23 Recreational/Water-based Parks
16 Resource-based Parks
3 River Corridor Parks

Financially sound into the future

This year, Colorado Parks and Wildlife will begin reducing agency
expenditures by $9.9 million per year. That’s some tough cuts
across the agency but they’re necessary cuts to make sure that the
agency and its mission are protected for future generations.

CPW Land and Water Acres
Ownership and Management as of 7/1/2013
State Parks Properties

© CPW

Adventure seekers from across the globe flock to Colorado for its
rugged mountains, majestic wildlife and amazing places but we
aren’t just the dream destination for the hard core and extreme
thrill seekers. We are the beginning of the experience for anyone
looking to do more, see more, be more and feel more.

Through the use of conservation easements purchased from
and donated by willing private landowners, Colorado Parks and
Wildlife has protected hundreds of thousands of acres of wildlife
habitat. It has been done in a way that keeps traditional uses like
farming and ranching on the land. Much more habitat has been
protected by agriculture and landowner groups and by individual
landowners who have enhanced habitat through their own efforts.
Programs like the Walk-In Access Program and access easement
opportunities make it possible for private landowners to find new
revenue streams that can make a difference to a family’s bottom
line. From a listing on the Colorado Birding Trail to participation
in a wildlife viewing event, many private landowners are also
learning the financial rewards that come with being part of the
state’s $2.2 billion annual watchable wildlife industry.

�The Habitat Stamp:
Small stamp, big difference

When the sky opened up and dropped record rainfall in northeastern Colorado in September, Colorado Parks and Wildlife staff
rushed to help. CPW swift water rescue teams deployed to the area
and agency staff, with their boats, trucks and equipment converged
to help with rescue efforts and to protect evacuated neighborhoods.
While all of that occurred, our properties were not without their
own challenges. St. Vrain State Park suffered significant damage
and was closed throughout the fall and winter (expected to reopen
spring 2014). Eldorado Canyon State Park - one of the world’s best
known technical rock climbing areas - lost a main road, picnic
areas and the park was shut down for weeks. Dozens of state wildlife area properties along the South Platte River were damaged and
many were permanently changed as the power of nature pushed
through the countryside.

© CPW

The Colorado Wildlife Habitat
Stamp continues to make a
big difference. As a testament
to that difference, in 2013, the
Colorado legislature approved
a renewal of the program
through 2026. Conservation
easements purchased through
the program have protected
more than 150,000 acres of
important habitat so far in the
state - keeping agricultural
land in the hands of the
private land stewards who
know it best. In addition, the
Habitat Stamp is responsible
for opening up access to some
great places to hunt and fish.

Flood damages parks and properties:

GOCO invests in Colorado

© CPW

Great Outdoors Colorado invests a portion of Colorado Lottery revenues
in parks, trails, wildlife habitats, river corridors, playgrounds and open
spaces in all 64 counties. Voters approved a constitutional amendment
to create GOCO, which is managed by an independent board and uses
no tax dollars. Half of GOCO funds, or more than $400 million to date,
is dedicated to Colorado Parks and Wildlife projects. The list includes
building and enhancing parks, trails and facilities; providing water for
recreation, preserving natural areas, and protecting wildlife species;
funding for Youth Corps
crews, and support for
environmental educational
programs, volunteer programs and youth outreach.
In 2013, CPW opened
FY 12-13
Award
Staunton State Park thanks
Parks
Purpose
$11,553,600
to $10 million in GOCO
Wildlife Purpose
$18,847,618
investments in land acquisition and infrastructure
Lottery (Parks)
$13,563,176
development.

CPW merger brings convenience

© CPW

Colorado Parks and Wildlife was created in 2011 when Colorado
State Parks and the Colorado Division of Wildlife were merged
through legislative action. Today, the merger means more convenience for the customers of the agency. Need a park pass or boat
registration? Now you can do that at 18 more offices around the
state. Need to get a hunting or fishing license? Now you can buy it
at 42 state parks. And there are more conveniences to come in 2014
as a new agency website launches and new programs are offered at
locations across Colorado.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

2015 Fact Sheet
Colorado Parks and Wildlife charts a sustainable path

Colorado Parks and Wildlife (CPW) remains a leader in natural resource management and outdoor recreation.
Funded primarily through user fees, Great Outdoors Colorado (GOCO) grants, and federal and state grants,
CPW strives to ensure efficient and effective use of financial and human resources. Through planning and by
following Colorado's Lean Program, CPW evaluates programs and makes spending adjustments accordingly.
In fiscal year 13-14, CPW invested a portion of its reserve funds to help rebuild habitat and infrastructure after unprecedented flooding. CPW
also provided local grants for trails, improved wildlife habitat, successfully worked with the public to develop a 5-year Big Game Season
Structure and collaborated with partners to provide outdoor experiences for youth and families, to name a small fraction of accomplishments.
Wildlife and outdoor recreation represent an important part of Colorado’s economy and quality of life, accounting for an estimated
$34.5 billion in total economic significance. Hunting, angling, wildlife viewing and state park visitation contributes roughly $6.1 billion in
economic effects statewide. Through ongoing partnerships, strategic planning and wise resource investments, CPW is poised to continue
providing quality wildlife management and outdoor recreation during the coming year.

5%
Sale of goods,
donations, interest
and other revenues:

4%

41%

3%

Severance tax and
General fund¹:

License sales and
other wildlife fees:

Reserve funds:

$8,735,560

$80,248,080

$9,653,867

$4,963,054

15%

Park entrance and
camping fees, reservations
and registrations:

16%

$28,370,985

Lottery and Great
Outdoors Colorado:

16%

$31,596,886

Federal and state grants:

$30,672,308

Total Revenue (FY 13-14)
$194,240,740

¹CPW received $471,773 from GF in
FY13-14 (0.2% of total revenue)

Total Expenditures (FY 13-14)
$194,240,740

9%
Capital improvements/
equipment:

$17,873,872

30%

1%

Operating expenses:

Land and water
acquistion:

© CPW/VERDON TOMAJKO

$56,860,112

$2,659,424

6%

8%

46%

Grants to others:

Payments to other
agencies:

Personal services:

$13,429,600

$14,820,921

$88,596,811

cpw.state.co.us

�Colorado Parks &amp; Wildlife Facts and Figures
Economic Significance of Outdoor Recreation
Total Economic Effect²
All outdoor recreation activities³
$34,505,000,000
Highlighted activities
Fishing⁴
$1,916,000,000
Hunting⁴
$919,000,000
State park recreation⁵
$1,032,000,000
Wildlife watching⁴
$2,280,000,000

CPW Land and Water Acres
Ownership and Management as of 7/1/2014

Jobs
313,000

State Parks Properties

Acres

43 �state parks⁸

16,413
10,882

220,385

Ownership⁹

% of Total Acres

Colorado Parks and Wildlife (Fee Title)
Non-fee title

TBD

19,541

22%
78%

⁸This includes Lone Mesa State Park that is currently open only for limited hunting access.

² This includes estimates of all direct, indirect and induced contributions.
³ This includes a broad set of outdoor recreational activities performed on state, federal, county, local and other lands.
⁴ Source: Economic contributions of Outdoor Recreation in Colorado, 2014, Southwick Associates and CPW.
⁵ �Estimated with visitor expenditure data taken from the 2009 State Parks Market Assessment Study (Corona
Research) and a conservative industry economic multiplier. Jobs supported by state parks has not yet been calculated.

⁹CPW owns about 22% of the total acres. All other land and water managed under agreements with
different government and private organizations.

Conservation Easements in State Parks

© CPW

4,551

(Held by CPW)

Award

Parks purpose:
Wildlife purpose:
Lottery (parks):

$13,300,476
$5,285,059
$13,011,351

© CPW/KEN PAPALEO

FY 13-14

Wildlife Properties, Third-Party Conservation
Easements and Public Access Areas
346 State wildlife areas
79 State habitat areas
19 State fish hatcheries

Conservation Easements in Wildlife Areas

222,589,592

�Boating and water-based recreation excluding fishing

37,328,022

Fishing

26,411,408

Motorized recreation

14,712,057

Wildlife viewing, including bird watching

14,456,827

�Tent camping (not backpacking)

14,158,319

State Land Board public access
State Land Board total surface control (CPW)

Ownership or Management Details

6,474,549

Walk-in Hunting Access Programs

6,091,660

Small game/upland bird hunting

2,964,683

Small Game Walk-In Access (WIA)
Big Game Walk-In Access (BGAP)
Ranching for Wildlife

Total Hunting and Fishing License Sales		
Hunting Licenses
Fishing Licenses
Resident
369,353
668,647
Non-resident
107,273
339,598
⁷ Source: 2009 State Parks Market Assessment Study (Corona Research)
© CPW/KEN PAPALEO

% of Total Acres
45%
22%
22%
10%
1%

Acres
247,000
147,090
1,200,000

© CPW/THOMAS KIMMEL

Visitation and Participation, FY 13-14
State Park Visitation			
Percent of Colorado residents who visited state parks		
Most popular activities when visiting a state park		
			

