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                  <text>ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Woodling (1985),
NDIS (2014), and Paul Foutz (pers.
comm., CPW). The remaining maps
were created using county occurrence.
For birds, the following resources were
used: Andrews and Righter (1992),
First Colorado Breeding Bird Atlas
(Kingery 1998), Second Colorado
Breeding Bird Atlas (COBBAII
2015), and CFO (2015). Reptile and
amphibian maps were created using
Hammerson (1999) and NDIS (2014).
Mammal maps were created using
NDIS (2014), Boyle (2006), Grunau et
al. (1999), and Schorr (200 1).
Habitat Condition Scorecards

Wildlife species profiles have been created for priority wetland-dependent birds, mammals,
fish, amphibians, and reptiles. Example species covered include (clockwise from top left):
least tern, Preble's meadow jumping mouse, redbelly dace, and northern leopard frog.

The purpose ofthese profiles is
to provide landowners andland
managers with easy-to-use guidelines
for understanding habitatneeds of
priority wetland-dependent wildlife.

2014). Additional wetland-dependent
priority species more specific to the
mountains or western Colorado are
not included in this series, but may be
developed in ilie future.

Introduction

Colorado Distribution

This series of species profiles consists
of 18 profiles for 26 wetlanddependent wildlife species; three
profiles cover guilds with multiple
related species: the dabbling duck
profile includes seven species; the frog
profile and the redbelly dace profile
each include two species. These 26
species were selected because they
were included in research on habitat
preferences completed as part of
projects in the Lower Soutl1 Platte
River Basin (Ortega 2013) and the
Lower Arkansas River Basin (Ortega

County occurrence for species in the
series is provided on the following
page. A checkmark does not suggest
regular occurrence in a county; rather,
it may indicate that the species has
been observed at least once or may
hypothetically occur. If a species is
ch ecked, please refer to tl1e profile for
more detailed information.
Maps

Colorado distribution maps were
created using multiple credible sources.
Maps were created for fish using
Hydrologic Unit Codes, according to

Habitat Scorecards are included with
each species profile for the purpose
of assessing habitat conditions before
and after implementation of wetland
habitat improvement projects. Please
check the instructions for each species.
In most cases, habitat variables should
be assessed during late spring or early
summer. Each habitat variable has assigned values for ranges of conditions;
the variables and their assigned values
are weighted according to importance
of each variable as perceived by experts
in the field.
Getting Help

Most variables can easily be measured
or determined by anyone. In some
cases, users may wish to solicit
assistance from CPW staff, such as
District Wildlife Managers, Wetland
Focus Area Committees, or Private
Land Biologists at Natural Resources
Conservation Service and U.S. Fish
and Wildlife Service. Questions
can also be addressed to the CPW
Wetlands Program Coordinator,
Brian Sullivan at (970) 472-4306 or
brian.sullivan@state.co.us.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

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*The black rail is not on t he official list of priority species, but is a species of interest in the Lower Arkansas River Basin.
Citations
Andrews, R., and R. Righ ter. 1992. Colorado Birds.
Denver Museum of Natural History, Denver,
Colorado.

Hammerson, G. A. 1999. Amphibians and Reptiles in
Co/orndo: A Colorado Field Guide. University Press
of Colorado and Colorado Division of Wildlife.

lloyle, S. 2006. North American river otter (LonIra
ca11nde11sis): a technical conservation assessment.
USDA Forest Service, Rocky Mountain Region.

Kingery. H. E.. Editor. 1998. Colorado breeding
bird atlas. Colorado Bird Atlas Partnership and
Colorado Division of Wildlife, Denver, Colorado.

COBBAII (Colorado Breeding Bird Atlas II). 2015.
Maps available on line at http://bird.atlasing.org/
Atlas/CO/.

NDIS (Natural Diversity Information Source). 2014.
On-line mapping tool formerly available through
Natural Resource Ecology Laborato ry, Colorado
State University, Fort Collins, Colorado.

CFO (Colorado Field Ornithologists). 20 15. Colorado
county birding. http://coloradocoun tybirding.org/
CountySelector.aspx.
Grunau, L., R. Schorr, D. Green , B. Roscnlu nd, C.
Pague, and J. Armstrong. 1999. Conservation and
management plan for the Preble's meadow jumping
mouse on the U. S. Air Force Academy. Colorado
Natural Heritage Program unpublished report
to the Natural Resources Branch. U.S. Air Force
Academy.

O rtega, C. P. 2013. Habitat quality for wetlanddependent priority wildlife species in the Lower
South Platte River Basin, Colorado: species
assessments and mon itoring protocol. Report
prepared for Colorado Natural Program. Colorado
Parks and Wildlife, and U.S. Environmental
Protection Agency, Region 8.

Ortega. C. P. 2014. Habitat quality for wetlanddependent priority wildlife species in the
Lower Arkansas River Basin, Colorado: species
assessments and monitoring protocol. Report
prepared for Colorado Natural Program, Colorado
Parks and Wildlife, and U.S. Environmental
Protection Agency, Region 8.
Schorr, R. A. 200 I. Meadow jumping mice (Znp11s
l,udsoni11s pre/,lei) on th e U.S. Air Force Academy,
El Paso County. Colorado. Colorado Natural
Heritage Program unpublished report to the
Natural Resources Branch. U.S. Air Force Academy.
Woodling. J. I985. Colorado's little fish: a guide to the
minnows and other lesser known fishes in the
state of Colorado. Colorado Division of Wildlife,
Denver, Colorado.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�ASSESSING HABITAT QUALITY FOR PRIORITY W ILDLIFE SPECIES IN COLORADO W ETLANDS

Species
Distribution
Range
American bitterns breed from the
mid United States through noriliern
Canada. In Colorado, American
bitterns occur in non-contiguous areas
throughout ilie state and are most
common in the San Luis Valley.

The American bittern (Botaurus /entiginosus, Family Ardeidae) is a secretive, marsh bird found
throughout Colorado at lower elevations.

Species Description
Identification
The American bittern is a type of
heron with a haunting low-frequency
dunk-a-doo vocalization that sounds
similar to a metal stake being driven
into mud. Even with their large size
(24-33 inches), they can be difficult to
find because they are secretive, fairly
solitary, and cryptic among emergent
vegetation, especially dry vegetation.
Preferred Habitats
American bitterns occur primarily in
emergent marshes. However, depending on size and condition of the
habitat, they might be found in beaver
ponds, oxbows, herbaceous riparian
wetlands, warm water sloughs, wet
meadows and reclaimed gravel pits.

Diet
American bitterns consume a wide
variety of animal matter, such as amphibians, snakes, small mammals, fish,
crayfish, crabs, insects, and spiders.
They frequently feed at the interface of
water and emergent vegetation.
Conservation St atus
American bitterns are listed as a Nongan1e Species of Management Concern
by the U. S. Fish and Wildlife Service.
States list them as endangered, in1periled, at risk, species of special concern,
or species of greatest conservation
need. In Colorado, American bitterns
are listed as a Tier 2 Species of Greatest
Conservation Need (CPW 201 5). The
International Union for Conservation of Nature lists the population as
decreasin g, but because of its extensive
range and low rate of decline, it is con sidered ofleast concern.

•

Breed ing

•

Year..-ound

•

Wintering

North America map used by permission from Cornell
Lab of Ornithology's Birds of North America Online
(http:/bna.birds.corncll.edu/bna). Colorado map based
on Andrews and Righter (1992), Kingery ( 1998}, COBBAIi (201 5), and CFO (201 5).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Preferred Habitat Conditions
Distance to disturbances

buffer &gt;220 yards

Dominant vegetation

tall, dense and robust emergent vegetation

Height of herbaceous vegetation

3-6.5 feet

Interspersion

complex patterns that maximize interface between
water and vegetation

Landscape context

proximity to other wetlands on the landscape

Percent emergent cover

60-80% with high amount of edge

Residual cover (litter)

m ix of dead and live vegetation, with moderate
litter depth

Size of habitat

large wetlands &gt;25 acres

Water depth (predominant)

at least some open water 3-25 inches deep

Water quality

unpolluted water that will support preferred foods

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for American bitterns.

Acknowledgements
Pete Walker (Colorado Parks and Wildlife) and
Colin Lee (Rocky Mountain Bird Observatory/
Colorado Parks and Wildlife/Natural Resources
Conservation Service) reviewed an earlier
version and provided input on preferred habitat
conditions.
Suggested Reading and Citations

Hydrology

•
•
•

Preserve shallow wetlands interspersed with water.
Avoid complete drawdowns to maintain food base.
Throughout breeding season, maintain water levels between 3-25 inches.
Avoid drawdowns before mid-August.

Vegetation

•
•
•
•
•

Preserve wetlands with robust vegetation.
Occasionally set back succession witl1 disking and/or drying.
Maintain wide and dense vegetated buffer around emergent wetlands.
Limit mowing, burning, and deferred grazing to once every 2-5 years, and avoid
immediately around wetland, especially during breeding.
Use no-tillage or minimal tillage practices; avoid conventional tillage.

Contamination

•

Reduce application of agricultural chemicals and other toxins.
Reduce siltation and eutrophication.

Land Use

•

Restrict livestock grazing within 220 yards of wetland edge.

Conservation

•
•

Preserve wetlands greater than 25 acres, preferably larger.
Maintain mosaic of wetlands in different successional stages.

Andrews, R., and R. Righter. 1992. Colorado
Birds. Denver Museum ofNa1ural History,
Denver, Colorado.
Brown, M .. and J. J. Dinsmore. 1986. Marsh size
and isolation for marsh bird management.
Jo11rnal of Wildlife Management 50: 392-397.
COBBAII (Colorado Breeding Bird Arias 11).
2015. Maps available online at hllp://bird.
atlasing.org/ Atlas/CO/.
CFO (Colorado Field Ornithologists). 2015.
Colorado county birding. Maps available
at hllp://coloradocounlybirding.org/
Count ySclector.aspx.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlitc Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Dechant, J. A., M. L. Sondrcal, D. H. Johnson,
L. D. lgl, C. M. Goldade. A. L. Zimmerman,
and 8. R. Euliss. 2003. Effects of managemenl
practices on grassland birds: American
Bittern. Northern Prairie Wildlife Research
Cen1er, Jamestown, North Dakota.
Gibbs,). P., and S. M. Melvin. 1992. American
Bittern. In Migratory nongame birds of
management concern in the northeastern
United States. K. Schneider and D. Pence,
Eds. U.S. Fish and Wildlife Service, Newton
Corner, Massachusetts.
Kingery, H. E., Editor. 1998. Colorado Br.:eding
Bird Atlas. Colorado Bird Atlas Partnership
and Colorado Division of Wildlife, Denver,
Colorado.
Lowther, P., A. F. Poole, J. P. Gibbs, S. Melvin, and
F. A. Reid. 2009. American Bittern (Botaurus
/entiginos11s). The Birds of North America No.
018. Cornell Lab of Ornithology.
Rehm, E. M., and G. A. Baldassarre. 2007. The
influence of interspersion on marsh bird
abundance in New York. Wilson /011rnal of
Ornitliology 119: 648-654.
Wiggins, D.A. 2006. American Billern (Bota11rus
lentiginosus): a technical conservation
assessment. USDA Forest Service, Rocky
Mountain Region.

COLORADO PARKS &amp; W ILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for American Bitterns (v. Jan 2016)
Assessment of habitat before and after restoration or management actions
Project Name:_ __ _ _ _ __________ _______ ____ Date(s) of Assessment: ____ __ _ _ _
Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.
Key habitat variable and conditions

Value

Before

Size of habitat

&gt;25 acres

10.8

&gt; 12 - 25 acres

7.2

2.5 - 12 acres

3.6

Residual cover (litter) depth

&gt; 15 - 25 inches

10.3

&gt;8 - 15 inches OR &gt;25 - 40 inches

6.8

4-8 inches

3.4

Percent of water with emergent vegetation

&gt;60-80%

10.3

&gt;30 - 60% OR &gt;80 - 100%

6.8

15 - 30%

3.4

Water quality

No visual evidence of turbidity or other pollutants

10.3

Some turbidity or presence of other pollutants, but limited to sma ll and localized areas within the wetland. Water
may be slight ly clo udy.

6.8

Wat er is cloudy or has unnatural oil sheen, but the bottom is still visible. Note: If the sheen breaks apart when you
run your finger through it, it is a natural bacterial process and not water pollution.

3.4

Dominant vegetation

Robust wetland herbaceous plants (e.g., cattail, bulrush, reedgrass) OR tall sedges &gt;8 inches

10.3

Low sedges, rushes, or grasses &lt;8 inches

6.8

An nual and perennial forbs OR open willows and shrubs

3.4

Height of herbaceous vegetation

&gt; 3 - 6.5 feet

9.7

&gt; 1.5 - 3 feet

6.5

1 - 1.5 feet OR &gt;6.5 feet

3.2

Distance to developed areas (urban or industrial land uses)

&gt; 250 yards

9.7

&gt; 100 - 250 yards

6.5

50 - 100 yards

3.2

Interspersion

B or C or D

9.7

E

3.2

0 g @ @ 0
A

Interspersion patterns refer to the diagram
(sti ppled = water, solid = vegetation)

B

C

E

D

.

Percent of wetland with some water

&gt;70 - 100%

9.7

&gt;50 - 70%

6.5

10- 50%

3.2

Predominant depth of water

9.2

2 - 8 inches
&gt;8 - 40 inches

6.1

&lt;2 inches or &gt;40 - 50 inches

3.1

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
Bald eagles live almost entirely within
North America. Their distribution is
highly scattered throughout North
America and in Colorado. Up-to-date
maps of their distribution in Colorado
are unavailable due to their sensitive
status.

Bald eagles (Haliaeetus leucocephalus, Family Accipitridae) prefer tall, mature trees for
perching and nesting. From these perches, they watch for fish or other small prey.