485,227
12,945

Colorado Parks and Wildlife (fee title)
Perpetual conservation easement
Third-party conservation easement
Leasehold interests (excluding SLB leases)
Perpetual public access/ROW easement

Big game hunting
© CPW/KEN PAPALEO

194,875

State Land Board Access

�RV Camping (and “developed camping”)

⁶ Source: 2014 Statewide Comprehensive Outdoor Recreation Plan (SCORP)

Acres

(Held by third parties; funded by CPW)

2014 Statewide Outdoor Recreation Activity Days⁶
Activity Days (by Colorado Residents)
�Trail recreation- non-motorized: hiking, walking, biking

Acres
678,949
162,306

11,948,406
46%⁷
Hiking, fishing,
camping, boating
1,562,071
Combination
77,200
–

Employees and Volunteers
CPW Employees (FY 13-14)
Authorized Permanent Employees
Temporary employees

886.5
1,632

Volunteers (Calendar Year 2013)
Total volunteer hours
298,150
Overall number of volunteers
6,524
Average hours per volunteer
46
Overall value
$6,600,000
(based on 2013 Independent Sector average hourly rate)
© CPW/THOMAS KIMMEL

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

2016 Fact Sheet
World class recreation and resource management
Economic significance of the outdoors
Accounting for an estimated $34.5 billion annually in total economic impact, wildlife and
outdoor recreation represent important parts of Colorado’s heritage, quality of life and
economy. Angling, hunting, state park visitation and wildlife viewing contribute roughly
$6.1 billion annually in economic effects statewide. Fishing and hunting have a total combined economic effect in Colorado of more than $2.8 billion. Colorado Parks and Wildlife
(CPW) is playing a pivotal roll in the Governor's Colorado the Beautiful initiative.

© CPW/WAYNE D. LEWIS

Parks offer much more than terrific views
Colorado’s 42 state parks protect habitat, provide recreation and many offer hunting and
fishing opportunities. Park rangers, volunteers and partners provide classes on safe boating
practices, beginning hunting and fishing
clinics and wildflower identification to
name a few examples. This past year,
active duty military and veterans could
enter any Colorado state park in August
for free. Over 6,600 people took advantage of this exciting opportunity.

CPW manages the largest elk herd in North America,
with an estimated population of 264,000 animals. This elk
herd provides hunters with healthy locally sourced food.
Additionally people from around the world enjoy watching
elk in their natural habitat. CPW also stocks more than 90
million fish throughout the state from 19 state fish hatcheries
and rearing units. Coloradans have fishing access to over 2,000
natural lakes, 800 reservoirs and 9,500 miles of streams.

© CPW/WAYNE D. LEWIS

Conservation benefits wildlife
and habitat
Through ongoing partnerships with
other conservation groups and working
farms and ranches, strategic planning
and wise resource investments, CPW continues to provide quality wildlife management
and outdoor recreation.

© CPW/VICK SHENDEL

CPW manages over 350 State Wildlife Areas and protects over
900 wildlife species in Colorado. In 2015, CPW protected
31,955 acres of sage-grouse habitat, cooperated on black-footed ferret reintroductions,
stocked native fish and managed distribution
of native fish across the state.
Outdoor recreation, a native priority
The top reasons Coloradans choose to live
here are the state’s clean environment, access
to public lands and outdoor recreation
opportunities and residents’ ability to maintain
a healthy outdoor lifestyle. Over 80 percent of
Coloradans participate in trail-related activities,
making these the most popular forms of
outdoor recreation.

Economic Significance of
Outdoor Recreation
SHOWN IN BILLIONS OF DOLLARS

$

2.2 1.9
$

WILDLIFE 1
WATCHING

$

FISHING

1

1.0 .91
$

STATE PARK 2
RECREATION

HUNTING

1

¹ �Source: Economic contributions of Outdoor Recreation in
Colorado, 2014, Southwick Associates and CPW.
² �Estimated with visitor expenditure data taken from the 2009
State Parks Market Assessment Study (Corona Research) and a
conservative industry economic multiplier.

2014 Statewide Outdoor
Recreation Activity Days³
SHOWN IN MILLIONS OF DAYS

222 37.3

Trail recreationnon-motorized: hiking,
walking, biking

Boating and waterbased recreation
excluding fishing

26.4 14.7
Fishing

Motorized recreation

14.4

14.1

Wildlife viewing,
including bird watching

Tent camping
(not backpacking)

³ �Source: 2014 Statewide Comprehensive Outdoor
Recreation Plan (SCORP)

cpw.state.co.us

�Colorado Parks &amp; Wildlife At-A-Glance
F U N DI NG SOU RC E S : $213.9 *

USE OF F U N DS : $213.9 *

43% $91.8 • Personnel Services

50% $107.4 • Licenses, Passes, Fees and Permits

25% $53.5 • Operating

18% $38.5 • Federal, State Grants and Loans

12% $26.0 • C� apital Equipment/Improvements

17% $36.7 • L� ottery and Great Outdoors Colorado

7% $15.7 • �Grants to Others

7% $13.7 • S� ales, Donations, Interest and Other

6% $13.4 • P� ayments to Other Agencies

4% $8.8 • Registrations

4% $8.2 • R� eserve Fund Increase

4% $8.8 • General Fund and Severance Tax

3% $5.3 • Land and Water Acquisitions

*All figures shown in millions

GOCO and Lottery
FY14-15 Award

Visitation and Participation, FY14-15
State Park Visitation: 12,464,445
Total Hunting and Fishing Licenses Sales: 1,623,522

HUNTING LICENSES

FISHING LICENSES

COMBINATION

Resident

382,320

686,973

80,242

Non-Resident

107,898

366,089

-

CPW Vision Statement
Colorado Parks and Wildlife is a national
leader in wildlife management, conservation,
and sustainable outdoor recreation for
current and future generations.
Employees and Volunteers
Authorized Permanent Employees - 886
Temporary Employees - 1,696
Volunteers: 6,084
Volunteer Hours: 307,080
Volunteer Dollar Equivalent: $7.1 million
FTE Equivalent: 148

Schools in the Outdoors
CPW connects our youngest citizens to
the places they live by connecting students
and teachers to outdoor learning resources.
We are committed to extending learning
beyond the classroom through:
• �Schools and Outdoor Learning
Environments Program (SOLE)
• �Outdoor Understanding for Teachers
(OUT), Teaching Environmental Science
Naturally (TEN) and other teacher
professional development opportunities
• �National Archery in the Schools Program
• �Field trips to Colorado state parks and
state fish hatcheries
For more information on CPW financial
sustainability please visit
cpw.state.co.us/financialsustainability

© CPW

Working Together
Connecting all Coloradans to the outdoors
requires a network of committed volunteers
and partner organizations working together.
Without our 6,000 volunteers, hundreds of
partner organizations and friends groups
working alongside agency staff, our education
and outreach programs would only be able to
accomplish a fraction of what we do every day.
We engage our partners and volunteers by:
• �Hosting the annual Partners in the
Outdoors conference
• �Providing hundreds of opportunities for
volunteer projects
• �Training volunteers to represent CPW as
certified instructors
• �Inviting volunteers to represent CPW at
community events and in state parks

Parks Purpose - $17,810,153
Wildlife Purpose - $11,800,000
Lottery - $12,800,000

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman St, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us
CPW-2016-1/29/16

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

2017
Fact Sheet
A REVIEW OF STATEWIDE RECREATION AND CONSERVATION PROGRAMS
“Price is what you pay. Value is what you get.” — Warren Buffett.
Sportsmen and women provide 80 percent of Colorado Parks and
Wildlife’s (CPW) funding for wildlife through hunting, fishing and
recreational shooting. As Warren Buffett might say, the price of buying
a hunting or fishing license corresponds to the value received. CPW
finds it increasingly difficult to provide the value sportsmen expect for
the prices being paid. The last price increase for resident licenses was
in 2005. Since that increase, the costs of doing business have climbed
while revenues from resident licenses have remained unchanged. We
cut $40 million from the wildlife budget and defunded 50 positions in
order to keep our costs down. Without increased revenue, CPW will
have to continue scaling back wildlife management and conservation in
Colorado, impacting wildlife and the quality of outdoor recreation.
That’s why in 2016, CPW held 18 public meetings across the state asking for input on the price of resident hunting and fishing licenses. More
than 85 percent of meeting attendees voiced support for a resident
license fee increase. Future conversations between CPW and sportsmen
and women will focus on how to best fulfill CPW’s mission to perpetuate the state’s wildlife resources.
By state and federal law, funding sources for wildlife are kept completely
separate from parks funding. The cost of doing business in state parks is
also rising. The Colorado Parks and Wildlife Commission is authorized
to adjust park pass and camping fee prices but that authority is constrained by a statute that limits the amount of total revenue parks fees
can generate. An increase in the sales of parks passes caused revenues to
meet that cap. As a result, even though state parks are facing rising costs
and increasing pressure to accommodate more visitors, the Commission cannot raise fees to cover these costs.
Commission authority to set park fees is set through the legislative process. CPW is communicating with our partners about the situation and
discussing opportunities for the future. Preserving the Commission’s
authority to set fees puts CPW in a better position if we need to bring in
more revenue to meet demand and keep pace with the rate of inflation.