Species Description
Identification
O ur national emblem , the bald eagle,
is distinguished by its white head and
tail contrasting with a dark brown
body in its full adult form. Immature
bald eagles (first four years) may have
light or brown heads and tails; they can
best be distinguished from immature
golden eagles by the light leading edge
of the wing contrasting with darker
flight feathers.
Preferred Habitats
The most important wetland habitats
for bald eagles are those connected
with large bodies of water, such as rivers and reservoirs. Bald eagles use tall
trees and dead snags along tl1e edge
of waterbodies to perch and watch for
prey.

Diet
Bald eagles are opportunistic foragers. They prefer fish when available,
but also consume small mammals and
birds, especially during winter.
Conservation Status
The U. S. Fish and Wildlife Service
removed bald eagles from the Federal
list of endangered species in 2007
(USFWS 2007). In Colorado, bald
eagles are listed as a Tier 2 Species of
Greatest Conservation Need (CPW
2015). l11e number of bald eagle pairs
breeding in Colorado increased from
14 pairs during Colorado Breeding
Bird Atlas I ( 1987-1994) to 148 pairs
during Colorado Breeding Bird Atlas
II (2007- 2011).

-

Breeding

Possible occurence

North America map used by permission from Cornell
Lab of Ornithology's Birds of North America Online
(http:/bna.birds.corncll.cdu/bna). Colorado map based
on Andrews and Righter (I992), Kingery ( 1998), COB·
BAIi (20 15), and CFO (20 15).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 11 92 • cpw.state.co.us

�Preferred Habitat Conditions
Distance between tall trees and
open water with fish

the closer the better, best within 1 mile

Distance from human disturbance

varies with locatio n; 100 to &gt;500 yards

Dominant vegetation

open canopy and easy-to-access large trees with
open horizontal branches

Tree size

mature, tall, large-d iameter trees for perching and
nesting

Management Recommendations
Acknowledgements
Chase Taylor (Rocky Mountain Bird
Observatory) reviewed an earlier version and
provided input on preferred habitat conditions.

This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wilcllife priority wetland-dependent wilcllife. A number of
practical steps can be taken to improve habitat for bald eagles.

Suggested Reading and Citations

Hydrology
• Maintain water levels to maximize prey availability and foraging.
• Manage for a diversity of fish.

Andrews, R., and R. Righter. 1992. Colorado
Birds. Denver Museum of Natural History,
Denver, Colorado.
Berry, M. E., C. E. Bock, and S. L. Haire. 1998.
Abundance of diurnal raptors on open space
grasslands in an urbanized landscape. Condor
100: 601-608.

Vegetation
• Thin dense forests (particularly in the vicinity of reservoirs), burn by prescription,
and selectively harvest to achieve desired open canopy.
• Avoid clearcutting.
• Manage for age diversity and species diversity.
• Protect existing tall, large-diameter trees, including open-top trees and snags.
• Promote growth of trees in areas where lacking.
• Create super-canopy trees (much taller than the rest) by cutting.
• Revegetate with trees where appropriate.
• Create or protect openings that are ~8.5 acres within 2 miles of shoreline; maintain these openings with sparsely-spaced roosting and/or nesting trees.

Caton, E. L., B. R. McClelland, D. A. Patterson,
and R. E. Yates. 1992. Characteristics of
foraging perches used by breeding bald eagles
in Montana. Wilson Bulletin 104: 136-142.
Chester, D. N., F. Stauffer, T. J. Smith, D. R.
Luukkonen, and J. D. Fraser. 1990. Habitat
use by nonbreeding bald eagles in North
Carolina. Journal of Wildlife Management 54:
223-234.
COBBAll (Colorado Breeding Bird Atlas II).
20 I 5. Maps available online at http://bird.
atlasing.org/Atlas/CO/.
CFO (Colorado Field Ornithologists). 2015.
Colorado county birding. Maps available
at http://coloradocountybirding.org/
CountySelector.aspx.

Contamination
• Reduce contaminants.

CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.

Land Use
• Reduce visual contact witl1 disturbance using vegetation strips 8-100 yards wide.
• Minimize development along shorelines.

Kingery, H. E., Editor. 1998. Colorado Breeding
Bird Atlas. Colorado Bird Atlas Partnership
and Colorado Division of Wildlife, Denver,
Colorado.

Conservation
• Preserve undeveloped shores and adjacent forests.
• Maintain mosaic of wetlands in different successional stages.

Grubb, T. G., and R. M. King. 1991. Assessing
human disturbance of breeding bald eagles
with classification tree models. Journal of
Wildlife Management 55: 500-511.
USFWS. 2007. Bald eagle fact sheet: natural
history, ecology, and history of recovery. U. S.
Fish and Wildlife Service.

~

u:
~

~

u:
~

oi

"'"'
"'0"'

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~0

::.

!2
0

~
~

~

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Habitat Scorecard for Bald Eagles (v.Jan 2016)
Project Name:_ _ _ _ __ _ _ _ _ _ _ _ _ _ _ _ __ __ _ __ _

Date(s) of Assessment: _ _ _ _ _ _ _ __

Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Dominant vegetation

Open canopy with dominant snag or tall trees with open horizontal branches

18.5

Distance to permanent open water capable of supporting fish from a live or dead tree &gt;30 feet tall

&lt;55 yards

17.5

55 ya rds - 1 mile

11.7

&gt;1 - 2 miles

5.8

Percent total canopy cover that is &gt;6.5 feet

10 - 40%

17.5

&gt;40 - 60%

11.7

&gt;60 - 100%

5.8

Height of tallest tree within 100 yards of focal area

&gt;50 feet

17.5

&gt;30-S0feet

11 .7

15 - 30 feet

5.8

Distance from development

&gt;0.3 mile
0.2 - 0.3 miles

10.3

100 yards - 0.2 miles

3.4

6.8

Apparent water depth

&lt;20 feet deep within 100 yards of shore

14.5

&lt;20 feet deep within 50 yards from shore but deeper beyond

9.7

&gt;20 feet deep within SO yards from shore

4.8

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
Black rails are widely, but sparsely,
distributed through North America,
Central America, and the Caribbean.
In Colorado, they are known from
five counties: Bent, Lincoln, Otero,
Prowers, and Pueblo.

Black rails (Lateral/us jamaicensis, Family Rallidae) are both the smallest and most secretive
rails in North America. They hide within dense marsh vegetation.

Species Description
Identification
Secretive amongst dense vegetation
and no larger than a sparrow (about
6 inches long), black rails are difficult
to detect in the field. Their gravely
kee-kee-deer song and their haunting
kookaburra-like call are heard more at
night than during the day.
Preferred Habitats
Black rails rely most frequently on
dense emergent marshes, including
beaver ponds.
Diet
From limited knowledge, the diet of
black rails appears to consist primarily
of invertebrates and some seeds.

■ Breeding

Conservation Status
Black rails appear to have declined over
the last century, due almost entirely to
habitat loss and destruction. They are
listed as endangered in Arizona, threatened in California, and near threatened by the International Union for
Conservation of Nature. Though they
are not listed as a Species of Greatest
Conservation Need in Colorado (CPW
2015), they are locally important in the
Lower Arkansas River Basin.

Year-round
• .. • Local breeding/summer records
••• Year-round records
.. Local winter rocords
•

Isolated breeding/summer records
lsolaled winter records

Possible occurence

Nonh America map used by permission from ComeU
Lab of Ornithology's Birds of North America Online
(lmp:/bna.birds.cornell.edu/bna). Colorado map based
on Andrews and Righter ( 1992), Kingery (1998), COBBAIi (201 5), and CFO (2015).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Preferred Habitat Conditions
Depth of water and water
fluctuations

&lt;1.2 inches water with little fl uctuation

Dominant vegetatio n

cattails, rushes, sedges, and grasses; in Colorado,
occupied habitats are dominated by cattails

Herbaceous cover height

tall vegetation, presumably fo r protection

Interspersio n

interspersion patterns favo ring high vegetation
density

Percent of emergent vegetation

dense or thick

Residua l cover (litter) depth

mixture of new and residual growth

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for black rails.
Hydrology
• Reduce fluctuations of water levels through the nesting season to maintain moist
soil or shallow water.
• Improve water control devices where needed.
• Impoundments should provide sloping shoreline that provides 25% shallow water
averaging 1 inch.
Vegetation
• Maintain emergent vegetation.
Contamination
• Reduce contaminants where needed.
Conservation
• Reduce h abitat destruction.
• Manage wetlands on regional scale.

Acknowledgemen ts
Tony Leuker ing (formerly with Rocky Mountain
Bird Observatory) reviewed an earlier version and
provided input on preferred habitat conditions.
Suggested Reading and Citations
Andrews, R., and R. Righter. 1992. Colorado
Birds. Denver Museum of Natural History,
Denver, Colorado.
COBBAII (Colorado Breeding Bird Alias 11).
2015. Maps available on line at h11p://bird.
atlasing.org/ Atlas/CO/.
CFO (Colorado Field Ornithologists). 201 5.
Colorado county birding. Maps available
at hllp://coloradocountybirding.org/
CountySelector.aspx.
CPW (Colorado Parks and Wildlife). 201 5.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Eddleman, W. R., R. E. Flores, and M. Legare.
1994. Black rail (Latem//11s jamaice,isis). The
Birds of North America No. 123. Cornell Lab
of Ornithology.
Flores, R. E., and W. R, Eddleman. 1993.
Nesting biology of the California black rail
in southwestern Arizona. Western Birds
24:8 1-88.
Flores, R. E., and W. R, Eddleman. 1995.
California black rail use of habitat in
southwestern Ariwna. /011rnnl of Wildlife
Management 59:357-363.
Kingery, H. E., Editor. 1998. Colorado Breeding
Bird Atlas. Colorado Bird Atlas Partnership
and Colorado Division of Wildlife, Denver,
Colorado.
Legare, M. L., and W.R. Eddleman. 200 1. Home
range size, nest-site selection and nesting
success of black rails in Plorida. Journal of
Field Ornithology 72: 170- 177.
Tsao, D. C., J. Y. Takckawa, I. Woo, J. L. Yee,
and J. G. Evens. 2009. I Jome range, habitat
selection, and movements of California black
rails at tidal marshes at San Francisco Bay,
California. Condor 111:599-610.

COLORADO PAR.KS &amp; WILDLI FE • 6060 Broadway, Denver, CO 802 16 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Black Rails (v. Jan 2016)
Project Name: ___ __ _ _____________________ Date(s) of Assessment: _________
Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Value

Key habitat variable and conditions

Before

Dominant vegetation
Robust wet land herbs (cattails, bulrush, reedgrass, etc.), Tall sedges, rushes, and grasses

17.2

Water depth and hydrology
17.2

&lt;1.2 inch water and little to no water fluctuation or moist soil throughout

Percent of emergent vegetation
&gt;90 - 100%

17.2

&gt;80-90%

11.5

70-80%

5.7

Height of emergent vegetation
Open (little to no vegetation)

18.5

Residual cover depth
&gt;20 - 80 inches

16.4

&gt; 12 - 20 inches

10.9

8- 12 inches

5.5

Interspersion
15.6

E
D

10.3

0 g

B

A

Interspersion patterns refer to the above diagram
(stippled = water, solid = vegetation)

.

Total (of 100 possible): add all numbers in before or after columns

C

D

(9 @

E

0

After

�Several species are included in the Dabbling Duck guild. Top row from left: American
wigeon (Anos americana), blue-winged teal (A. discors), cinnamon teal. (A. cyanoptera).
Second row from left: gadwall (A. strepera), green-winged teal (A. crecca), mallard (A.
p/atyrhynchos). Bottom: northern pintail (A. acuta).

Species Description
Preferred Habitats

The most important wetland habitats
for dabbling ducks during spring and
fall migration include beaver ponds,
emergent marshes, warm water
sloughs, moist soil units, wet meadows,
and herbaceous riparian wetlands.
During winter, most small wetlands
freeze and ducks congregate in deeper
water, such as open river channels,
warm water sloughs, reservoirs, and
deep gravel pits, or on open sandbars.
During the breeding season, most dabbling ducks nest in upland vegetation.

Diet

Most dabbling ducks consume
far more invertebrates during the
breeding season compared with other
times of year. During non-breeding
seasons, the diet varies according to
species but includes seeds, aquatic
vegetation, tubers, and crop grains.
Conservation Status

The population status diffe rs among
species. All ducks in this guild are
federally protected game birds in the
United States, Can ada, and Mexico.
Colorado Parks and Wildlife designated these ducks as priorily species
because they provide valuable hunting
and viewing opportunities.

Species Distribution
Range

The ducks in this guild are widely
distributed. With the exception of
cinnamon teal, all have a distribution
beyond the Americas. TI1ey are
found throughout most of Colorado
during at least part of the year. In
general, outside of winter, the greatest
concentrations include the San Luis
Valley, North Park, and the Front
Range within the South Platte River
Basin.

COLORADO PARKS &amp; W1LDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

sedges, rushes, grasses, forbs, and aquatic
vegetation

Density of plants desirable to
ducks

abundant (desireable plants are often seed bearing
species such as pondweeds, dock, sedges, and
some grasses)

Emergent vegetation within open
water

21-50% for diurnal use
61-80% for nocturnal use

Interspersion

complex patterns that maximize interface between
water and vegetation

Landscape context

proximity to other wetlands on the landscape

Size of habitat

&gt;20 acres for wet meadows
&gt;2 acres for other wetlands excluding reservoirs

Submergent vegetation

31-60%

Water depth (predominant)

4-12 inches

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for dabbling ducks.
Hydrology
• Maintain water depths 4-12 inches.
• Time drawdowns in summer to coincide with desired vegetation.
• Drawdown gradually for the greatest diversity of vegetation.
• Re-flood in late summer or early fall for fall migrants.
Vegetation
• Consider establishing submerged aquatic vegetation.
• Consider revegetating with native plants during drawdown if devoid of vegetation
for long periods.
• Use disturbance techniques to set back succession.
• Control undesirable vegetation, especially robust plants, exotics, and woody
vegetation.
• Control woody vegetation at young age.
• Create 50:50 interspersion or hemimarsh conditions (l:l open water to emergent
vegetation).
• Manage for diversity of native plants.
• Use drawdowns to accelerate decomposition.
Land Use/ Other
Limit time of grazing and maintain appropriate stocking rate; where possible,
protect wetland with fencing.

Acknowledgements
Brian Sullivan (Colorado Parks and Wildlife)
reviewed an earlier version and provided input
on preferred habitat conditions.
Suggested Reading and Citations
Austin, J. E., and M. R. Miller. 1995. Northern
pintail (A11as awta). The Birds o f North
America No. 163. Cornell Lab of Ornithology.
Drilling, N., R. Titman, and P. Mckinney. 2002.
Mallard (Anas platyr/1y11cl10s). The Birds
of North America No. 658. Cornell Lab of
Ornithology.
Gammonley. J. H. 1996. Cinnamon teal (Anas
cya11optera). The Birds of North America No.
209. Cornell Lab of Ornithology.
Johnson, K. 1995. Green-winged teal (/\nas
crecca). The Birds of North America No. 193.
Cornell Lab of Ornithology.
Leschack, C. R., S. K. McKnight, and G. R. Hepp.
1997. Gadwall (/\11as strepcra). The Birds
of North America No. 283. Cornell Lab of
Ornithology.
Mowbray, T. I 999. American wigeon (/\11as
americana). The Birds of North America No.
40 I. Cornell Lab of Ornithology.
Naugle, D. E., R. R. Johnson, M. E. Estey. and K.
F. Higgins. 2001. A landscape approach to
conserving wetland bird habitat in the prairie
pothole region of eastern South Dakota.
Wetlands 21:1-17.
Rohwer, F. C., W. P. Johnson, and E. R. Loos.
2002. Blue-winged teal (A 11as discors). The
Birds of North America No. 625. CorneU Lab
of Ornithology.

Conservation
• Minimize disturbance by humans.
• Control for burrowing mammals only if needed
to maintain integrity oflevees or to avoid
excessive vegetation removal or obstructions.
• Control fish that cause turbidity, e.g., carp.
• Provide diversity through wetland complexes on
landscape.

COLORADO PARKS &amp;WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Dabbling Ducks (v. Jan 2016)
Project Name:_ _______________ _ _ __ _ ______ Date(s) of Assessment: _ _ _ __ _ _ __
Instructions: Select appropriate checklist: (1) Emergent Wetlands, Playas, and Impoundments, (2) Wet Meadows, or (3)
Sandbars. Enter one value that best describes migratory (spring/fall) conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.
Emergent Wetlands, Playas, and Impoundments
Key habitat variable and conditions

Value

Before

Dominant vegetation
Sedges, rushes, grasses, forbs, and aquatic vegetation

18.7

Robust wetland herbs (cattail, bulrush, reedgrass, etc.)

12.5

Open willows / shrubs, Closed ca nopy trees (&gt;50% cover)

6.2

Percent of emergent vegetation within water
21 - 50%

18.7

5 - 20%

12.5

50 - 100%

6.2

Predominant depth of water
4-12 inches

18.7

&gt; 12 - 25 inches

12.5

&gt;25 - 40 inches

6.2

Percent submergent vegetation
&gt;30 - 60%

17.8

&gt; 10 - 30%

11 .8

0 - 10%

5.9

Interspersion
C or D

15.0

B

10.0

A or E

5.0
A

Interspersion patterns refer t o the above diagram
(sti pp led = w ater, solid= vegetation)

Size of habitat

B

0 0

&gt;2 acres

C

D

E

(9 ~ 0
11.1

&gt;0.5 - 2 acres

7.5

0.25 - 0.5 acres

3.7

Total (of 100 possible): add all numbers in before or after columns

After

�Habitat Scorecard for Dabbling Ducks (v. Jan 2016)
Project Name:. _________________ _ _ _ _ __ _ _ __ Date(s) of Assessment: _ _ __ _ _ _ __
Instructions: Select appropriate checklist: (1) Emergent Wetlands, Playas, and Impoundments, (2) Wet Meadows, or (3)
Sandbars. Enter one value that best describes migratory (spring/fall) conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.
Wet Meadows (natural or irrigation-influenced)
Key habitat variable and conditions

Value

Before

Dominant vegetation
Sedges, rushes, grasses, forbs, and aquatic vegetation

28.2

Robust wetland herbs (cattail, bulrush, reedgrass, etc.)

18.8

Open willows/ shrubs, Closed canopy trees (&gt;50% cover)

9.4

Percent of herbaceous vegetation that is too dense for a duck to move through
0-20%

28.2

&gt;20-50%

18.8

&gt;50 - 80%

9.4

Height of herbaceous vegetation
8- 20 inches

26.7

&gt;20 - 80 inches

17.8

&gt;80 inches

8.9

Size of habitat
&gt;20 acres

16.9

&gt;5 - 20 acres

11.3

2.5 - 5 acres

5.6

Total (of 100 possible): add all numbers in before or after columns

After

�Habitat Scorecard for Dabbling Ducks (v. Jan 2016)
Project Name:.______ _ _ _ _ _ _ _ _ _ _ _ __ _ _ _ _ _ _ __ Date(s) of Assessment: _ __ _ _ _ _ __
Instructions: Select appropriate checklist: (1) Emergent Wetlands, Playas, and Impoundments, (2) Wet Meadows, or (3)
Sandbars. Enter one value that best describes migratory (spring/fall) conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Sandbars
Key habitat variable and conditions

Value

Before

Dominant vegetation
Sedges, rushes, grasses, forbs, and aquatic vegetation

25.6

Robust wetland herbs (cattail, bulrush, reedgrass, etc.)

17.1

Open willows / shrubs, Closed canopy t rees (&gt;50% cover)

8.5

Percent of herbaceous vegetation that is too dense for a duck to move through
0- 20%

25.6

&gt;20-50%

17.1

&gt;50-80%

8.5

Percent of herbace ous vegetation that is easy for a duck to move through
0 - 30%

25.6

&gt;30 - 60%

17.1

&gt;60 - 100%

8.5

Percent cover of woody vegetation &gt;6.6 feet (2 meters) in height
0-20%

23.2

&gt;20- 40%

15.4
7.7

&gt;40 - 100%
Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALIT Y FOR PRIORIT Y WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range

Least terns range extensively along
coasts and major rivers in North
America, Central America, the Caribbean, and northern South America. In
Colorado, least terns breed only in the
Lower Arkansas River Basin. During
migration, they occasionally occw- in
the Lower South Platte River Basin,
and less frequently on the west slope.
a~\1

The least tern (Sternula albifrons, FamilyLaridae) is a small, ground nesting bird that nests in
open, sandy soil.They are rare throughout their range, including in Colorado.

Species Description
Identification

Diet

At 8-9 inches in length and slightly
smaller than an American robin,
the least tern is the smallest tern of
North America. Their white forehead
contrasting with a black cap is easily
seen in flight. They have a yellow bill
tipped in black.

The diet of least terns consists
primarily of small (1-4 inches long),
slender(½ inch) fish that swim near
the surface. To a lesser degree, they
also feed on aquatic invertebrates.

Preferred Habitat s

Interior least terns nest on sandy
shores of reservoirs and gravel pits;
although not usually considered wetlands, these habitats are often adjacent
to or associated with wetlands along
the shores. Sandbars, particularly along
the Arkansas River, represent potential
nesting habitat, however, the regulation
of water probably precludes least terns
from successfully nesting. They forage
in streams, reservoirs, marshes, gravel
pits, and other impounded wetlands.

Conservation Status

The interior population is Federally
listed as endangered. In Colorado,
least terns are listed as endanged and
a Tier 2 Species of Greatest Conservation Need (CPW 2015). During
the second Colorado Breeding Bird
Atlas (2007-2011), least terns were
observed and confirmed breeding in
only two priority blocks: Adobe Creek
Reservoir and just south of Neesopah
and Neegronda Reservoirs. During
the previous Colorado Breeding Bird
Atlas (1987-1994), they had also been
confirmed as breeders in the vicinity of
Neenoshe Reservoir.

North America map used by permission from Cornell
Lab of Ornithology's Birds of North America Online
(hnp:/bna.birds.comeU.cdu/bna). Colorado map based
on Andrews and Righter (1992), Kingery ( I998), COBBA II (20 I5), and CFO (20 I 5).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

open view with little to no vegetation

Height of vegetation

&lt; 6 inches

Location of nesting habitat patch

next to water with an unobstructed view

Nesting substrate

sand or fine gravel, free from silt and/or clay

Percent canopy cover

none

Percent herbaceous cover

0-5%

Size of nesting habitat if other ideal
conditions met

&gt; 0.2 acres

Acknowledgements
Dr. Eileen Kirsch (U. S. Geological Service)
reviewed an earlier version and provided input on
preferred habitat conditions.

Size of unvegetated patch on
vegetated bar

&gt; 1 acre

Suggested Reading and Citations

Water depth at nest sites

0 inches

Andrews, R., and R. Righter. 1992. Colorado
Birds. Denver Museum of Natural History,
Denver, Colorado.

Woody debris on ground

presence of woody debris near nests (not well
understood)

COBBAII (Colorado Breeding Bird Atlas 11).
20 15. Maps available on line at http://bird.
atlasing.org/ Atlas/CO/.

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A nwnber of
practical steps can be taken to improve habitat for least terns.
Hydrology/ Substrate
• Manage and time flows to benefit habitat (scouring) while minimizing nesting
mortality.
Create sand and gravel river islands.
Control vegetation by flooding after breeding and lower water level prior to arrival
in spring.
Add sand to islands as needed.
Add small amount of woody debris where needed.
Vegetation
• Control vegetation to create or maintain sparse vegetation.
Land Use
• Curtail cattle grazing near potential nesting sites.
Conservation
Provide exclosures/enclosures or electric fencing where needed to protect from
predators.
Close beaches to human use during breeding season.
Create educational signage to protect breeding birds from human disturbance.

CFO (Colorado Field Ornithologists). 201 5.
Colorado county birding. Maps available
at http://coloradocountybirding.org/
CountySelecLor.aspx.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Ki ngery, H. E.. Editor. 1998. Colorado Breeding
Bird Atlas. Colorado Bird Alias Partnership
and Colorado Division of Wildlife, Denver,
Colorado.
Kirsch, E. M. 1996. Habitat selection and
productivity of least terns on the Lower Platte
River, Nebraska. Wildlife Monograpl,s I 32:
3-48.
Kirsch, E. M .. and/. G. Sidle. I 999. Status o f the
interior population of least tern. Journal of
Wildlife Management 63: 470-483.
Koenen. M. T., D. M. Leslie, /r., and M. Grcgorr1996. Habitat changes success of artificial
nests on an alkaline salt flat. Wilson B11/leti11
108: 292-301.
Siulc, J. G., and E. M. Kirsch. J 993. Least tern
and piping plover nesting at sand pits in
Nebraska. Colonial Waterbirds I 6: I39-148.
Sidle, ). G., and W. F. Harrison. I 990. Recovery
plan for the interior population o f the least
tern Sterna a11tillaru111. U.S. Fish and Wildlife
Service.
Sidle, /. G., D. E. Carlson, E. M. Kirsch, and J. J.
Dinan. I992. Flooding: mortality and habitat
renewal for least terns and piping plovers.
1992. Colonial Waterbirds 15: I32-136.
Thompson, B. C., J. A. Jackson, ). Burger, L. A.
Hill, E. M. Kirsch, and J. L. Atwood. 1997.
Least tern (Stern 11/a a11tillarum). llte Birds
of North America No. 290. Cornell Lab of
O rnithology.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-11 92 • cpw.state.co. us

�Habitat Scorecard for Least Terns (v. Jan 201 6)
Project Name:___________________________ Date(s) of Assessment: _ _ _ _ _ __ _ _
Instructions: Select appropriate checklist: (1) Nesting (e.g., sandbars, reservoir edges, gravel pits) or (2) Foraging (e.g., emergent
marshes, ponds, stream channels, warm water sloughs). Enter one value that best describes early to mid-summer conditions of
each habitat variable, using the numbers in the value column. Habitat variables are in shaded boxes; ranges of condition are directly
below each variable. If condition is outside range or is not described, enter a zero.

Nesting Habitat (e.g. sandbars, reservoir edges, gravel pit s)
Key habitat variable and conditions

Value

Before

Dominant vegetation
Open bare ground

11.8

Low grass

3.9

Location: Juxtaposition of habitat patch to water
Totally surrounded by water

11.8

Partially connected and adjacent to water

7.9

Not adj acent to water

3.9

Location: Distance with unobstructed view (from appropriate patch)
&gt;275 yards

11.8

&gt; 100 - 275 yards

7.9

SO - 100 yards

3.9

Percent total canopy cover 6.6 feet
NO canopy cover &gt; 6.6 feet

11.8

Substrate
&gt; 75% coarse and/or fine sand

11.3

Size of habitat patch (Answer for ONLY one, using best option)
Size of entirely unvegetated bar or island
&gt;0.25 acre

10.7

&gt;0.06 - 0.25 acre

7.1

0.02 - 0.06 acre (~900 sq feet- ~2,500 sq feet)

3.6

Size of unvegetated patch on otherwise vegetated bar or island
&gt; 1 acre

10.7

0.5 - 1 acre

7.1

0.25 - 0.5 acre

3.6

Predominant water depth
No water

10.7

Vegetation height
&lt;6 inches

9.4

6-20 inches

3.1

Total (of 100 possible): add all numbers in before or after columns

After

�Habitat Scorecard for Least Terns (v.Jan 2016)
Project Nam e:_ __ _ _ _ _ _ __ __ _ _ _ _ __ _ _ _ _ __ _ _

Date(s) of Assessment: _ __ _ _ _ _ __

Instructions: Select appropriate checklist: (1) Nesting (e.g., sandbars, reservoirs, and gravel pits) or (2) Foraging (e.g., emergent
marshes, ponds, stream channels, warm water sloughs). Enter one value that best describes early to mid-summer conditions of
each habitat variable, using the numbers in the value column. Habitat variables are in shaded boxes; ranges of condition are directly
below each variable. If condition is outside range or is not described, enter a zero.