Bob D. Broscheid, Director
Colorado Parks and Wildlife

Charlie Meyers State Wildlife Area
Roxborough State Park

Bull elk

cpw.state.co.us

PHOTOS BY © WAYNE D. LEWIS/CPW

Let’s keep working together to overcome financial challenges and conserve Colorado’s beautiful places and wildlife.

�CPW Adopts New Principles on Conservation and Outdoor Recreation
These principles provide an opportunity to unite Coloradans on the
importance of our natural resources, and our responsibility to one
another in promoting and protecting them. Colorado became the first
state to adopt the principles at the November meeting of the Parks and
Wildlife Commission.

5.

6.

We believe the uniquely American public-land heritage is a privilege
and a birthright, and Colorado’s abundant open space and outdoor
recreation opportunities contribute to our quality of life and economic
vitality. Combined with the North American Model of Wildlife
Management and private-land conservation, Coloradans and our
visitors enjoy spectacular landscapes in which to work, play and live.

2.
3.

4.

Outdoor recreation and conservation require that a diversity of
lands and waters be publicly owned, available for public access
and cared for properly.
Within Colorado’s diversity of land and waters, private land
provides a balance of conservation and access for outdoor
recreation and conservation of landscapes.
Both recreation and conservation are needed to sustain
Colorado’s quality of life. Both are beneficial to local economic
well-being, for personal health and for sustaining Colorado’s
natural resources
All recreation has impact. Coloradans have an obligation to
minimize these impacts across the places they recreate and the
larger landscape through ethical outdoor behavior.

Parks have implemented
numerous cost-saving
measures to stretch
existing budgets, such as
installing energy efficient
lighting and compacting trash to reduce waste
disposal costs.

© WAYNE D. LEWIS/CPW

1.

7.

Proactive management solutions, combined with public
education, is necessary to care for land, water and wildlife, and
to provide the protections needed to maintain quality recreation
opportunities.
Physical, biological and social science must inform the
management of outdoor recreation.
Stable, long-term, and diverse funding sources are essential to
protect the environment and support outdoor recreation.

Did you know? The CPW outdoor recreation annual impact in Colorado is $6.1 billion.

$1
Billion

$919
Million

Hunting and wildlife viewing are the only recreational activities that benefit every county in the state — urban and rural.

Legislative Bill Introduced
In order to sustain Colorado state parks and wildlife opportunities, the
legislature is considering a bill that would allow the Parks and Wildlife
Commission the limited authority to address current financial
challenges.
Funding is needed to:
•
•
•
•
•

Repair high-priority dams to ensure public safety and maintain
recreational opportunities.
Address public access for hunting and fishing in Colorado.
Increase efforts to improve wildlife habitat.
Renovate hatcheries in order to continue stocking reservoirs and
streams.
Continue inspection programs to keep Aquatic Nuisance Species
out of Colorado’s waters.

•
•

Maintain and improve park facilities including trails, campgrounds and visitor centers.
Improve outdoor recreation educational opportunities and programs to get more Coloradans outside.

Colorado Parks and Wildlife relies on user fees, NOT general tax
dollars. However, the fees have not kept pace with operational costs
and inflation.
•
•
•
•

Resident hunting and fishing license fees have not increased since
2005.
Park entrance fees have not increased since 2010.
Since 2009, CPW has cut or defunded 50 positions and reduced
$40 million from its wildlife budget.
CPW is facing a forecasted wildlife budget shortfall of $14 million
annually just to maintain current operation.

© WAYNE D. LEWIS/CPW

$1.9
Billion

Hunters

State Park Visitors
© WAYNE D. LEWIS/CPW

© KEN PAPALEO

$2.3
Billion

Anglers

© WAYNE D. LEWIS/CPW

Wildlife Viewers

�Colorado Parks and Wildlife At-A-Glance
Colorado Parks and Wildlife relies on user fees, not general tax dollars,
to pay for wildlife management and state parks. However, those fees
are not keeping pace with rising operational costs and inflation. CPW’s
financial sustainability has reached its limit. Outdoor recreation is vital
to Colorado’s economy.

Hunting contributes $919 million in total economic impact to
Colorado, fishing contributes $1.9 billion and wildlife viewing adds an
additional $2.3 billion. Colorado state parks contribute $1 billion to
Colorado’s economy, attracting more than 12 million visitors each year.

Funding Sources: $212.40*

Use of Funds: $212.40*
43% $92.0 - Personal Services

54% $114.0 - Licenses, Passes, Fees and Permits
19% $39.6 - Lottery and Great Outdoors Colorado

26% $54.6 - Operating

15% $30.8 - Federal and State Grants/Loans

11% $23.8 - Capital Equipment/ Improvements

5% $9.7 - Sales, Donations, Interest and Other

6% $12.9 - Payments to Other Agencies

4% $9.3 - Registrations

6%

4%

5% $11.2 - Reserve Fund Increase

$9.0 - General Fund and Severance Tax

*All figures shown in millions.

2%

Visitation and Participation, FY15-16

$12.6 - Grants to Others

$5.3 - Land and Water Acquisitions

Total Hunting and Fishing Licenses Sales

Cheyenne Mountain
State Park

1,715,057

State Park Visitation

13,615,312
Colorado Parks and Wildife relies on its
permanent and temporary staff, as well as
skilled volunteers, to execute its mission.
Authorized Permanent
Employees. . . . . . . . . . . . . . . . . . . . . . . . . . 886
Temporary Employees . . . . . . . . . . . . 1,708
Volunteers. . . . . . . . . . . . . . . . . . . . . . . . 5,842
Volunteer Hours . . . . . . . . . . . . . . . . 298,884
Volunteer Dollar
Equivalent. . . . . . . . . . . . . . . . . . . $7,041,707
FTE Equivalent. . . . . . . . . . . . . . . . . . . . . . 144

GOCO and Lottery
FY15-16 Awards

Since 1992, Great Outdoors Colorado (GOCO) has
awarded more than $400 million from its share of
Colorado Lottery proceeds to Colorado Parks and
Wildlife projects and programs.
GOCO to Parks Purpose . . . . . . . . . . . $19,011,044
GOCO to Wildlife Purpose. . . . . . . . . $11,800,000
Lottery to State Parks. . . . . . . . . . . . . $14,400,000

Fishing
Licenses
686,973
394,888

© WAYNE D. LEWIS/CPW

Hunting
Licenses
Resident
377,952
Nonresident 109,712

Combination
(Small Game/Fishing)

83,590
—

Steamboat Lake State Park

© WAYNE D. LEWIS/CPW

Employees and Volunteers

© WAYNE D. LEWIS/CPW

Colorado state parks are visited by almost half of Coloradans,
making more than 12 million visits to our state parks each year and
contributing $1 billion to the state economy. Colorado’s population
is projected to grow by more than 2 million people in the next 25
years, putting more pressure on our parks and other natural areas.
Here too there must be a balancing of recreational interests with
conservation of natural resources.

Sportsmen and women
provide 80 percent of
Colorado Parks and
Wildlife’s funding for
wildlife through hunting,
fishing and recreational
shooting.

�Throughout 2016, the team has worked
with landowners on 20 pending
projects, totaling 4,412 acres as well as
2,600 linear feet of water channels. This
work has offered significant technical
assistance on 32 other habitat projects
totaling 18,584 acres.

New State Trails Plan

Colorado Parks and Wildlife will host
its first-ever Outdoor Adventure Expo​
Sept. 23-24 at Cherry Creek State Park.
This free event invites participants for
a taste of the many outdoor recreation
possibilities at CPW’s 41 state parks
and other properties. From biking to
boating, clay pigeons to climbing, the
Outdoor Adventure Expo will provide a
little something for everyone.

Trail use, both motorized and nonmotorized, is the No. 1 recreational
activity in Colorado. Each year, Coloradans access trails more than 250 million
times.Trails are also Colorado’s top outdoor recreational “gateway” activity.
Over the past decade, CPW’s Trails
Program distributed more than $54
million for trail grants, capital improvements that benefit trail use and winter
trail grooming operations. These funds
created job opportunities, especially
through the Youth Corp organizations,
which received about $1.6 million to
build and maintain trails.
Strategic Goals:
1. Provide statewide leadership for trail
recreation in Colorado.
2. Facilitate trail development and management across Colorado.
3. Serve as the advocate for trails.
4. Emphasize responsible stewardship for
all phases of trail development and use.
5. Advance ethical use and cooperation
among trail recreationists.
6. Seek financial sustainability for the
Trails Program.

Discover archery at the Outdoor
Adventure Expo.

© KEN PAPALEO

© WAYNE D. LEWIS/CPW

Water channels have been
enhanced by the Private Lands
Wildlife Biologist Program.