Foraging Habitat (e.g. emergent marshes, ponds, stream channels, warm water sloughs}
Key habitat variable and conditions

Value

Before

Interspersion
Aor B

27.1

C

18.0

D

9.0
A

Interspersion patterns refer to the above diagram
(stippled = water, solid = vegetation)

Dominant vegetation

B

C

D

E

0 0 ® @ 0

Open water (no vegetation)

24.3

Sparse emergent vegetation

16.2

Presence of small fish
Abundant small fish 1 - 4 inches

24.3

Abundant aquatic invertebrat es

16.2

Distance to potential nesting habitat
&lt;3 miles

24.3

3-6 miles

16.2

&gt;6- 9 miles

8.1

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
The patchy distribution of Lewis's
woodpecker is restricted to western
North America, from southern New
Mexico to mid British Columbia and
from California to eastern Colorado.
In Colorado, they can occur almost
anywhere and have been known to
breed in well over half of Colorado's
counties.

Lewis's woodpeckers (Melanerpes lewis, Family Picidae) nest in open cottonwood or
ponderosa pine stands and hunt insect in flight.

Species Description
Identification
Named after the explorer Meriwether
Lewis, Lewis's woodpeckers differ from
most other woodpeckers in their foraging habits, including catching insects
in flight. They are approximately 11
inches long with hefty bodies. Their
backs and wings are a greenish black
and bellies a pinkish to salmon rose.
Their green color is due to reflection
from feather structure rather than
from green pigment; therefore, the
intensity of green varies with lighting,
and they can appear almost black in
poor lighting.
Preferred Habitats
Cottonwood gallery forest is the most
important wetland type used by Lewis's
woodpeckers. However, they also
use riparian shrub wetlands and wet
meadows, and they forage over stream
channels.

Diet
Lewis's woodpeckers forage
opportunistically on locally abundant
insects, including outbreaks in burned
forests and hatches over water and wet
meadows. During summer months,
they feed primarily on insects; in fall
and winter, they switch to feeding
heavily on mast fruits (especially
acorns) and agricultural grains or
waste.
Conservation Status
Almost all populations of Lewis's
woodpeckers have declined. The most
recent Colorado Breeding Bird Atlas
(COBBA 2007-2011) indicates that
they were observed in 36% fewer
blocks than during the first COBBA
{1987-1994). In Colorado, Lewis's
woodpeckers are listed as a Tier 2
Species of Greatest Conservation Need
{CPW2015).

■ Wintering

-

Breeding

Possible occurence

North America map used by pennission from Cornell
Lab of Ornithology's Birds of North America O nline
(h11p:/bna.birds.corneU.edu/bna). Colorado map based
on Andrews and Righter ( 1992), Kingery ( 1998), COBBA 11 (2015), and CFO (2015).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 802 16 • (303) 297-1 192 • cpw.state.co.us

�Preferred Habitat Conditions
Clusters of large-diameter trees

5-15 tree clusters

Dominant vegetation

open canopy with snags (standing dead trees) and
som e shrub or brushy layer

Height of trees

&gt;50 feet

Percent bare ground

very little (&lt;5%)

Percent cover of canopy trees

prefer open canopy &gt; 15-30%

Percent mowed or fallow fields
within 100 yards

&gt;40%

Percent of mast fruits (e.g., acorns)
or grain (e.g., corn) within 100
yards

&gt;35%

Percent shrub and/or brushy layer

prefer some shrub/brush 10-33%

ttl
~

Size of habitat

&gt;15 acres

0

ffi
:a:
0
f;j

:,

8

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for Lewis's woodpeckers.

Hydrology
• Manage stream flow to encourage cottonwood regeneration.
Vegetation
• Retain at least 50% of snags &gt;9 inches in diameter during post-fire salvage logging.
• Retain large snags.
• Protect dumps of large trees and snags.
Encourage regeneration of cottonwoods/snags in riparian zones.
• Maintain open forests.
• Maintain understory shrub community.
Pesticides
• Reduce pesticide exposure.
Land Use
• Lin1it browsing by livestock.
Silviculture
• Maintain six or more snags per acre.
• Where snags are unavailable, consider topping trees.
• Provide no-logging buffer zone to
protect nesting habitat.

~

Acknowledgements
Dr. Kerry Vierling (Department of Fish and
Wildlife Science, University of Idaho) reviewed
an earlier version and provided input on
preferred habitat conditions.
Suggested Reading and C itations
Abele, S. C., V. A. Gaines, and 0. Garton. 2004.
Lewis"s woodpecker (Mela11erpes lewis): a
technical conservation assessment. USDA
Forest Service, Rocky Mountain Region.
Andrews, R., and R. Righter. 1992. Colorado
Birds. Denver Museum of Natural History,
Denver, Colorado.
Bock, C. E., H. H. Hadow, and P. Somers. I 971.
Relations between Lewis' and red-headed
woodpeckers in southeastern Colorado.
Wilson Bulletin 83: 237-248.
COBBAII (Colorado Breeding Bird Atlas 11).
20 15. Maps available online at http://bird.
atlasing.org/Atlas/CO/.
CFO (Colorado Field Ornithologists). 2015.
Colorado county birding. Maps available
at http://coloradocountybirding.org/
CountySelector.aspx.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Gebauer, M. 2004. Lewis's woodpecker,
Mc/a11crpes lewis. Accounts and Measures for
Managing Identified Wildlife 2004: 1- 12.
Kingery, H. E., Editor. 1998. Colorado Breedi11g
lJird Alias. Colorado Bird Atlas Partnership
and Colorado Division o f Wildlife, Denver,
Colorado.
Saab, V. A.. and J. G. Dudley. I 998. Responses
of cavity-nesting birds to stand-replacement
fire and salvage logging in ponderosa
pine/ Douglas-fir forests of southwestern
Idaho. RMRS-RP-11. U.S. Department of
Agriculture, Forest Service, Rocky Mountain
Research Station, Ogden, Utah.
Saab, V. A.. R. E. Russell, and J. G. Dudley. 2007.
Nest densities of cavity-nesting birds in
relat ion to postfire salvage logging and time
since wildfire. Condor 109: 97-108.
Vierling, K. T., V. A. Saab, and B. W. Tobalske.
20 I 3. Lewis's woodpecker (Mehmerpes lewi.,).
11,e Birds of North America No. 284. Cornell
Lab of O rnithology.
License for Francesco Veronesi photo: https://
creativecommons.org/licenses/by-sa/2.0/
deed.en

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Lewis's Woodpeckers (v. Jan 201 6)
Project Name: _ _ _ _ _ _ _ _ _ __ _______________ Date(s) of Assessment: _ _ _ _ _ _ _ __
Instructions: Enter one value that best describes early to mid-sum mer conditions of each h abitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Dominant vegetation

Open canopy with snags

15.9

Tree height

&gt;50 feet

15.9

&gt;35 - 50 feet

10.6

15 - 35 feet

5.3

Percent cover of dominant canopy trees

&gt;15-30%

14.4

&gt;30 - 50%

9.6

5 - 15% OR &gt;50 - 80%

4.8

Percent shrub or brush layer

10- 33%

12.6

&gt;33 - 50%

8.4

&gt;50%

4.2

Percent bare ground

0-5%

12.6

&gt;5 - 15%
&gt;15 - 35%

4.2

8.4

Number of mature tree clusters

5 - 15 clusters

11.2

3 - 4 clusters

7.4

1 - 2 clusters

3.7

Size of habitat

&gt; 15 acres (&gt;6 ha)

6.2

&gt;2- 15 acres (&gt;2-6 ha)

4.1

1.2 - 5 acres (0.5 - 2 ha)

2.1

Percent mowed or fallow fields within 100 yards

&gt;40- 100%

6.2

20-40%

4.1

&lt;20%

2.1

Percent mast fruits (e.g., acorns) or grain (e.g., corn) within 100 yards

&gt;35- 100%

5.6

20-35%

3.7

&lt;20%

1.9

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
Long-billed curlews breed in the
western United States, including
eastern Colorado, and in southwestern
Canada. During migration, long-billed
curlews occur sporadically in western
Colorado and regularly throughout
eastern Colorado.

Long-billed curlews (Numenius americanus, Family Sco/opacidae) have a distinctive long bill
that curves downward. They are can be found near playas and ponds in eastern Colorado.

Species Description
Identification
The long-billed curlew, at 20-26 inches
in length, is the largest shorebird in
North America. Their primitivesounding curlee vocalizations are
considered a harbinger of spring. Th eir
down-curved, sickle-shaped bill is the
largest among shorebirds and inspired
their genus name, Numenius, derived
from the Greek word, noumenios,
meaning of the new crescent moon.
Preferred Habitats
Long-billed curlews are considered a
grassland species, but they are rarely
observed far from water. In Colorado,
they are usually associated with ponds,
reservoirs, playas, and wet meadows.
Diet
Long-billed curlews probe or peck
for invertebrates, including mollusks,
worms, crustaceans, spiders, and

insects, particularly grasshoppers.
They also eat some vertebrate species,
including fish, amphibians, and bird
eggs/nestlings.
Conservation Status
Populations oflong-billed curlews
have experienced overall declines in
many areas, especially throughout the
eastern United States, due primarily to
habitat loss and historic over-hunting.
In Colorado, long-billed curlews are
listed as a Tier 2 Species of Greatest Conservation Need (CPW 201 5).
The Breeding Bird Survey indicates a
significant population decline in Colorado, and the Colorado Breeding Bird
Atlas indicates a decrease in distribution. Confirmed breeding observations
oflong-billed curlews were reported in
~60% fewer priority blocks during the
second Colorado Breeding Bird Atlas
(2007-2011) compared to the first atlas
(1 987-1994).

-

Breeding

Possible occurance

North America map used by permission from Cornell
Lab ofOrnithology's Birds of North America Oruine
(http:/bna.birds.cornell.cdu/bna). Colorado map based
on Andrews and Righter (1992), Kingery ( 1998), COBBAil (2015), and CFO (2015).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominan t vegetation

short grasses, sedges, and short annual forbs

Height of herbaceous vegetation

8-20 inches

Landscape context

mosaic of short grasslands, meadows, agricultural
fields, and wetlands

Percent cover of emergent
vegetation

0-33%

Size of habitat

&gt;50 acres

Water depth in wet meadows

0 or hummocks

Water depth in wetlands

0-6 inches

Management Recommendations

Acknowledgements
Colin lee (Rocky Mountain Bird O bservatory/
Colorado Parks and Wildlife/ Natural Resources
Conservation Service) reviewed an earlier
version and provided input on preferred habitat
conditions.

This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for long-billed curlews.

Suggested Reading and Citations

Vegetation

Andrews, R., and R. Righter. 1992. Colorado
Rirds. Denver Museum of Natural History,
Denver, Colorado.

•
•
•
•
•
•

Create and maintain diversity of vegetation within wet meadows by mowing,
rotational burning, and/or grazing as appropriate.
Remove residual vegetation.
Control tall d ense vegetation.
Use periodic prescribed burns to control tall vegetation.
Control woody plants and shrubs within appropriate grasslands.
Control height of vegetation through appropriate livestock grazing.
Use livestock grazing to maximize low vegetation but reduce grazing intensity
during breeding period to reduce trampling effects.

Pesticides

•
•

Avoid controlling grasshoppers with pesticides.
Reduce use of herbicides on grasslands.

Conservation

•
•

Protect breeding birds from human
disturbance.
Create 220-325-yard buffers around
suitable habitat.

COBBAII (Colorado Breeding Bird Atlas 11).
20 15. Maps available online al hllp://bird.
atlasing.org/ Atlas/CO/.
CFO (Colorado Field Ornithologists). 20 15.
Colorado county bird ing. Maps available
al http://coloradocou ntybirding.o rg/
Count ySclcctor.aspx.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colo rado. Denver,
Colo rado.
Dark-Smiley, D. N., and D. A. Keinath. 2004.
Species assessment for long-billed curlew
(Nwnenius nmeriw 1111s) in Wyoming. Repo rt
to Bureau of Land Management, Wyoming
State Office, Cheyenne, Wyoming.
Dechant, J. A., M. l. Sondreal, D. H. Johnson, l.
D. lgl, C. M. Goldade, P. A. Rabie, and B. R.
Euliss. 2003. Effects o f management practices
on grassland birds: long-billed curlew.
Northern Prairie Wildlife Research Center,
Jamestown, ND.
Dugger, B. D., and K. M. Dugger. 2002. longbilled curlew (N11111e11it1s nmerica1111s). The
Birds of North America No. 628. Cornell Lab
of Ornithology.
Kingery, H. E., Editor. 1998. Colorado Breedi11g
Rird A/Ins. Colorado Bird Atlas Partnership
and Colorado Division of Wildlife, Denver,
Colorado.
Mueller, J. 2000. Long-billed curlew (Nt1111e11it1s
nmericnnus). Fish and Wildlife Habitat
Management Leaflet, Number 7. Natural
Resources Conservation Service, U. S.
Department of Agriculture.
Pampush, G. J., and R. G. Anthony. 1993. Nest
success, habitat utilization and nest-site
selection of long-billed curlews in the
Columbia Basin, Oregon. Condor 95:957-967.

:,:

"'..:
Cl

:a:
-&lt;

"':.:
i3
&gt;::

"'"'u

o(.:, -

"'"'-&lt;

:.:
0

..
:,:

~

Saalfeld, S. T., W. C. Conway, D. A. Haukos, M.
Rice, S. l. Jones, and S. D. Fellows. 2010.
Multiscale habitat selection by long-billed
curlews (N11111eni11s nmerica1111s) breeding in
the United Stales. Waterbirds 33: 148- 161.
Scdg wick, J. A. 2006. l ong-billed curlew
(Nwnenius nrnericanus): a technical
conservation assessment. USDA Forest
Service, Rocky Mountain Region.
Linccnse for Michael Baird photo: Im ps://
crcativccommons.org/licenses/by/2.0/deed.cn

COLORADO PARKS &amp;WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Long-billed Curlews (v.Jan 2016)
Project Name: _ _ __ _ _ _ __ _ _ _ __ _ __ _ _ _ _ _ _ _ _ Date(s) of Assessment: _ _ _ _ _ _ _ __
Instructions: Select appropriate checklist: (1) Playas or other wetlands or (2) Wet Meadows. Enter one value that best describes
early to mid-summer conditions of each habitat variable, using the numbers in the value column. Habitat variables are in shaded
boxes; ranges of condition are directly below each variable. If condition is outside range or is not described, enter a zero.

Playas or Other Wetlands
Key habitat variable and conditions

Value

Before

Percent of water with emergent vegetation
0-33%

25.2

&gt;33 -50%

16.8

&gt;50- 70%

8.4

Predominant depth of water
0-6 inches

25.2

&gt;6- 7 inches

16.8

&gt;7 - 8 inches

8.4

Size of habitat
&gt; 50 acres

19.9

&gt; 12 -50 acres

13.3

7- 12 acres

6.6

Height of herbaceous vegetation
8-20 inches

19.9

&gt;20 -40 inches

13.3

&gt;40 inches

6.6

Dominant vegetation
Low grasses &lt;8 inches

9.8

Low sedges, rushes &lt;8 inches OR Annua l or perennial forbs

6.5

Tall grasses &gt;8 inches

3.3

Total (of 100 possible): add all numbers in before or after columns

After

�Habitat Scorecard for Long-billed Curlews (v. Jan 201 6)
Project Name: _ _ __ _____________ __________ Date(s) of Assessment: _ _ _ __ _ _ __
Instructions: Select appropriate checklist: (1) Playas or other wetlands or (2) Wet Meadows. Enter one value that best describes
early to mid-summer conditions of each habitat variable, using the numbers in the value column. Habitat variables are in shaded
boxes; ranges of condition are directly below each variable. If condition is outside range or is not described, enter a zero.

Wet Meadows
Key habitat variable and conditions

Value

Before

Height herbaceous cover
8 - 20 inches
&gt;20 - 40 inches

28.7
19.1

&gt;40 inches

9.6

Predominant depth of water
0 o r hum mocks

28.7

Size of habitat
&gt;50 acres

21.4

&gt; 12 - 50 acres

14.3

7 - 12 acres

7.2

Percent of herbaceaous vegetation t hat is too dense or hard for animal movement
0 - 20%

11.2

&gt;20-50%

7.4

&gt; 50-80%

3.7

Dominant vegetation
Low grasses &lt;8 inches

10.0

Low sedges, rushes &lt;8 inches OR Annual or perennial forbs

6.7

Tall grasses &gt;8 inches

3.3

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
Piping plovers have a disjointed,
narrow distribution restricted to North
America with rare records in Ecuador.
They occur in eastern Colorado as very
rare migrants and have been known
to nest at a few reservoirs: Adobe
Creek, John Martin, Neegronda, and
Neeskah.

The piping plover (Charadrius melodus, Family Charadriidae) is a rare shorebird that nests on
sparsely vegetated, often sandy or rocky beaches.

Species Description
Identification
The piping plover is a small shorebird
(about 7 inches long). During the
breeding season, the piping plover has
a single black neck band and short
black-tipped orange bill. The piping
plover makes a distinctive high -pitched
pipe-pipe-pipe-pipe-pipe... call, emitted
during flight. Other common vocalizations include a rattling threat call (bee,
bee, bee... ) and an alarm call (woo-up,
woo-up or kee-ah kee-ah).
Preferred Habitats
In Colorado, piping plovers use
sparsely-vegetated sandy shores of
reservoirs and gravel pits where they
nest directly on the ground in simple
scrapes that may or may not be lined
with small pebbles or shells.

Diet
The diet of piping plovers consists
entirely of invertebrates.
Conservation Status
The interior (non-coastal) population
of piping plovers is Federally listed
as threatened. In Colorado, piping
plovers are listed as a threatened, Tier
2 Species of Greatest Conservation
Need (CPW 2015). They nest on
beaches, and are vulnerable to human
disturbance and increased predation;
therefore, conservation strategies
include boosting reproductive success
with nesting exclosures and closing
nesting beaches to humans during the
breeding season.

■ Breeding
Vllnt~

North America map used by permission from Cornell
Lab of Ornithologys Birds of North America Online
(http:/bna.birds.cornell.edu/bna). Colorado map based
on Andrews and Righter (1992), Kingery (1998), COBBAIi (2015), and CFO (2015).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

none to very sparse clumps of short grasses or
sedges

Landscape context

ungrazed and low human disturbance

Size of habitat

beaches more than 20 yards wide; the larger, the
better

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for piping plovers.
Hydrology/ Substrate
• Engineer and maintain sandbars.
• Create sand and gravel river islands.
• Provide for course material, e.g. pebbles.
• Enlarge or widen habitat where possible.
• Lower summer flows to expose more habitat.
• Manage and tin1e flows to benefit habitat (scouring) while minimizing nesting
mortality.
• Control vegetation by flooding after breeding, and lower water level prior to arrival in spring.
Add sand to islands as needed.
Vegetation
• Clear vegetation from existing sandbars.
• Control vegetation to create or maintain sparse vegetation.
Land Use / Other
Exclude cattle.
Create nutrient-rich foraging habitat.
Conservation
• Close breeding beaches to human use.
• Create educational signage to protect breeding birds from human disturbance.
• Provide exclosures where needed.
• Consider exclosure design and use wiili caution; monitor success.
• Remove predators.
• Create artificial habitat.
• Protect all potential habitat, regardless of size.

Acknowledgements
Dr. James Fraser (Department of Fish and
Wildlife Conservation, Virginia Tech) reviewed
an earlier version and provided input on
preferred habitat conditions.
Suggested Reading and Citations
Andrews, R., and R. Righter. I 992. Colorado
Birds. Denver Museum of Natural I listor y,
Denver, Colorado.
Brown, M. B., }. G. Jorgensen, S. E. Steckler, M.
J. Panella, W, R. Silcock, and C. M. Thody.
201 1. A review of interior least tern and
piping plover management, conservation,
and recovery on the Lower Platte River,
Nebraska. Joint report of the Tern and Plover
Conservation Partnership and the Nongame
Bird Program at the Nebraska Game and
Parks Commission, Lincoln, Nebraska.
Catlin, D. H., J. D. Fraser, J. f-1. Felio, and J. B.
Cohen. 20 11. Piping plover habitat selection
and nest success on natural, managed, and
engineered sandbars. Journal of Wildlife
1Wa11ageme11t 75: 305-3 10.
COBBAIi (Colorado Breeding Bird Atlas II).
2015. Maps available online at http://bird.
atlasing.org/ Atlas/CO/.
CFO (Colorado Field Ornithologists). 20 I 5.
Colorado county birding. Maps available
at http://coloradocount)'birding.org/
CouncySelector.aspx.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver.
Colorado.
Ki ngery, H. E., Editor. I998. Colorado Bl'ceding
Bird Atlas. Colorado Bird Atlas Partnership
and Colorado Division of Wildlife, Denver,
Colorado.
Dcblinger, R. D., J. J. Vaske, and D. W. Rimmer.
1992. An evaluation of different predator
exclosures used to prolect Atlantic Coast
piping plover nests. Wildlife Society B111/etin
20: 274-279.
Elliott-Smith, E., and S. M. Haig. 2004. Piping
plover (Charadrius melodus). The Birds of
North America No. 002. Cornell Lab of
Orni1hology.
Rimmer, D. W., and R. D. Dcblingcr. 1990. Use of
predator exclosures to protect piping plover
nests. Journal of Field Ornithology 6 1:2 17-223.
Sidle, J. G .. D. E. Carlson, E. M. Ki rsch, and J. J.
Dinan. 1992. Flooding: mortality and habitat
renewal for least terns and piping plovers.
Colonial Waterbirds 15: 132- 136.
License for Mdf and Dick Daniels photos: https://
creativccommons.org/licenscs/by·sa/ 3.0/
deed.en

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Piping Plovers (v. Jan 2016)
Project Name:_________ _ _ _ _ _ __ _ _ _ _ _ __ _ _ __ Date(s) of Assessment: _ __ __ _ _ __
Instructions: Enter one value that best describes early to mid-sum mer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Percent open sand or gravel area
&gt;80 - 100%

20.2

&gt;50 -80%

13.5

40-50%

6.7

Size of habitat
&gt;22 yards wide

20.2

&gt; 16 - 22 yards wide

13.5

11 - 16 yards wide

6.7

Percent total canopy cover &gt;6.6 feet
0%

20.2

1-5%

13.5

&gt;5 - 20%

6.7

Percent herbaceous cover without woody vegetation
0 - 5%

20.2

&gt;5 - 10%

13.5

&gt; 10 - 20%

6.7

Dominant vegetation
Low grasses (&lt;8 inches}, annual forbs, OR open (unvegetated}
Perennia l forbs

Total (of 100 possible): add all numbers in before or after columns

19.2
12.8

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range

Sandhill cranes breed in a variety
of northern regions, including
northwestern Colorado. During
migration, sandhill cranes can occur
almost anywhere in Colorado.

Sandhill cranes (Grus canadensis, Family Gruidae) are impressive birds with a wide wingspan,
red eye patch, and loud trumpeting call.

Species Description
Identification
With a length of 3½-4 feet and wingspan of 6-7 feet, sandhill cranes are
hard to miss, but they are sometimes
mistaken for great blue herons. Their
graceful dancing helps establish and
maintain pair bonds, which last a lifetime, and their warbling or trumpeting
calls can be heard from a mile away.
Preferred Habitats

Sandhill cranes occupy numerous
wetland habitats, including emergent
marshes, seeps and springs, wet meadows, moist soil units, playas, reservoirs,
and streams. They rely heavily on grain
crops; therefore, wetlands close to
crops are preferred.
Diet

During migration, sandhill cranes
depend on corn and other grains, such
as wheat, barley, and oats. Other food

items include snails, crayfish, insects,
roots, tubers, small vertebrates, and
waterfowl eggs.
•

Breeding

Conservation Status

-

Year-round

There are several subspecies of sandhill
crane. The greater sandhill crane (G.
c. tabida), listed as a Tier 1 Species of
Greatest Conservation Need in Colorado (CPW 2015), winters primarily
in New Mexico, with spring and fall
stopovers in the San Luis Valley of
Colorado. Grus c. canadensis migrates
through the eastern plains of Colorado.
Although two other subspecies (G. c.
pulla and G. c. nesiotes) are Federally
endangered, sandhill crane populations appear to be stable or increasing
in most areas. In Colorado, breeding
records were confirmed in 40% more
priority blocks during the second
Colorado Breeding Bird Atlas (20072011 ) than the first atlas (1987-1994),
suggesting they use more areas for
breeding than was known historically.

•

Nonbrccdlng

-

Breeding

Possible occurence

Nonh America map used by pcnnission from Cornell
Lab of Ornithology's Birds of North Am erica Online
(http:/bna.birds.corncll.edu/bna). Colorado map based
on Andrews and Righter (1992), Kingery ( 1998), COBBAIi (20 15), and CFO (201 5).

COLORADO PARKS &amp;WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Distance from shore where water is 50-100 yards
4-8 inches deep
Dominant vegetation

open water in roosting wetlands; low grasses,
annual forbs and crop plants in foraging wetlands

Habitat size

&gt;2.5 acres

Height of vegetation

&lt;3.3 feet

Percent emergent cover in roosting 0-20% (open view and little to no emergent
wetlands
vegetation)
Percent herbaceous cover in
foraging wetlands

80-100%

Water depth (predominant)

4-8 inches in roosting wetlands;
0 or hummocks in foraging wetlands

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for sandhill cranes.
Hydrology
• Manage hydrology to maintain adequate width and depth (4-8 inches deep) for
roosting.
• Maintain flowing water to prevent spread of disease.
Vegetation
• Maintain availability of vegetation tl1at produces food.
• Control woody vegetation where needed, especially along shorelines.
• Maintain wetland vegetation at early seral stage.
• Control encroaching coarse emergent vegetation (e.g., cattail, bulrush).
• Use appropriate cattle grazing or fall burning to maintain grass height &lt;10 inches.
Land Use / Other
• Maintain juxtaposition of roosting and feeding sites within 2.5 miles.
Discourage land use changes that reduce availability of small grains.
• Provide grit (e.g., pebbles and small gravel) at roost sites if needed.
• Remove unused fences, towers, and utility lines.
• Avoid development of new roosting habitat adjacent to dangers (e.g., utility lines).
Conservation
• Monitor harvest rate; adjust as needed to maintain desirable population numbers.
• Form and maintain partnerships between agencies and agricultural producers.