Outdoor Adventure Expo

RECREATION

The Private Lands Wildlife Biologist
Program is a partnership with Bird
Conservancy of the Rockies (BCoR),
Colorado Parks and Wildlife (CPW) and
the Natural Resources Conservation
Service (NRCS). Projects enhanced
2,517 acres and 963 linear feet of
instream channels statewide utilizing
$377,792 in federal Farm Bill funds and
$97,500 in partner funding.

OUTREACH

CONSERVATION

Partnering for Conservation

Conservation

is at the heart of Colorado Parks and Wildlife’s mission. And no other scientific principle is more responsible
for creating Colorado’s enormous abundance of fish, wildlife and its world-class outdoor recreation. Yet, in spite of its importance to our state,
conservation — in the context of wildlife and outdoor recreation management — is a concept that’s often overlooked and misunderstood.

Established in 1897, CPW has a rich history of managing Colorado’s wildlife and recreational resources through conservation science. In fact,
CPW’s own mission statement embodies the primary themes of conservation: “The mission of Colorado Parks and Wildlife is to perpetuate the
wildlife resources of the state, to provide a quality state parks system, and to provide enjoyable and sustainable outdoor recreation opportunities
that educate and inspire current and future generations to serve as active stewards of Colorado’s natural resources.”

The new program is a partnership
between Colorado Parks and Wildlife, the State Library and local public
libraries across Colorado designed to
encourage learning at a state park and
provide a new unique service offered
by the libraries. There are currently 574
backpacks with a free parks pass that
are available for check out from 287
libraries statewide. In 2017, Colorado
state colleges and university libraries
will be included in the program. In
a Dec. 2016 library survey, 53 library districts reported 2,671 checkouts in 2016
and there were 454 current holds at the
time of the survey for those waiting to
check out the program.

EDUCATION

Check Out Colorado
State Parks

OUTREACH

Whether you’re a hunter, angler, hiker, kayaker or just someone who enjoys Colorado’s wildlife and wild places, here are just a few of the things
that Colorado Parks and Wildife is doing to further its mission:

Schools and Outdoor
Learning Environments (SOLE)

The SOLE program started in 2013 as a pilot program through Colorado Parks and
Wildlife by funding from Great Outdoors Colorado (GOCO)​with one simple mission —
­
connect kids to the outdoors. The program is designed to increase interest and
participation in the outdoors. We connect Colorado 4th-grade students and their
families to local environmental education programs and outdoor locations.

SOLE Last Year 2015-16
• 24 participating schools 12 Den-

SOLE 2016-17
• 34 participating schools 16 Den-

•

•

•
•
•
•

ver area and 12 non-Denver area
Colorado schools
1,707 students in program, 69 4th-grade classrooms participated
67 Field Trips
24 Family Nature Nights
36 Partner Organizations
Art Contest: There were three
Grand Prize Winners, sponsored
by Cabela’s in Thornton, Lone Tree
and Grand Junction

•
•
•
•

ver area and 18 non-Denver area
Colorado schools
2,327 students in program, 69 4th-grade classrooms participated
97 Field Trips
32 Family Nature Nights
65 Partner Organizations
Art Contest: Deadline is Apr. 7,
2017

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

2018 Fact Sheet
A REVIEW OF STATEWIDE CONSERVATION AND RECREATION PROGRAMS
Fishing in Sylvan Lake State Park

MIKE DELLIVENERI/CPW

Colorado’s Heartbeat

VIC SCHENDEL/CPW

American white pelican

MIKE DELLIVENERI/CPW

Hunting has a $919 million
impact to Colorado’s economy.

cpw.state.co.us

For more than a century, conservation work
has been the primary mission of Colorado
Parks and Wildlife (CPW). Nationwide, wildlife agencies were created to ensure the prosperity of both game and nongame species.
CPW employees are dedicated professionals
who work passionately for Colorado’ resources every day. And the agency is fortunate to
be supported by dedicated sportsmen and
sportswomen who cherish Colorado’s parks
and wildlife.
Colorado’s natural beauty, land, water and
wildlife are the top priorities for the agency.
We take our responsibility to operate our 41
state parks — each of which protects valuable
landscapes — and to ethically manage wildlife in public trust very seriously. The work
of CPW staff is done with dedication and an
unwavering sense of duty.
Conserving our parks, landscapes, waters
and wildlife is a calling for CPW employees.
These resources provide a sense of place and
purpose for all Coloradans. Not only do we
gaze at these natural wonders with awe, we
also feel their comforting and healing beauty
deep in our soul. When the sun rises on
a meadow and the mist lifts, you can hear
nature’s song — Colorado’s heartbeat.
Our mission underscores the importance of
caring for our land, water and wildlife. These
natural resources are what makes Colorado
special. Not only do these resources perpetuate the intrinsic values of our state’s citizens,
they also support a robust economy and our
vibrant lifestyle. Assuring the sustainability
of our natural resources and the variety of
recreation opportunities they provide is vital
to Colorado’s future.
Hunters and anglers play a major role in sustaining Colorado’s natural resources through
their willingness to pay for conservation
through the purchase of hunting and fishing
licenses. The health and well-being of our

land, water and wildlife could not be accomplished in Colorado without the support and
investment made by these individuals.
Through the support of hunters, anglers, park
users and all Coloradans, CPW is working
on long-term sustainability for our resources.
We cannot predict the future, but are planning for it.
Colorado faces a growing number of challenges as we see unprecedented population
growth, urban sprawl, habitat loss, continued debates over water use and a growing
segment of citizens who are not connected to
nature and its care.
It is now more important than ever to work
together to chart our course in achieving a
shared vision and investment in Colorado.
We need to rise above our differences to meet
the demands of the people and the needs
of wildlife. It is on each of us to work on
largescale conservation, and to invest in the
resources that sustain us.
We want to emerge from the challenges
stronger and equipped with better strategies
to maintain and improve Colorado’s outdoor
future. You have entrusted CPW with a very
important mission. As the outdoor recreation
industry continues to become one of the
nation’s largest economic sectors, we must
focus on the intersection of conservation and
outdoor recreation. We must protect our public lands and ensure the health of our wildlife
resources. Despite the challenges we face,
CPW remains a leader in outdoor recreation
and wildlife management, and we owe a lot of
that success to you.

Bob D. Broscheid
Director, Colorado Parks and Wildlife

�Colorado Parks and Wildlife
Funding At-A-Glance
Colorado Outdoor
Partnership

SOURCE OF FUNDS
$218.7

TOTAL FUNDING

$121.50

LICENSES, PASSES,
FEES &amp; PERMITS

$32.40

FEDERAL, STATE
GRANTS &amp; LOANS

$38.60

LOTTERY &amp; GREAT
OUTDOORS COLORADO

$11.80

SALES, DONATIONS,
INTEREST &amp; OTHER

$9.50

REGISTRATIONS

$4.90

GENERAL FUND &amp;
SEVERANCE TAX

100%
56%

The Colorado Outdoor Partnership (CO-OP) is a
collaborative of leaders across the state of Colorado that
are innovating together to ensure our private and public
lands remain viable to support our diverse wildlife, outdoor
recreation, agricultural heritage and economic well-being.

15%
18%

THE CO-OP HAS COMMITTED TO:

5%

• Promoting the importance and stewardship of public and

private lands and waters to support sustainable habitat
conservation and responsible outdoor recreation.

4%

• Supporting, improving and strengthening public and

2%

private funding to conserve these resources.

THE VISION STATEMENT OF THE CO-OP:

USE OF FUNDS
$218.7

TOTAL BUDGET

$92.40

PERSONAL
SERVICES

$55.10

OPERATING

$19.00

CAPITAL EQUIPMENT/
IMPROVEMENTS

$16.10

GRANTS TO OTHERS

In 2050, Colorado’s people and economy thrive because
of our healthy lands, water, wildlife, working farms and
ranches, and improved hunting, angling and outdoor
recreation opportunities for all.

100%
42%
25%
9%

CPW Launches Integrated Parks
&amp; Wildlife Purchasing System

7%

$13.30

PAYMENTS TO
OTHER AGENCIES

$17.10

RESERVE FUND
INCREASE

4%

$5.70

LAND &amp; WATER
ACQUISITIONS

3%

6%

The new purchasing system sells all CPW
products, including licenses, park passes and state park
camping reservations.

ADVANTAGES OF THE NEW SYSTEM:
• Each customer has a personalized account
• More purchasing options for customers in

ALL $ FIGURES SHOWN IN MILLIONS

•
•
•
•
•

MIKE DELLIVENERI/CPW

Waterton Canyon

•
•
•
•

one central location
Keep your contact and payment information current
Receive receipts, confirmations and special alerts
View your purchase history
Change/adjust camping reservations
Check your preference points, applications and
draw results
Update your license applications up until the
application deadline
Limited Licences applications are now paperless
License payments are due when a customer
draws rather than upfront
24/7 customer service assistance

�DNR Dashboard —
A Guide to Success
The dashboard provides a summary of how
we are doing across key prioritiy areas that
helps fulfill the Governor’s vision for a truly
Health Colorado. Colorado offers numerous
opportunities for outdoor recreation and
serves a growing and diverse population of
people with varied demands for recreational
activities. Outdoor recreation at state parks,
as well as hunting and fishing opportunities,
contribute a combined estimated $6.1
billion to Colorado’s economy. As increasing
numbers of tourists and Coloradans take
advantage of the state’s outdoor resources,
care must be taken to maintain the beauty
and accessibility of state parks and the
activities therein.