Acknowledgements
Rick Schnaderbeck (U. S. Fish and Wildlife
Service) reviewed an earlier version and provided
input on preferred habital condilions.
Sugges1ed Reading and Citations
Armbrusler, M. J. 1987. Habitat suilability index
models: grealer sandhill crane. U. S. Fish and
Wildlife Service Biol. Rep. 82( 10.140). 26 pp.
Andrews, R., and R. Righler. 1992. Colorado
Birds. Denver Museum of Natural History,
Denver, Colorado.
COBBAII (Colorado Breeding Bird Atlas 11).
2015. Maps available online al htlp://bird.
atlasing.o rg/ Alias/CO/.
CFO (Colorado Field Orni1hologis1s).2015.
Colorado coun1y birding. Maps available
at hltp://coloradocoun1ybirding.org/
Coun1ySclcc1or.aspx.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Aclion Plan: A Stralcgy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Iverson, G. V., P. A. Vohs, and T. C. Tacha. 1987.
Habi1a1 use by mid-conlincnl sandhill cranes
during spring migra1ion. /011mnl of Wildlife
Management 5 I: 448-458.
Kingery, H. E., Editor. 1998. Colorado Breedi11g
Bird Atlas. Colorado Bird Alias Parlncrship
and Colorado Division of Wildlife, Denver,
Colorado.
Laubhan, M. K., and J. H. Gamrnonley. 2001.
Agricultural producers' perceptions of
sandh ill cranes in the San Luis Valley of
Colorado. Wildlife Society 811/leti11 29: 639645.
Littlefield, C. D., and G. L. Ivey. 2000.
Conservation assessmenl for grea1er sandhill
cranes wintering on the Consumnes River
floodplain and deha regions of California.
Prepared for lhe Na1urc Conservancy.
Lovvorn, J. R., and C. M. Kirkpalrick 1981.
Roosting behavior and habitat of rnigram
greater sandhill cranes. Journal of Wildlife
Management 45: 842-857.
Tacha, "I'. C., S. A. Nesbitt, and I~ A. Vohs. I 992.
Sandhill crane (Grus ca11adensis). "fhe Birds
of North America No. 03 I. Cornell Lab of
Ornithology.

COLORAD O PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Habitat Scorecard for Sandhill Cranes (v.Jan 2016)
Project Name:___________________________ Date(s) of Assessment: ________ _
Instructions: Select appropriate checklist: (1) Roosting (e.g., emergent marshes, playas, moist soil units, reservoirs) or (2) Foraging
(e.g., emergent marshes, wet meadows, moist soil units, recharge ponds). Enter one value that best describes early to mid-summer
conditions of each habitat variable, using the numbers in the value column. Habitat variables are in shaded boxes; ranges of
condition are directly below each variable. If condition is outside range or is not described, enter a zero.

Roosting Habitat (e.g., emergent marshes, playas, moist soil units, reservoirs)
Key habitat variable and conditions

Value

Before

Predominant depth of water
&gt;4-8 inches

23.1

&gt;8 - 12 inches or 2 -4 inches

15.4

Percent of emergent vegetation
0-20%

20.8

&gt;20-40%

13.9

&gt;40 - 100%

6.9

Distance from shore where water is 4-8 Inches deep
&gt;55 - 165 yards

19.6

&gt;25 - 55 yards

13.1

15- 25 yards

6.5

Dominant vegetation
Open (little to no vegetation)

18.5

Grasses

12.3

Interspersion

A

9.0

SorC

6.0
3.0

D

A

Interspersion patterns refer to the above diagram
(stippled= water, solid= vegetation)
Size of habitat

B

0 0

C

E

D

~ @

0

&gt;2.5 acres

9.0

&lt;2.5 acres

3.0

Total (of 100 possible): add all numbers in before or after columns

After

�Habitat Scorecard for Sandhill Cranes (v.Jan 2016)
Project Name:_ _ _ _ __ _ _ _ _ _ _ _ _ _ _ __ _ __ _ __ _ Date(s) of Assessment: _ _ _ _ _ _ _ __
Instructions: Select appropriate checklist: (1) Roosting (e.g., emergent marshes, playas, moist soil units, reservoirs) or (2) Foraging
(e.g., emergent marshes, wet meadows, moist soil units, recharge ponds). Enter one value that best describes early to mid-summer
conditions of each habitat variable, using the numbers in the value column. Habitat variables are in shaded boxes; ranges of
condition are directly below each variable. If condition is outside range or is not described, enter a zero.

Foraging Habitat (e.g., emergent marshes, wet meadows, moist soil units, recharge ponds)
Key habitat variable and conditions

Value

Before

Dominant vegetation
Low grasses, annual forbs, and crops

20.8

Size of habitat
&gt; 2.5 acres

20.8

&lt; 2.5 acres

6.9

Percent of herbaceous vegetation (no woody plants)
&gt;80- 100%

19.8

&gt;60-80%

13.2

30-60%

6.6

Predominant depth of water
0 or hum mocks

19.8

Height of herbaceous cover
8- 20 inches

18.8

&gt;20 - 40 inches

12.5

&gt;40 inches

6.3

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species Distribution
Range

Short-eared owls exist throughout
much of the world. In Colorado,
both their breeding d istribution
and migratory distribution appear
sporadic, but in general, they are more
common in the grasslands of eastern
Colorado.

Short-eared owls (Asio flammeus, Family Strigidae) use both grasslands and wetlands for
habitat They hunt small mammals within the vegetation of wet meadows and marshes.

Species Description
Identification

At 15 inches long, the short-eared owl
is considered a medium-sized owl.
They hunt mostly at dawn and dusk
and nest on the ground. They have a
wide array of vocalizations, including
a rapid hoo-hoo-hoo-hoo-hoo-hoo song.
Preferred Habitats
Short-eared owls use emergent
marshes, playas, and wet meadows,
where they sometimes hunt. Extensive
grasslands may represent their most
important habitat; therefore, the
wetland habitats closest to large tracts
of grasslands will be of highest value to
short-eared owls.

Diet
The diet of short-eared owls consists
almost entirely of small mammals,
especially voles, mice, and shrews. Less
frequently, they eat birds.

•

Y. .l ◄OUnd

•

Nonbr-'l"G

Conservation Status

Although populations of short-eared
owls have significantly declined in
many areas, including the United
States and Canada, the International
Union of Conservation of Nature
identifies them as a species ofleast
concern because the large global
population and extensive range do
not justify placing them in a more
vulnerable category. In Colorado, they
are listed as a Tier 2 Species of Greatest
Conservation Need (CPW 2015).

-

Brooding

Possible occurence

North America map used by permission from Cornell
Lab of Ornithology's Birds ofNortl1 America Online
(http:/bna.birds.cornell.edu/bna). Colorado map based
on Andrews and Righter ( 1992), Kingery (1998), COBBAil (2015), and CFO (2015).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 8021 6 • (303) 297- 1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

grasses

Height of vegetation

&lt;20 inches

Landscape conte..'&lt;t

mosaics of grasslands, agricultural fields, and
preferred wetlands

Percent herbaceous cover

70-100%

Residual cover

2- 8 years old and 40-60%

Size of habitat

&gt;125 acres

Water depth

0 for nest sites

Management Recommendations
This fact sheet contains easy-to-use guid elines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for short-eared owls.
Vegetation
• Mow, burn, or occasionally graze as appropriate to maintain 2-8 years of accumulation of residual vegetation.
Land Use
Reduce amount of land continually grazed.
Hang strean1ers from fences to avoid collisions.
Remove unused fences.
Conservation
• Protect large open areas, especially native grassland.
• Preserve wetland/grassland mosaic on landscape scale.
• Use Conservation Reserve Program to protect habitat for nesting and foraging.

Acknowledgements
Rick Schnaderbcck (U. S. Fish and Wildlife
Service) reviewed an earlier version and provided
input on preferred habitat conditions.
Suggested Reading and Citations
Andrews, R., and R. Righter. 1992. Colorado
Birds. Denver Museum of Natural History,
Denver, Colorado.
Clark. R. J. 1975. A field study of the short-eared
owl, Asio nammcus (Pontoppidan), in North
America. Wildlife Mo11ograpl,s 47: 1-67.
COBBA II (Colorado Breeding Bird Atlas Ii).
2015. Maps available on line at hllp:1/bird.
atlasing.orgl Atlas/CO/ .
CFO (Colorado Field Ornithologists). 201 5.
Colorado county birding. Maps available
at http://coloradocountybirding.org/
CountySelcctor.aspx.
CPW (Colorado Parks and Wildlife). 201 5.
State Wildlife Actio n Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Dechant, J. A., M. L. Sondreal, D. H. Johnson, L.
D. Igl, C. M. Goldade, M. P. Nenneman, and
8. R. Euliss. 1998 (revised 200 I). Effects of
management practices on grassland birds:
short-cared owl. Northern Prairie Wildlife
Research Center, Jamestown, North Dakota.
IOpages.
Herkert, J. R., S. A. Simpson, R. L. Westemeier,
T. L. Esker, and J. W. Walk. I999. Response
of northern harriers and short-eared owls to
grassland management in Illinois. Journal of
Wildlife Management 63:517-523.
Kingery, H. E.. Editor. I 998. Colorado Breeding
Bird Atlas. Colorado Bird Atlas Partnership
and Colorado Division of Wildlife, Denver,
Colorado.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Short-eared Owl (v. Jan 2016)
Project Name: _______________________ _ ___ Date(s) of Assessment: _ _ __ _ __ __
Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Height of herbaceous vegetation

8- 20 inches

15.9

&gt;20 - 40 inches

10.6
5.3

&gt;40 inches
Estimate of percent of grasslands or grasslands with some agriculture within S miles

&gt;35%

15.9

&lt;35%

5.3

Percent herbaceous vegetation (e.g., grasses; do not include woody vegetation)

&gt;70 -100%

15.9

&gt;50-70%

10.6

30-50%

5.3

Percent residual cover

&gt;40 - 60%

14.3

&gt;20-40%

9.5

10-20%

4.8

Size of habitat

&gt;250 acres

14.3

&gt; 125 - 250 acres

9.5

60 - 125 acres

4.8

Dominant vegetation

Grasses (tall and short)

11.9

Open canopy trees &lt;50% cover

7.9

Predominant depth of water

0 or dry nesting sites with residual cover available

11.8

&lt;1 inch

7.9

1 - 8 inches

4.0

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
Preble's meadow jumping mice occur
only in Colorado and Wyoming. In
Colorado, they are known to occupy
the counties along the Front Range
from ilie Wyoming border through El
Paso County.

Preble's meadow jumping mouse (Zapus hudsonius preblei, Family Dipodidae) is a rare small
mammal that lives in dense riparian vegetation along the Front Range.

Species Description
Identification
Preble's meadow jumping mice grow
to approximately nine inches in length,
including their five and a half-inch
tails. They are mostly nocturnal and
hibernate in upland areas for eight
months of the year, making them difficult to observe.
Preferred Habitats
During summer months, the most
important wetland types occupied
by Preble's meadow jumping mice
include riparian areas and adjacent
wet meadows. During the summer,
they prefer dense shrub, grass and forb
ground cover along creeks, rivers, and
associated waterbodies. From early
fall through the spring, they hibernate
underground in burrows that are
typically at the base of vegetation and
have a northerly aspect.

Diet
The diet of Preble's meadow jun1ping
mice includes, but is probably not
limited to, grass seeds, invertebrates,
fungi, and insect larvae.
Conservation Status
The Preble's meadow jumping mouse
is listed as threatened by ilie U.S.
Fish and Wildlife Service (USFWS
1998). In Colorado, they are listed as a
threatened, Tier 1, Species of Greatest
Conservation Need (CPW 2015). The
Colorado Natural Heritage Progran1
considers ilie subspecies to be globallyimperiled (GS T2) and criticallyimperiled wiili the state (Sl).

-

Known occurence

Posslblo occurenco

Full species range map from USFWS (2004). Colorado
map based on Grunau el al. ( 1999), and Schorr (200 I).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions

;:,,~-.~tr~~

Distance from riparian habitat
patch to water

&lt;10 yards

Dominant vegetation

a combination of grasses, shrubs, forbs, and trees

~·

Habitat size

&gt;60 acres

Herbaceous vegetation height

&gt;20-40 inches

~

Landscape context

ungrazed

~

Percent ground cover

densely vegetated; between grasses, forbs, and
shrubs, close to 100%

Species richness

&gt;20 plant species

Management Recommendations

.,,

..

5
""~
:i:

~
.,
0

This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for Preble's meadow jumping mouse.
Hydrology

•
•

Maintain or restore natural regime.
Control erosion within stream corridor.

0:

0

Acknowledgements
Robert Schorr (Colorado Natural Heritage
Program) reviewed an earlier version and
provided input on preferred habitat conditions.
Suggested Reading and Citations

Vegetation

CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.

•
•
•
•
•

Grunau, L., R. Schorr, D. Green, B. Rosenlund, C.
Pague, and J. Armstrong. 1999. Conservation
an&lt;l management plan for the Preble's
meadow jumping mouse on the U. S. Air
Force Academy. Colorado Natural Heritage
Program unpublished report lo the Natural
Resources Branch, U.S. Air Force Academy.

Provide, restore, and enhance riparian vegetation.
Provide, restore, and enhance upland shrub.
Provide, restore, and enhance grassland habitat.
Control noxious weeds, avoiding May through October.
Consider providing woody debris.

Conservation

•

Use best strategies to minimize disturbance.

Meaney, C. A., A. K. Ruggles, B. C. Lubow, N.
W. Clippinger. 2003. Abundance, survival,
and h ibernation of Preble's meadow jumping
mice (Zapus liudso11ius preblei) in Boulder
County, Colorado. Southwestern Naturalist
48: 610-623.
Schorr, R. A. 2001. Meadow jumping mice
(Zap,u l,udso11ius preblei) on the U.S. Air
Force Academy, El Paso County, Colorado.
Colorado Natural Heritage Program
unpublished report lo the Natural Resources
Branch, U.S. Air Force Academy.