GOAL : Increase state park visits. In June
2017, Colorado Parks and Wildlife counted
14.8 million vehicle visits to Colorado’s state
parks over the previous 12 months. This
was a nearly 10% increase over the prior
fiscal year. Based on the last five years, CPW
expects 12.8 million state park visitors in FY
2017–18.
GOAL: Maintain the number of hunting
and fishing participants due to a variety
of favorable outside factors including
weather and healthy herd numbers. CPW
experienced increased participation in 2016
and 2017.

Since 1992, Great Outdoors Colorado
(GOCO) has awarded more than $​550​million
from its share of Colorado Lottery proceeds.

$15,250,585

GOCO TO PARKS PURPOSE

$15,250,585

GOCO TO WILDLIFE PURPOSE

$13,347,498

OUTCOME MEASURE

OUTCOME
BASELINE
(JUNE 2015)

ACTUAL
(JUNE 2016)

ACTUAL
(JUNE 2017)

OUTCOME TARGET

TARGET
DATE

State park visits

12,272,000

13,515490

14,823,913

12,856,071

2017

Hunting &amp; fishing
participants

960,000

1,066,536

1,100,609

960,000

2017

By developing new and maintaining existing
trails for the public while conserving wildlife
habitat through restoration and improvement
projects, Colorado is engaging the next
generation of land stewards and protecting
the abundant natural resources that make
this state so special.

Enhancing land stewardship efforts across
Colorado ensures the sustainability of
millions of acres of land for the enjoyment
of generations to come. Active land
management including agricultural leases,
mineral development and leasing lands for
recreational activities supports Colorado’s
commitment to land stewardship.

​ OCO and
G
Lottery FY16 –17

LOTTERY TO STATE PARKS
(10% DIRECT DISTRIBUTION)

Volunteer FY 16-17
Numbers
Volunteers play an important role for CPW.
They help us get the job done and are a vital
part of the CPW family.

​304,460

VOLUNTEER HOURS

OUTCOME
BASELINE
(JUNE 2016)

ACTUAL
(JUNE 2016)

ACTUAL
(JUNE 2017)

OUTCOME
TARGET

TARGET
DATE

6,174

Funding for the trails

$7.8 MILLION

$7.8 MILLION

$7.8 MILLION

$7.8 MILLION

2018

Number of acres of wetlands
restored, maintained or
enhanced per year

$7,3​49,664.40

2,583

2,583

3,828

6,650

2018

Number of acres of prairie
grassland habitat restored,
maintained or enhanced per year

40,000

40,000

40,000

40,000

2018

Stream miles occupied by native
greenback cutthroat trout

2

2

10

17

2018

OUTCOME MEASURE

TOTAL VOLUNTEERS​

VOLUNTEER VALUE

14​5.8​

FTE EQUIVALENT

DUSTIN DOSKOCIL/CPW

Roxborough State Park

�Bull elk

Colorado’s Archery in the
Schools Program
it is estimated that at least 35,000 kids are
shooting archery in school thanks to CPW
staff. Each region helped fund archery
equipment grants to schools above and
beyond what the statewide archery budget
would allow for. Also, we were fortunate
to have NASP contribute $12,000 in
equipment credits towards our Colorado
schools this year (compared to $7,000 the
previous year).

Colorado
Outdoors Online
Since 1938, Colorado Outdoors magazine
has promoted wildlife-related recreation
and championed the tireless work and
dedication of CPW staff. Launched in
2013, Colorado Outdoors Online, the
magazine’s companion blog, engages
a variety of outdoor recreationists
from hunters and anglers to bikers and
wildlife watchers in dialogue around the
importance of maintaining state lands and
facilities and managing wildlife.
With nearly 200,000 visitors per year,
the blog serves as the ideal platform for
highlighting partner and volunteer efforts,
sharing conservation work and providing
how-to articles to inspire generations
of Coloradans to get outside. Through
story-telling, the blog showcases the
often unseen efforts of the agency. Visit
coloradooutdoorsmag.com to read the
hundreds of stories.

CPW WEB TRAFFIC
DEVICE
CATEGORY

SESSIONS

% NEW
SESSIONS

NEW USERS

MOBILE

3,969,396

48.23%

1,914,261

DESKTOP

3.963,835

45.55%

1,805,442

TABLET

745,250

46.59%

347,197

TOTAL

8,678,481

46.86%

4,066,900

The demand for this program continues
to grow and CPW region and statewide
staff are always willing to step up and
help train teachers to teach archery at
their schools. Each of the 295 schools are
teaching about 120 kids archery each year,

Here’s how the program has expanded this school year as compared to the year before:
CATEGORY

2016–17#

2017–18#

% INCREASE IN ONE YEAR

Equitment grant funds awarded
(Including nasp* funds)

$27,000

$42,800

59%

Maintenance grants funded

12

23

92%

New program grants funded

9

17

89%

Colorado schools in program
(Required reporting nationally implemented in spring 2017)

189

295

56%

Basic archery instructor classes

4

7

75%

Basic archery instructor refresher classes

1

1

NO CHANGE

Basic archery instructor (teachers) trained

61

126

106%

State tournament participants

287

369

29%

COTREX (Colorado Trail Explorer)
WE’RE MAPPING
COLORADO’S TRAILS
The Colorado the Beautiful vision,
launched by Governor Hickenlooper
in 2015, laid out plans to ensure that
within a generation, every Coloradan
will live within 10 minutes of a park, trail
or vibrant green space. CPW is leading
this effort and aims to map every trail in
the state as part of its mission to provide
enjoyable and sustainable outdoor
recreational opportunities that educate
and inspire current and future generations
to serve as active stewards of Colorado’s
natural resources.
The Colorado Trail System (CTS) through
COTREX, the online program promotes
and enhances the Colorado the Beautiful
mission through a world-class system
of recreational trails in Colorado. CTS
is designed to enhance the outdoor
recreationists’ experience by providing:

• A comprehensive and up-to-date map of

trails in Colorado;

• Increased opportunities for discovering

new trails;

• Eﬃciencies for planning an outing and

finding access to trailheads;

• Better tools for exploring

Colorado’s outstanding network of
recreational trails.

The Colorado Trail Explorer (COTREX)
is powered by the CTS mapping project by
serving as the connection between people,
trails and technology.
Shrub jay

MIKE DELLIVENERI/CPW

VIC SCHENDEL/CPW

It was a year of significant growth for
CPW’s National Archery in the Schools
Program across the state.

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

2019 Fact Sheet
A REVIEW OF STATEWIDE CONSERVATION AND RECREATION PROGRAMS

Pheasant hunting in Eastern Colorado

cpw.state.co.us

VIC SCHENDEL/CPW

Bull moose in State Forest State Park

MIKE DELLIVENERI/CPW

Fishing in the Arkansas River

DUSTIN DOSKOSIL/CPW

Conservation at our Core
All of us at Colorado Parks and Wildlife are
very grateful for the support of our hunters,
anglers, parks visitors and so many other
Coloradans that understand, appreciate and
participate in the conservation work that is
at the core of CPW. It shows in the purchases
of hunting and fishing licenses and parks
passes, as well as in the over 6,000 volunteers
that contribute time and energy to our parks
and projects. It shows in the number of calls
placed to Operation Game Thief to prevent
resource theft, and the wide-ranging group
of residents that attend public meetings, and
submit comment forms on different wildlife
and park management plans statewide. It is
because of your time, support, and contributions that the agency is able to enact our
mission: to perpetuate the wildlife resources
of the state, provide a quality parks system,
and provide enjoyable and sustainable recreation opportunities that educate and inspire
current and future generations to actively
participate in the conservation of Colorado’s
natural resources.

has faced over years of stagnant funding.
The bill holds us accountable to 10 key goals
we plan to work toward addressing by 2025.
These goals build off of the agency’s Strategic
Plan and long term vision for wildlife and
natural resource management. We embrace
the goal of stemming the decline of sportsmen and women in our state, and to improve
and increase big game populations through
investing in habitat and conservation. We
hope to begin planning the next Colorado
State Park for our residents and visitors. We
need to continue recruiting and to retain
highly qualified employees to carry out our
mission. And we strive to engage all outdoor
recreationists in responsibly maintaining and
managing the natural resources that make up
the very heart of our state.

As part of our current generation of stewardship, we want to recognize our hunters,
anglers, parks visitors and those purchasing
registrations and permits for recreational
vehicles. We thank you for your many contributions to conservation in Colorado through
those purchases, as your participation in
these activities is the primary source of
funding in managing the health of our land,
water and wildlife. Colorado’s quality of life,
outdoor heritage, and economic prosperity
are dependent on the health and sustainability of these natural treasures that you help us
conserve and protect every day.