..
:i:

(j

p~

0""
0:

6
z

0

"'
0"'
0:

l)

Q

!&gt;

~

Schorr, R. A. 2003. Meadow jumping mice
(Zapus /iudso11i11s preblei) on the U.S. Air
Force Academy, El Paso County, Colorado:
Populations, Movement an&lt;l Habitat from
2000-2002. Colorado Natural Heritage
Program unpublished report lo the Natural
Resources Branch, U.S. Air Force Academy.
Trainor, A. M, T. M. Shenk, and K. R. Wilson.
2007. Microhabitat characteristics of Preble's
meadow jumping mouse high-use areas.
Journal of Wildlife Management 71: 469-477.
Trainor, A. M., T. M. Shenk, and K. R. Wilson.
2012. Spatial, temporal, and biological factors
associated with Preble's meadow jumping
mouse (Zapus hudso11i11s prel,lei) home range.
Journal o/Mammalogy 93: 429-438.
U.S. Fish and Wildlife Service (USFWS). 20 I 3.
Endangered and threatened wildlife and
plants: 12-month finding on two pelilions lo
dclisl the Preble's meadow jumping mouse.
Federal Register 78: 31680-31712.
USFWS. 2015. Draft recovery plan Preble's
meadow jumping mouse (Zapus /111dso11i11s
preblei). U.S. Fish and Wildlife Service,
Region 6, Lakewood, Colorado.

E
~

Q L---l_..:I_ __.;;...=:;;,,

COLORADO PARKS &amp; WfLDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 11 92 • cpw.state.co.us

�Habitat Scorecard for Preble's Meadow Jumping Mouse (v.Jan 2016)
Project Name:._ _ __ _ __ _ _ _ __ _ _ _ _ _ __ __ __ _ _ __ Date(s) of Assessment: _ _ _ _ _ _ _ __
Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Dominant vegetation
Combination of sedg es, grasses, annual and perennial forbs and dense shrubs

13.2

Combination of sedges, grasses, and dense shrubs

8.8

Combination of sedges, grasses, and more open shrubs

4.4

Percent grass cover
&gt;40%

13.2

&gt;20-40%

8.8

10- 20%

4.4

Percent shrub cover
&gt;80%

13.2

&gt;50-80%

8.8

5 - 50%

4.4

Distance from habitat patch to water
&lt;10 yards

12.5

10- 20 ya rds

8.4

&gt;20- 40 yards

4.2

Species richness (number of species in vegetation community)
&gt;20 species

11.9

10 - 20 species

7.9

&lt;10 species

4.0

Habitat size
&gt;60 acres

10.6

50- 60 acres

7.0

&lt;50 acres

3.5

Percent forb cover
&gt;35 - 50%

10.4

&gt; 15 -35%

6.9

5-15%

3.5

Height of herbaceous vegetation (grasses and forbs)
&gt;20 - 40 inches

9.9

8- 20 inches

6.6

&lt;8 inches

3.3

Land use within 100 yards
Ungrazed

5.1

Grazed lightly

3.4

Grazed moderately

1.7

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range

Northern river otters historically
ranged throughout most of the United
States and Canada but were extirpated
from much of their range in the west.
Reintroductions of otter to Colorado
began in 1976, and they are now
found in small numbers throughout
most of western Colorado with a
more scattered distribution in eastern
Colorado.

Northern river otters (Lontra canadensis, Family Mustelidae) occupy streams and beaver
ponds across Colorado's western slope.

Species Description
Identification

Northern river otters vary in length
from 26-42 inches, with males larger
than females. Males weigh on average
25 pounds; females weigh about 18
pounds. Their long tapered tails make
up about one third their length.

Conservation Status

Northern river otters are listed as
least concern by the International
Union for the Conservation of Nature.
In Colorado, they are listed as a
threatened, Tier 2, Species of Greatest
Conservation Need (CPW 2015).

Preferred Habitats
River otters may inhabit nearly every
aquatic habitat; however, they are most
likely to occupy beaver ponds, stream
channels, and warm water sloughs.
Diet
Otters favor fish but also consume
crayfish, mollusks, frogs, snakes,
turtles, salamanders, birds, mammals,
and fruit.

~

Known occurenco

Possible occurence

North America map used from lntemel Center for
Wildlife Damage Management 01ttp://icwdm.org/
handbook/carnivor/RiverOllers.asp). Colorado map
based on Boyle (2006) and NDJS (2014).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Beaver

often associated with beaver activity or old beaver
structures

Dominant vegetation

healthy riparian vegetation

Habitat size

longer reaches usually better

Shore complexity

undercut banks and other complex features

Structures and debris

habitat complexity, including woody debris and
logjams

Stream order

&gt;4th order

Management Recommendations
Tilis fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife p riority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for northern river otters.
Hydrology
• Maintain natural regime to extent possible.
• Protect instream and seasonal flow regimes.
• Enhance and/or protect stream and habitat connectivity.
Vegetation
• Maintain or recreate conditions favoring healthy riparian vegetation.
• Encourage new recruitment of trees.
• Manage for age diversity.
Land Use / Other
• Enhance and/or protect bank structure.
• Enhance and/or protect flood plain.
• Enhance and/or protect good water quality.
• Protect potential upland denning sites from flooding.
• Manage for prey base (e.g., fish ).
Conservation
• Monitor and control commercial harvest to maintain desirable population
numbers.
• Form and maintain partnerships across jurisdictional boundaries to protect
connectivity and long stream reaches.

Acknowled gements
Eric Odell and Scon Wait (Colorado Parks
and Wildlife) reviewed an earlier version and
provided input on preferred habitat conditions.
Suggested Reading and Citations
Boyle, S. 2006. North American river otter
(Lo11tra ca11nde11sis): a technical conservation
assessment. USDA Forest Service, Rocky
Mountain Region.
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Crowley, S., C. ). Johnson, and D. Hodder. 2012.
Spatial and behavioral scales of habitat
selection and activity by river oilers at latrine
sites. /011mnl of Ma111111nlogy 93: 170-182.
Dubuc, L. J., W. B. Krohn, and R. B. Owen, Jr.
1990. Predicting occurrence of river otters
by habitat on Mount Desert Island, Maine.
Journal of Wildlife Management 54: 594-599.
Gorman, T. A., J. D. Erb, B. R. McMillan, and D.
). Marlin. 2006. Space use and sociality of
river o tters (Lo11tra c,mndensis) in Min nesota.
/011mal of Mn111111alogy 87: 740-747.
Melquist, W. E., and M. G. Hornocker. 1983.
Ecology of river otters in west central Idaho.
Wildlife Mo11ograp/is 83: 3-60.

COLORADO PARKS &amp; W CLDLJFE • 6060 Broadway, Denver, CO 80216 • (303) 297-11 92 • cpw.state.co.us

�Habitat Scorecard for River Otters (v. Jan 201 6)
Project Name:_________ _ _ _ _______________ Date(s) of Assessment: _________
Instructions: Enter on e value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Woody objects, e.g., logs, log jams

&gt;5 large logs

17.8

1 - 4 large logs

11.9

Beaver activity (current or remains)

Current activity or recent remnants

17.8

Historic remnants

11.9

Percent canopy cover &gt;6.6 feet

&gt;50%

17.0

&gt;30-50%

11 .3

20-30%

5.7

Height of canopy cover

&gt;50 feet

17.0

&gt;16- SOfeet

11.3

8 inches - 16 feet

5.7

Number of complex shore features (e.g., undercut banks)

&gt;5
1 -4

17.0
11.3

Percent of water &gt; 8 inches deep

&gt;90%

13.4

&gt;80-90%

8.9

40-80%

4.5

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range

Arkansas darters occur only in parts of
Arkansas, Colorado, Kansas, Missouri,
and Oklahoma. In Colorado, they a.re
known from eight drainages within
the Arkansas River Basin: Upper
Arkansas, Fountain, Chico, Upper
Arkansas-Lake Meredith, Horse,
Upper Arkansas-John Martin, Big
Sandy, and Rush.

Etheostoma cragini

Arkansas Darters (Etheostoma cragini, FamilyPercidae) are small native fish of the Colorado
plains. They inhabit pools and channels within small streams.

Species Description
Identification

Arkansas darters are 2½- to 3-inch fish
belonging to the perch family. During
April and May, the belly of breeding
males changes from white to bright
orange.
Preferred Habitats

Arkansas darters inhabit spring-fed
streams, stream channels, and pools
near shorelines, often with undercut
banks.
Diet

A large portion of the Arkansas
darter's diet consists of snails, but they
conswne many other aquatic animals,
including insects, crustaceans, other
invertebrates, and fish eggs.

Conservation Status

Arkansas darters appear on several
lists of concern: Federal candidate
species (U.S. Fish and Wildlife
Service), sensitive (Bureau of Land
Management), threatened and Tier 1
Species of Greatest Conservation Need
(CPW 2015), and near threatened
(International Union for Conservation
of Nature). Flows on a landscape or
segment scale are needed for the fish
to move between or an1ong pools and
to disperse. Large-scale withdrawals
of ground water and dewatering
streams are thought to have caused
extirpations.
Known occurence

North America map used from US. Geologic Survey
(http://nas.cr.usgs.gov). Colorado map by 1-1IB 8
watershed based on Woodling (1985), NDIS (20 14), and
Paul Foutz (pers. comm., CPW).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

floating, broad-leafed vegetation, especially watercress (Nasturtium officinale)

Landscape context
Stream order

heavy grazing unsuitable

Stream width

varies considerably through range; in Colorado
1.5-22 yards

Substrate

sand, silt, gravel

Sunlight

high exposure to sunlight

first and second but may use larger streams for
dispersal

Water depth (predominant)

varies through range; in Colorado 4-20 inches

Water origin

spring fed

Water quality

clear, pH between 7-8.5

Water temperature

varies across seasons; in Colorado 55-86°F

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildJife. A number of
practical steps can be taken to improve habitat for Arkansas darters.
Hydrology
• Protect springs and connecting streams.
• Maintain refugia on landscape scale.
• Maintain corridors by allowing natural disturbances.
• Protect individual pools.
• Maximize water conservation.
• Protect natural function.
• Promote natural flow regimes.
• Restore and/or enhance connections for migration and dispersal.
Vegetation
• Implement tarnarisk control, with an emphasis on long term management for native vegetation.
Contamination
• Eradicate predatory and non-native fish.
• Avoid pesticide and herbicide use near small streams.
Land Use / Other
• Reduce impact from livestock grazing.
Conservation
• Translocate to establish new populations.
• Identify suitable habitat for potential translocations.

Acknowledgements
Paul Foutz (Colorado Parks and Wildlife)
reviewed an earlier version and provided input
on preferred habitat conditions.
Suggested Readjng and Citations

CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Fuller, P. 2014. Et/reostoma cmgi11i. USGS
Nonindigenous Aquatic Species Database.
Gainesville, Florida.
Labbe, 1'. R.. and K. D. Fausch. 2000. Dynamics
of intermittent stream habitat regulate
persistence of a threatened fish al multiple
scales. Ecological Applications 10: 1774- 179 I.
Layher. B. 2002. Recovery plan for the Arkansas
Darter, Etl,eostoma cragi11i Gilbert. in Kansas.
Prepared for Kansas Department of Wildlife
and Parks.
Miller, D. L. 1984. Distribution, abundance, and
habitat of the Arkansas darter Etl,eostoma
cragi11i (Percidac) in Colorado. Southwestern
Na turalist 29: 496•499.
NDIS (Natural Diversity Information Source).
20 14. On·line mapping tool formerly
available tlirough Natural Resource Ecology
Laboratory, Colorado Stale University, Fort
Collins, Colorado.
Taber, C. A.. 8. A. Taber, and M. S. Topping.
1986. Population structure, growth and
reproduction of the Arkansas darter,
Etl,eostoma cragi11i (Percidae). Southwestern
Naturalist 31: 207·214.
USFWS (United Slates Fish 311d Wildlife Service).
2010. Species Profile for Arkansas darter

(Etl,costoma cragi11i).
Woodling, J. 1985. Colorado's little fish: a guide
to the minnows and other lesser known fishes
in the state of Colorado. Colorado Division of
Wildlife, Denwr, Colorado.

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Habitat Scorecard for Arkansas Darters (v. Ja n 2016)
Project Name:_ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ __

Date(s) of Assessment: _ _ __ _ _ _ _ _

Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column . Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Morphology of stream
Permanent pools w ith connectivity to other waters

10.8

Substrate
Sand must be a component

10.8

Dominant vegetation
Aquatic floating (e.g. w atercress) and aquatic submerged or robust, e.g., cattail, bulrush, reedgrass

10.8

Water quality
No visual evidence of turbidity or other pollutants

10.2

Some turbidity or presence of other pollutants, but limited to small and localized areas within the wetland. Water
may be slight ly cloudy

6.8

Water is cloudy or has unnatural oil sheen, but the bottom is sti ll visible. Note: If t he sheen breaks apart when you
run your finger through it, it is a natural bacterial p rocess and not water pollution

3.4

Water origin
Spring fed

9.7

Stream order
1- 2

9.2

Sunlight
Full sun to 10% shade

9.2

Water depth
&gt;4 - 20 inches

8.5

&gt;20 - 40 inches

5.7

0 - 4 inches or &gt;40 inches

2.8

Stream width at bankfull
3.3 - 5.5 yards

8.5

&gt;5.5 - 22 yards

5.7

&gt;22 - 44 yards

2.8

Water temperature
59 - 63°F

8.1

&gt;63- 77°F

5.4
2.7

&gt;77 - 95°F

Landscape context
Ungrazed

4.2

Grazed moderately

2.8

Grazed heavily

1.4

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range

Brassy minnows range from Canada
south to Colorado and Utah and from
Montana through northeast North
America. In Colorado, they are found
in the Lower South Platte River Basin
and also in the backwaters of the
Colorado River.

Hybognathus hankinsoni

Brassy minnows (Hybognathus hankinsoni, Family Cyprinidae) are small native fish of the
Colorado plains. They inhabit pools and channels within small streams.

Species Description
Identification
Adult brassy minnows are 2 ½-3
inches in length and are variously
described in color, perhaps due to
geographic differences or changes
that occur through preservation
of specimens. The majority of
descriptions include a brassy color on
the sides. The centers of their large eyes
are even with the tips of their snouts.

Diet
The diet of brassy minnows consists
primarily of plankton.
Conservation Status

Brassy minnows occur in low numbers
and are listed as a Tier 1 Species of
Greatest Conservation Need (CPW
2015).