We are grateful every day for the opportunity
to work with, play in and protect the natural
resources of Colorado. It takes a lot of hard
work to maintain and improve habitat, conserve over 960 species, and manage 41 state
parks. But the lands, waters and wildlife that
our residents and visitors cherish are equally
loved by the hundreds of dedicated CPW
professionals that work to ensure the health
and prosperity of these resources every day.

Looking to the future, we are also extremely
thankful for the public support of The Hunting, Fishing and Parks for Future Generations
Act. Passed in May 2018, the bill provides the
agency with several opportunities to continue
working toward our mission, and to address
some of the specific challenges that CPW

All 10 goals can be found on our website, and
we will report our annual progress to the legislature through 2025. While they are large,
these goals are important to us and we know
they are important to all Coloradans.

The men and women of Colorado Parks and
Wildlife are honored to be the face of conservation in Colorado, but it is all of us that
make up its heart. We are thankful for the
opportunity to work with you in conserving
and protecting the beauty of our state.

Jeﬀrey M. Ver Steeg
Acting Director, Colorado Parks and Wildlife

3.19

�Colorado Parks and Wildlife
Funding At-A-Glance
SOURCE OF FUNDS
$231.8

TOTAL FUNDING

$125

LICENSES, PASSES,
FEES &amp; PERMITS

$38

FEDERAL, STATE
GRANTS &amp; LOANS

$34.2

LOTTERY &amp; GREAT
OUTDOORS COLORADO

$13.1

SALES, DONATIONS,
INTEREST &amp; OTHER

$12.1

GENERAL FUND &amp;
SEVERANCE TAX

$9.4

REGISTRATIONS

100%
54%

Colorado Parks and Wildlife (CPW) celebrated former Gov.
John Hickenlooper’s decision to sign the Hunting, Fishing
and Parks for Future Generations Act into law in May 2018.
The law, passing through the legislature with bipartisan
support, provided CPW its first significant cash infusion in
13 years.

16%
15%

Lawmakers agreed with CPW’s position that the increases
were necessary to continue to conserve wildlife resources,
maintain valuable infrastructure, ensure a quality state
park system and operate and maintain outdoor recreation
programs. The bill included 10 goals that the agency must
report on annually to demonstrate how these new funds are
being used.

6%
5%
4%

USE OF FUNDS
$231.8

TOTAL BUDGET

$93.6

PERSONAL
SERVICES

$61.8

OPERATING

$26.5

CAPITAL EQUIPMENT/
IMPROVEMENTS

$14.6

GRANTS TO OTHERS

The bill grants the Colorado Parks and Wildlife
Commission authority to enact modest increases to state
park entrance fees and both resident and non-resident
hunting and fishing licenses based on the Consumer Price
Index. The new revenue will allow CPW to begin to address
projected funding shortfalls of $30 million annually for
wildlife and $11 million annually for parks by 2025.

100%
40%
27%
11%

SCORP

6%

$15.9

PAYMENTS TO
OTHER AGENCIES

7%

$18.1

RESERVE FUND
INCREASE

8%

$1.3

LAND &amp; WATER
ACQUISITIONS

The 2019 Statewide Comprehensive Outdoor Recreation Plan
(SCORP) was finalized and released in January 2019. The plan
identifies four key priorities for all residents to address Colorado’s
needs for advancing conservation and sustainable recreation over
the next five years.

1%

Colorado’s SCORP focuses on these priorities:
1. Enhance sustainable access and opportunity to enjoy the
outdoors
2. Promote stewardship of natural, cultural and recreational
resources
3. Conserve lands, waters and wildlife
4. Ensure adequate funding to sustain Colorado’s outdoors for the
future

WAYNE D. LEWIS/CPW

ALL $ FIGURES SHOWN IN MILLIONS

Rocky Mountain Bighorn Sheep rams

Hunting, Fishing and
State Parks for Future
Generations Act

The SCORP is not a CPW plan, but Colorado’s plan. Our
government, industry and NGO partners all played a role in
developing the plan through a process of outreach meetings and
public comment. It will be critical for CPW and all partners to
share and model the key priorities with their networks, members,
staff and all of Colorado’s residents and visitors.

�Economic Impact of Outdoor Recreation
According to the 2017 Economic Contributions of Outdoor
Recreation in Colorado report by Southwick Associates, 92% of
Coloradans recreate outdoors. So it may be no surprise that the
outdoor recreation industry delivers an economic contribution
of $62 billion dollars to the state each year, and accounts for over
500,000 jobs in Colorado.
The most popular activities are walking and hiking followed by
picnicking, camping and fishing.
The staggering popularity of outdoor recreation, and its importance to our economy, means we must all play a role in ensuring
we enjoy our outdoors with conservation in mind.

Colorado the Beautiful Trail Grants
The Colorado the Beautiful Grant Program
is a partnership between CPW and Great
Outdoors Colorado (GOCO). Over $4 million
in grants were awarded to 11 land managers
to create or improve trails in Colorado. Grants
are awarded based on priorities including

connections to existing outdoor recreation
opportunities, regional trail connectivity,
proximity and benefit to local communities,
wildlife and natural resource mitigation,
collaborative partnerships and approach, and
community health benefits.

Hiking in Ridgway State Park

GOCO and
Lottery FY17–18
Since 1992, Great Outdoors Colorado
(GOCO) has awarded more than $580 million
from its share of Colorado Lottery proceeds.

$16,608,934

GOCO TO PARKS PURPOSE

$16,608,934

GOCO TO WILDLIFE PURPOSE

$14,073,680

LOTTERY TO STATE PARKS
(10% DIRECT DISTRIBUTION)

Volunteer FY 17–18
Numbers
Volunteers play an important role for CPW.
They help us get the job done and are a vital
part of the CPW family.

338,294

VOLUNTEER HOURS

6,134

TOTAL VOLUNTEERS

$8,352,478
DUSTIN DOSKOCIL/CPW

VOLUNTEER VALUE

162

FTE EQUIVALENT

�CPW Fishing App
CPW Fishing, the agency’s oﬃcial mobile
fishing app, is also available free via
both Google Play and the App Store.
For anglers seeking information on new
fishing locations or needing to check for
updated regulations while on the water,
CPW Fishing is an easy way to keep
angling details close at hand.
The app allows discovery of over 2,000
fishing locations in Colorado, and
provides information on the type of
fishing at each location, stream gauges,
species availability, accessibility, familyfriendliness and more

Live Life Outside
This year, CPW set out to create a tagline for
the agency that represented the mission of
the agency, while also serving as a rallying
cry for Coloradans who appreciate the
Colorado outdoors. We reached out to the
entire agency to collect staff insights, ideas
and thoughts about what is at the core of our
agency’s mission.

COTREX App

LIVE LIFE
OUTSIDE

The outcome was “Live Life Outside,” a tagline crafted by our field staff. In three words,
it highlights the passion and lifestyle of our
agency and the vision for all Coloradans —
while at the same time creating an opportunity to talk about why recreation and
conservation should be forever connected.

Hiking in Ridgway State Park

VIC SCHENDEL/CPW

In 2018, Colorado Parks and Wildlife
launched a new and improved version
of the COTREX trail mapping web
application. As part of the updates to the
overall application, CPW also introduced
free iOS and Android apps for COTREX
as well. Now, from either a computer or
phone, hikers, bikers, anglers and other
trail users can access routes and details of
almost 40,000 miles of trails in the state.
Trails are logged across all jurisdictions
and can be filtered by activity, distance,
diﬃculty, accessibility and more.

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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Plague in Gunnison’s Prairie Dogs
MAY 2014

Controlling Plague in Gunnison’s Prairie Dogs

Plague, caused by a non-native bacteria and carried by fleas can kill all prairie dogs in a colony, leading to local extinctions that threaten
the overall survival of the Gunnison’s prairie dog (Cynomys gunnisoni) in Colorado. Unmanaged plague has the potential to reduce
populations of the Gunnison’s prairie dog to the point that a listing as federally endangered or threatened becomes more likely. Such a
listing would not only underscore the need for additional conservation, it could also limit land uses for agriculture and recreation.
The Gunnison’s prairie dog serves key roles in southwest Colorado’s biological landscapes:
• A food source for many wildlife species including raptors and mammalian carnivores.
• Prairie dog colonies serve as habitat for mountain plover, burrowing owl, badger, weasel, snakes and other wildlife species.
Research Objectives
Since the Gunnison’s prairie dog is an important species in the state’s ecosystem, Colorado Parks and Wildlife (CPW) has determined that
controlling plague in this species is a priority conservation action. The CPW goal is to develop plague management tools to help manage
and stabilize Gunnison’s prairie dog populations on public lands and in cooperation with willing landowners.
CPW Plague Control Research Success
The U.S. Geological Survey’s National Wildlife Health Center developed a sylvatic plague vaccine (SPV) and tested the vaccine in the lab
in 2010. That laboratory research demonstrated that the vaccine is safe and protects prairie dogs from plague. Field trials are necessary to
confirm these findings in the wild.
In 2012, CPW conducted the first small-acreage field trials of the vaccine to measure prairie dog consumption of the peanut butter-flavored
baits, compare the colony population before and after the vaccine, and monitor the prairie dogs responses to the vaccine. Those field tests
demonstrated that the prairie dogs readily consumed the baits and the vaccine is safe with no observable negative effects.
In July 2013, CPW staff distributed the oral plague vaccine at three Gunnison’s prairie dog colonies in Gunnison and Teller Counties.
This began a three-year study to determine if use of the SPV increases or stabilizes survival of vaccinated prairie dogs compared to
non-vaccinated prairie dogs. These ongoing research efforts found:
• Prairie dogs captured at the treated colonies were tested and, in some cases, 90 percent had consumed the vaccine bait.
• The safety of the vaccine and the high uptake rates were also demonstrated when CPW researchers tested the vaccine on a black-tailed
prairie dog colony in Larimer County.
• Annual use of plague vaccine may be necessary to protect the young animals produced each year.
• The oral vaccine research project will continue for two more years (2015) to evaluate impacts on treatment and control colonies.

Vaccine baits contain a red dye which
marks the prairie dogs that eat them.

CPW researcher distributes plague vaccine
baits during field trials.

© CPW-Sean Streitch

© CPW-Sean Streitch

© CPW-Sean Streitch

CPW Research Outcome
Encouraged, in part, by CPW’s plague research and management projects, the U.S. Fish and Wildlife Service declined to list the Gunnison’s
prairie dog as endangered or threatened in November 2013.

Gunnison’s prairie dog eating plague
vaccine bait.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman, Suite 618, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us

�The range of the
Gunnison’s prairie
dog includes the
intermountain
valleys and
plateaus of
Southwestern
Colorado.

Historic CPW Plague Control Research and Management
Plague was found in flea samples collected from prairie dog colonies in northwest Colorado in 2007. CPW responded quickly in 2008,
manually dusting white-tailed prairie dog burrows with an insecticide that kills fleas, but is safe for wildlife, in the black-footed ferret
recovery area. The treatments were effective at controlling fleas and slowing the spread of plague, and CPW managers expanded the project
to protect Gunnison’s prairie dogs while continuing to research and develop additional plague management tools.
Starting in 2010, CPW managers dusted Gunnison’s prairie dog colonies in several Colorado counties with insecticide and found that:
• There was no observed mortality due to plague in the Gunnison’s prairie dog colonies that were dusted.
• Some nearby colonies that weren’t dusted experienced severe population declines due to plague.
• Researchers are seeking other more efficient methods to control plague on prairie dog colonies like vaccines.
Ongoing Plague Control Research
CPW researchers will continue to develop and refine new tools, like vaccines, to manage the
spread of plague while using insecticides to stabilize populations of Gunnison’s prairie dogs.
Other current projects include:
• Testing of new types of baits to efficiently distribute vaccine.
• Development of a comprehensive plague control program that utilizes both vaccines
and insecticide to effectively stabilize Gunnison’s prairie dog populations.
Partners: The U.S. Geological Survey’s National Wildlife Health Center, the U.S. Bureau
of Land Management, the City of Fort Collins, the National Park Service, the U.S. Forest
Service and many cooperating private land owners.
Funding: CPW’s Gunnison’s prairie dog research and management projects are funded
mainly through the Species Conservation Trust Fund, which is derived from severance tax
revenues on mineral extraction within Colorado.
General Plague and Human Health Background
The first confirmed human case of plague in Colorado occurred in 1940. Today, plague in
humans is rare with an average of seven cases annually in the U.S. Antibiotics are effective
for treatment of plague if administered in the early stages of the disease.

Gunnison’s prairie dog burrows are
“dusted” to kill fleas.

COLORADO PARKS &amp; WILDLIFE • 1313 Sherman, Suite 618, Denver, CO 80203 • (303) 866-3437 • cpw.state.co.us

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                <text>Plague, caused by a non-native bacteria and carried by fleas can kill all prairie dogs in a colony, leading to local extinctions that threaten the overall survival of the Gunnison’s prairie dog (Cynomys gunnisoni) in Colorado. Unmanaged plague has the potential to reduce populations of the Gunnison’s prairie dog to the point that a listing as federally endangered or threatened becomes more likely. Such a listing would not only underscore the need for additional conservation, it could also limit land uses for agriculture and recreation.</text>
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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Fact Sheet
MEDIA QUICK GUIDE

Colorado’s Declining Mule Deer Population
and CPW’s Proposed Predator
Management Strategy

Purpose and Background
The Commission has approved three other predator control plans since revising its
predator management policy in 2007. Approval was granted in 2011 to remove individual mountain lions preying on translocated desert bighorn sheep in the Middle Delores
River, due west of Montrose near the Utah border. The Commission also approved in
2011 a two-year proposal to remove mammalian predators in the Miramonte Basin,
about half way between Telluride and Utah to increase the recruitment of juvenile
Gunnison sage-grouse. Finally, approval was granted in 2013 to remove mountain lions
preying on Rocky Mountain bighorn sheep translocated to the area of the Hayman
burn near Cheesman Reservoir, southwest of Castle Rock.

© CPW FILE PHOTO

December 14th Commission Meeting in Ft. Collins
The purpose of this Commission meeting is to act on CPW’s staff request to approve
two predator management research proposals, per the Commission’s 2007 policy on
mammalian predator management.

Mule deer fawn

Mule Deer in Colorado
Mule deer are an important wildlife species to the state of Colorado and a large number of communities across the state for a variety of
reasons including hunting, wildlife viewing and associated economic benefits.
Although mule deer populations are doing well in some areas of the state, their numbers are well below the population objective established
for the Piceance Basin, (adjacent to the Roan Plateau – an area
with the largest mule deer herds in the state) and also below
objective in the upper Arkansas River valley.
CPW’s population objective for mule deer is approximately
560,000 statewide; the latest estimate puts the statewide population at under 450,000. CPW is not suggesting that the removal of
predators is the solution to increasing statewide deer numbers to
560,000. These two research projects are designed to evaluate the
extent to which predation is limiting deer population size in the
two specific study areas.

© CPW FILE PHOTO

In 2013, in response to a continued decline in mule deer numbers, CPW developed the West Slope Mule Deer Strategy. The
CPW Commission approved the strategy in December 2014.
CPW sought public input to develop the strategy, holding several public meetings across the state facilitated by an independent contractor to ensure objectivity and fairness.

Mule deer doe with three young fawns
COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�The West Slope Mule Deer Strategy contains seven components that address a variety of impacts to mule deer:
• Landscape‐scale habitat management to improve habitat
• Predator management where predation may be
limiting deer survival
• Protect habitat and mitigate development impacts
• Reduce the impacts of highways on mule deer
survival, movement and migration
• Reduce the impacts of human recreation on mule deer
• Regulate doe harvest and provide youth opportunity

© DAVID HANNIGAN

• Maintain a strong ungulate population and disease
monitoring program and conduct applied research to
improve management of deer populations
CPW will implement each component of the West Slope Mule
Deer Strategy when and where appropriate as funding allows.
CPW’s Research-based Predator Management Strategy
Colorado Parks and Wildlife is a science‐based wildlife management agency, employing some of the top researchers and
wildlife managers in the world. Research projects are compre-

Mountain lion
hensive, detailed and subject to scientific peer review.

The science on the impacts of predators on other wildlife populations is not clear cut or conclusive, contrary to recent claims – CPW would
like to make a significant contribution to that body of scientific wildlife knowledge.
Science
A paper published in 2012 in Mammal Review by Forrester and Wittmer reviewed 48 studies on mule deer survival and predation from the
past 30 years and quantified age-specific vital rates, population growth rates and causes of mortality. The authors wrote that:
• “The relative contributions of predation, forage availability and weather to observed [deer] population changes remains unclear and
controversial”
• “Results of predator control studies remain variable”
• “Predation was the largest proximate cause of mortality in both adults and fawns in all studies, including in fawns during winter”
• “the true effect of summer fawn predation in mule
deer dynamics is currently hard to identify”
• “Increasing wolf populations in Vancouver Island
also caused a decline in high-density black-tailed deer
population, and the predator removal study that was
conducted was successful in increasing deer populations”

• “the role of predation in multi-prey, multi-predator
systems at varying deer population densities needs
further investigation”
Black bear
COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

© CPW FILE PHOTO

• “suppression of both fawn and adult survival
simultaneously from predation and other mortality
sources can lead to marked and sustained population
declines”

�Other Theories
Ongoing CPW scientific research in the Piceance Basin has revealed that lack of quality deer habitat is not limiting this mule deer population, contrary to recent claims.
The Piceance Basin in northwest Colorado represents winter range supporting the largest migratory mule deer population in the state. This
area has been the focus of research and monitoring efforts since the late 1940’s and represents one the best documented mule deer populations in North America. Due to historic mule deer population declines, the Commission reduced total mule deer licenses in the area by 85
percent since 2007 and female licenses specifically have been reduced by about 99 percent; current license allocation in GMU 22 consists of
only 590 antlered deer and 20 antlerless deer licenses. Hunting is not the cause of the mule deer population decline.
Rationale
Based on CPW scientific data, researchers theorize that predation of young fawns in the spring in the Piceance Basin is the limiting factor
and CPW proposes to determine if reducing predator numbers in that specific area can result in increased numbers of deer.
With regard to the upper Arkansas, managers and researchers theorize that predation of adult does and fawns over the winter months have
limited this mule deer population in the past.
Since 1999, the leading cause of known mortality on mule deer in the Arkansas Valley has been mountain lions. From 2008 to 2012, higher
than normal rates of mountain lion predation were consistent with lower adult survival rates. The exact cause of these elevated predation
rates is unknown, but they could be associated with lion harvest quotas not being achieved during this time period. Like the Piceance
proposed study area, insufficient habitat is not considered to be a significant factor in fawn survival rates because biologists have only
documented minimal numbers of mule deer in these management units dying of starvation or suffering malnutrition.
Predation on deer by lions and bears is natural, but today’s ecological systems do not function completely naturally, therefore management
intervention is sometimes required to compensate for natural and manmade landscape changes – CPW is not proposing to completely
eradicate predators.
CPW is regularly called upon to help resolve human‐bear conflicts in Colorado and these conflicts occur to a large degree because of an
ever‐increasing number of people which further alters the natural system.

© CPW FILE PHOTO

Impact
Overall numbers of bears and mountain lions in the population units that include the Piceance study area should be minimally impacted
(anticipate removing about one percent of the lion population and two percent of the bear population). The predator treatment area represents 6 percent of mountain lion population management unit and 16 percent of black bear population unit. Furthermore, the proposed
reductions are consistent with the current mountain lion management objective in this rural area - to maintain relatively low predator
densities to reduce livestock conflicts. This area is currently one of four mountain lion population management units (out of 19 lion units
statewide) managed to minimize livestock conflicts.

Two does
COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�The total number of mountain lions removed during the
proposed nine-year upper Arkansas River research project
is not expected to differ significantly from the number of
lions that hunters would remove from the study area during the same nine-year period if the research proposal was
not implemented. If anything, the overall number of lions
removed during the study is likely to be lower than what
would have occurred by hunting alone.

Public Outreach
CPW initially proposed the Arkansas River predator management project to the Parks and Wildlife Commission at
its meeting in Craig in September 2015. Based on public
comment, CPW postponed seeking Commission approval
(initially scheduled for the November 2015 meeting in Wray)
Black bear
to revise the proposal based on public input and to include
a second research project in the Piceance Basin. At its June
2016 meeting in Pueblo, the Commission approved a reduction in mountain lion hunting quotas necessary to implement the Arkansas River research project, pending final Commission approval of the research plan at the December 2016 Commission meeting. At its
August 2016 meeting in Montrose, the Commission received an update on the timeline for considering the two proposed research
projects. CPW also held three public meetings specifically about the proposed research projects in addition to the three public Commission meetings. A meeting was held in Salida on August 15, in
Rifle on August 16 and in Denver on September 19.
In 2005, the Colorado Division of Wildlife funded a statewide
survey of public opinions and perceptions of mountain lions by
Corona Research, Inc. Some of the relevant findings were:
88.1 percent supported research to learn more about mountain
lion populations
Fifty-seven percent of respondents generally believed mountain
lions to be common in Colorado
A majority (61 percent) of respondents believed mountain lion
populations were stable or increasing in size as opposed to declining (22 percent).

© WAYNE D. LEWIS

© CPW FILE PHOTO

Lion and bear populations remain robust in the state ‐ the
overall number of lions and bears removed as part of this
study is very small when compared to both (Piceance and
upper Arkansas) local populations and when compared
to populations across the state (statewide lion population
estimate is 4,500 ‐ 5,500 and bear population estimate is
17,000 to 20,000).

Mule deer
COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

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                <text>The Commission has approved three other predator control plans since revising its predator management policy in 2007. Approval was granted in 2011 to remove individual mountain lions preying on translocated desert bighorn sheep in the Middle Delores River, due west of Montrose near the Utah border. The Commission also approved in 2011 a two-year proposal to remove mammalian predators in the Miramonte Basin, about half way between Telluride and Utah to increase the recruitment of juvenile Gunnison sage-grouse. Finally, approval was granted in 2013 to remove mountain lions preying on Rocky Mountain bighorn sheep translocated to the area of the Hayman burn near Cheesman Reservoir, southwest of Castle Rock.</text>
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                    <text>C O L O R A D O

P A R K S

&amp;

W I L D L I F E

Moose Reintroduction
NOVEMBER 2013

Moose in Colorado
Moose Background
Moose, (Alces alces), are the largest members of the deer family and are found
in the northern regions of North America, Europe and Asia. Moose prefer
habitat near lakes, rivers or wetlands, where they feed on young plants including shrubs, aquatic plants, grasses, mosses, willow, and aspen.
In 2013, moose inhabited northern forests in 15 states. Twelve states – including Colorado - offered moose hunting. The moose in Colorado belong to the
Shiras subspecies, which are smaller than the subspecies found in Alaska and
Canada. Colorado’s adult moose can run 35 miles per hour, weigh up to 1,000
pounds and stand six feet tall at the shoulder.
History of Moose in Colorado
Historical records dating back to the 1850s indicate that moose wandered into
northern Colorado from Wyoming, but were transient and never established
a stable breeding population. Most of the historic sightings involved hunters
seeing and/or harvesting a single bull moose.
Moose Reintroduction
In the 1950s, Colorado wildlife officials considered bringing moose to the state. There were concerns, voiced at public meetings on a series
of proposals, that moose would compete with cattle and sheep on grazing land, and potentially damage local ecosystems and other wildlife
species. State wildlife managers advocated moose reintroduction because of the abundant habitat available and lack of natural predators.
During the 1960s and the early 1970s, the Colorado Division of Wildlife (now Colorado Parks and Wildlife) worked with the U.S. Forest
Service, the Bureau of Land Management, the National Park Service, the general public and local ranchers on selecting the North Park area
in the Routt National Forest near Walden as the site for the first moose reintroduction.
In 1978, state wildlife experts transplanted 24 male and female moose from Wyoming and Utah to create a breeding population in North
Park and provide hunting opportunities. Additional moose from Wyoming, Utah and Colorado’s own growing population were introduced
to other areas of western Colorado over the years. The project succeeded in creating new hunting opportunities and a popular wildlife
viewing option. The state legislature, in 1995, declared Walden the “Moose Viewing Capital of Colorado.”
By 2012, the reintroduction program had established a breeding population of about 2,300 moose in Colorado. Moose hunting is available
in 39 game management units (GMUs). There were 16,500 applicants for 219 moose hunting licenses and 185 moose were harvested in
2012. While the moose population in other states has declined, Colorado’s moose population continues to grow.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Historical Highlights of Moose Reintroduction in Colorado
1978: North Park in Jackson County, near Walden
• 12 moose from Utah and 12 moose from Wyoming
• Dispersal: Boulder, Clear Creek, Gilpin, Grand, Larimer, Park, Routt and Summit Counties
1987: Laramie River Valley in northwestern Larimer County, near Glendevey
• 12 moose from Wyoming
• Dispersal: Boulder, Clear Creek, Gilpin and Park Counties
1991-1993: Upper Rio Grande River Valley in Mineral and Hinsdale Counties, near Creede
• 93 moose from Wyoming, Utah and North Park, Colorado
• Dispersal: Archuleta, Conejos, Gunnison, La Plata, Rio Grande, Saguache and San Juan Counties
2005-2007: Grand Mesa National Forest in Delta and Mesa Counties near Grand Junction
• 91 moose from Utah
• Dispersal: Eagle, Garfield, Gunnison, Mesa and Pitkin Counties
2009-2010: White River National Forest in Rio Blanco County near Meeker
• 19 moose from Utah and 20 from North Park
• Dispersal: Garfield, Moffat and Routt Counties
Reintroduction Partners: U.S. Forest Service, Bureau of Land Management, National Park Service, Utah Department of Natural Resources,
Wyoming Game and Fish, hunter and angler fees, Safari Club International and many other non-profit organizations. Plains Exploration &amp;
Production Company, Gunnison Energy Corporation., SG Interests, Ltd. have also contributed.

Recent Moose Conflicts
In recent years, moose-human conflicts have arisen as moose move into urban areas. Colorado Parks and Wildlife provides these tips for
avoiding conflicts:
• Keep a safe distance
• Move slowly
• Keep dogs on leashes
• Back off when moose put ears back, roll their eyes or appear aggressive
• Carry a wildlife pepper spray, UDAP or Counter Attack, to use if a moose becomes aggressive

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

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