Preferred Habitats
Brassy minnows occupy stream channels (particularly pools), back waters,
and beaver ponds.
Known occurence

North America m ap used from US. Geologic Survey
(http://nas.cr.usgs.gov). Colorado map by H UC 8
watershed based on Woodling (1985), NDIS (2014), and
Paul Foutz (pers. comm., C PW).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

Acknowled gements
Boyd Wright (Colorado Parks and Wildlife)
reviewed an earlier version and provided input on
preferred habitat conditions.

abundant aquatic vegetation, especially
submergent vegetation (plants growning fu lly
underwater)

Landscape context

connec tivity with other waters through the driest
months

Morphology of stream

permanent (deep) pools and backwaters

Predatory fish

absence of large predatory fish

Substrate

organic sediment on top of gravel

Water depth (predominant)

deep enough to persist th rough dry periods

Woody debris

abundant

Suggested Reading and Citations
CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Fuller, P., and M. Neilson. 20 I 5. l-lybog11at/1us
/1a11ki11so11i. USGS Nonindigenous Aquatic
Species Database, Gainesville, Florida.
NDlS (Natural Diversity Information Source).
2014. On-line mapping tool formerly
available through Natural Resource Ecology
Laboratory, Colorado State University, Fort
Collins, Colorado.
Quist, M. C., F. J. Rahel, and W. A. Hubert. 2005.
Hierarchical fauna! fillers: an approach to
assessing effects of habitat and nonnative
species on native fishes. Ecology of Fres/11vater
Fish 14: 24-39.

Management Recommendations

Scheurer, J. A., and K. D. Fausch. 2002. Brassy
minnow in Colorado plains streams:
identification, historical distribution, and
habitat requirements at multiple scales, Final
Progress Report. Colorado Water Resources
Research Institute and Colorado Division of
Wildlife, Aquatic Non-game and Endangered
Wildlife Program, Colorado Water Resources
Research Institute, Colorado State University,
Fort Collins, CO.

This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and W ildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for brassy minnows.
Hydrology
• Manage conditions to create or maintain permanent pools.
• Increase spring flows.

Scheurer.). A., K. D. Fausch, and Kevin Bcstgen.
2003. Mulliscale processes regulate brassy
minnow persistence in a Great Plains river.

Vegetation
• Manage riparian vegetation to create root mass that prevents streambank erosion.
• Manage for conditions that favor submergent vegetation.
• Manage for conditions in riparian area that favor recruitment of native woody
vegetation.

Trn11sactio11s of t/,e A111erica11 Fisheries Society
I32: 840-855.
Schlosser, I. J. 1988. Predation rates and the
behavioral response of adult brassy minnows
(l·Jybognathus hankinsoni) to creek chub and
smaJlmouth bass Predators. Copeia I988:
69 1-698.

Contamination
• Maintain native fish communities.
• Prevent invasion of exotic predators and competitors.
• Monitor streams for toxins and artificial hormones.

Woodling, J. 1985. Colorado's little fish: a guide
to the minnows and other lesser known fishes
in the state of Colorado. Colorado Division of
Wildlife, Denver, Colorado.

Land Use/ Other
• Encourage beaver where appropriate
to create suitable pools.
Conservation
Identify suitable habitat for potential
translocations.
Relocate in suitable habitat.

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COLORADO PARKS &amp; W ILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1 192 • cpw.state.co.us

�Habitat Scorecard for Brassy Minnows (v.Jan 2016)
Project Name:_ _ _ _ __ _ _ __________ _ _ __ __ __ _ Date(s) of Assessment: _ _ _ _ _ _ _ __
Instructions: Enter one value that best describes summer conditions of each habitat variable, using the numbers in the value
column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside range or
is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Stream morphology/landscape

Permanent pools connected with other waters during driest time of summer

21.4

Permanent pools with intermittent connection to other waters during driest ti me of summer
Shallow or temporary pools with connection to other waters during driest time of summer

14.2
7.1

Dominant vegetation

Abundant aquatic vegetation, including submergent veget ation

20.2

Predatory fish

General absence of predatory fish

20.2

Predominant water depth during driest time of summer
Answer for ONLY ONE appropriate stream segment, below, using best option
Upstream reach of streams inhabited by brassy minnow

&gt;15 inches

19.1

&gt;5 - 15 inches

12.8

3 - 5 inches

6.4

Size of unvegetated patch on otherwise vegetated bar or island

&gt;25 inches

19.1

&gt; 15 - 25 inches

12.8

5 -15 inches

6.4

Downstream reach of streams inhabited by brassy minnow

&gt;40 inches

19.1

&gt; 15 - 40 inches

12.8

10- 15 inches

6.4

Woody debris

Ample woody debri s in water to provide cover
Total (of 100 possible): add all numbers in before or after columns

19.1

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range

Northern redbelly dace are found in
many isolated populations across the
northern United States and Canada.
In Colorado, northern redbelly dace
are currently known only in the
West Plum Creek drainage, south of
Chatfield Reservoir. Southern redbelly
dace have a smaller distribution
but are also widely scattered into
isolated populations. In Colorado
southern redbelly dace are known
in five drainages: Upper Arkansas
River, Fountain Creek, Chico Creek,
Apishapa River, and Big Sandy Creek.

Red belly dace (Family Cyprinidae) are small native fish ofthe Colorado plains. Their name

comes from the reb color of their undersides during the summer months.

Species Description
Identification

Two dace are included in this guild:
northern redbelly dace (Phoxinus eos)
and southern redbelly dace (P. erythrogaster). The southern redbelly dace can
attain a length of up to 3 inches, while
the northern redbelly dace is usually
less than 2 inches in length. During
summer, males acquire the red on their
bellies that inspired their names.
Preferred Habitats

Both dace occupy stream channels and
off-channel wetlands.

Diet

Northern redbelly dace are
omnivorous, feeding on vegetation,
small invertebrates, and detritus
throughout the water column.
Southern redbelly dace graze from the
surface of substrate, rocks and other
submerged objects..

Known oceurenc&amp;

Conservation Status

Neither dace speceis is Federally
endangered; however, they are both
considered locally uncommon. Both
are endangered in Colorado, due to
low population numbers, and are listed
as Tier 1 Species of Greatest Conservation Need (CPW 2015).

Known occurence

Colorado occurcnce maps by HUC 8 watershed based
on Woodling (198S), NDIS (2011), and Paul Foutz (pcrs.

comm., CPW).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dom inant vegetation

plenty of vegetation , especially algae

Morphology of stream

slow- moving pools, undercut banks

Pred atory fish

absence ofla rge predatory fish

Predominant water depth

&gt;10 inches

Sunlight

ample shade

Water origin

spring-fed water up to third-order streams

Water quality

clear and unpolluted

Water temperature

cool; 63- 75°F

Woody debris

presence of woody debris as cover

Management Recommendations
Th.is fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A nwnber of
practical steps can be taken to improve habitat for redbelly dace.

Acknowled gements
Boyd Wright and Paul Foutz (Colorado Parks
and Wildlife) reviewed an earlier version and
provided input on preferred habitat conditions.
Suggested Reading and Citations

Hydrology/ Stream Morphology
• Restore streams to natural function, especially dewatered streams.
• Minimize changes to natural stream functions.
• Protect springs that flow into streams.
• Prevent erosion and siltation by maintaining buffers.
• Restore and/or enhance connections for migration and dispersal.
Contamination
• Prevent with barriers invasion by predatory and non-native fish.
• Prevent stocking of predatory fish and bullfrogs.
• Monitor streams for toxins and artificial hormones.
• Consider location of wastewater treatment facilities.
Conservation
• Restock appropriate streams with redbelly dace from closest stream.
• Restrict harvest and enforce regulations.
• Identify suitable habitat for potential translocations.

CPW (Colorado Parks and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conser ving Wildlife in Colorado. Denver.
Colorado.
Woodling, J. 1985. Colorado's little fish: a guide
to the minnows and other lesser known fishes
in the state of Colorado. Colorado Division of
Wildlife, Denver, Colorado.
Northern redbellv dace
Bestgen, K. R. 1989. Distribution and notes on
the biology of Pl1oxinus eos (Cyprinidae)
in Colorado. S011t/rwestem Naturnlist 34:
225-231.
MNHP and MFWP (Montana Natural Heritage
Program and Montana Fish, WildJife and
Parks). 2012. Northern redbelly dace,

Clrrosomos eos.
Nico, L., and P. Fuller. 20 15. Clrrosomus eos. USGS
Nonind igenous Aquatic Species Database,
Gainesville, Florida.
Propst, D. L., and C. A. Carlson. 1986. 111e
d istribution and status of warmwater fishes
in the Platte River d rainage, Colorado.
Southwestern Naturalist 31: 149- 167.
Stasiak, R. 2006. Northern redbelly dace
(Plwxinus eos): a technical conservation
assessment. USDA Forest Ser vice, Rod.-y
Mountain Region.
Southern redbelly dace
Bertrand, K. N., and K. B. Gido. 2007. Effects
of herbivorous minnow, southern redbellv
dace (Phoxi1111s erythrognster), on stream ·
productivity and ecosystem structure.
Oecologia 151: 69- 81.
Seilheimer, T. S., and W. L. Fisher. 20 l 0. Habitat
use by fishes in groundwater-dependent
streams of southern Oklahoma. American
Midland Naturalist 164:2 0 1-2 16.
Slack, W. 1~. M. ·c O'Connell, T. L. Peterson, J. A.
Ewin g Ill, and S. '[ Ross. 1997. lchth)•ofaunal
and habitat associations of d isjunct
populations of southern rcdbclly dace
Pl10xin11s erytrognster (Telcostci: Cyprinidae)
in Mississippi. American Midland Nnt11rnlist
137: 251-265.
Stasiak, R. H. 2007. Southern redbclly dace
(Phoxinus erytlrrogaster): a technical
conservation assessment. USDA Forest
Service, Rocky Mountain Region.

COLORADO PARKS &amp; W ILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297- 1192 • cpw.state.co.us

�Habitat Scorecard for Red belly Dace (v. Jan 201 6)
Project Name: _ _ __ _ _ _ _ _ __ _ _ _ _ _ _ _ _ _ _ _ _ _ __

Date(s) of Assessment: _ _ _ _ __ _ __

Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Stream morphology
Pools, overhung banks, slow water

13.3

Water quality
No visual evidence of tu rbidity or other pollutants

12.5

Some turbidity or presence of other pollutants, but limited to small and localized areas within the wetland

8.4

Water is cloudy or has unnatural oil sheen, but the bottom is still visible. Note: If the sheen breaks apart when you
run your finger through it, it is a natural bacterial process and not water pollution.

4.2

Dominant vegetation
Amp le vegetation, especially algae

12.5

Sunlight availability
Full shade

11.9

Some shade

7.9

No shade

4.0

Water origin/stream order
Answer for EITHER northern or sout hern redbelly dace
Northern redbelly dace
Spring-fed; stream order: 1 - 2

11.9

Southern redbelly dace
Stream order: 1 - 3

11.9

Predatory fish
General absence of predatory fish

11 .9

Predominant depth of water
10 inches

10.4

&gt;8 - 20 inches

7.0

6-8 inches

3.5

Woody debris
Presence of woody debris as cover

10.4

Water temperature
63 - 75°F

5.2

&gt;75°F

1.7

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORIT Y WILDLIFE SPECIES IN COLORADO W ETLANDS

Species
Distribution
Range

Northern leopard frogs range from the
northern United States and Canada to tl1e
more northern parts of the southwestern
United States. With the exception of a few
counties, they occur tl1roughout Colorado.
Plains leopard frogs have a much smaller
distribution than northern leopard frogs,
occurring through tl1e Great Plains
into southeastern Arizona and eastern
Colorado.

Northern Leopard Frog

Two species of leopard frogs occur in Colorado. Northern leopard frogs (Rana pipiens; primary
photo, brighter green) are more widespread than plains leopard frogs (R. b/airi; inset photo).

Species Description

Diet

Two leopard frogs are included in this
guild: northern leopard frog (Rana pipiens)
and plains leopard frog (R. blairi). They
are roughly the sam e size (3- 4 inches as
adults), but plains leopard frogs are usually
browner than the m ostly green northern
leopard frogs.

Adult leopard frogs eat primarily insects
and other invertebrates, iJ1cluding
crustaceans, mollusks, and worms, as
well as small vertebrates, such as other
amphibians and snakes. Leopard frog
tadpoles are herbivorous, eating mostly
free-floating algae, but also consuming
some animal material.

Prefe rred Habitats

Conservation Status

Due to their complicated life history traits,
leopard frogs occupy many habitats during
different seasons and stages of development, but they are closely associated with
wet environments. 1n general, leopard
frogs occupy three categories of habitat:
( 1) over-wintering habitat with deep water
that does not freeze solid; (2) foraging
habitat for adults, which may consist of
uplands, riparian areas, and wet meadows; and (3) breeding habitat suitable for
egg development and tadpole survival. In
general, plains leopard frogs breed in more
ephemeral ponds, while northern leopard
frogs use semi-permanent ponds.

Northern leopard frog populations have
declined throughout their range; they are
listed in all western states and Canada as
sensitive, threatened, or endangered. 1n
Colorado, northern leopard frogs are listed
as a Tier 1 Species of Greatest Conservation Need (CPW 201 5). Less is known
about plain s leopard frog populations, but
threats to plains leopard frogs are probably similar to those of northern leopard
frogs. They are listed as a Tier 2 Species of
Greatest Conservation Need in Colorado
(CPW 2015).

Identification

-

Known occurenco

Pou lble occuranco

Plains l eopard Frog

-

Known occurence

Possible occurence

Leopard frog distribution maps for Colorado adapted
from Hammerson {1999) and NDIS (20 14).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation in adult
foraging wetlands

varies but includes grasses, sedges, and forbs

Dominant vegetation in breeding
wetlands

emergent vegetation with buffer of grasses, sedges,
rushes, and spike rushes

Landscape context

0.6-1.2 miles between categories of habitat (see
above habitat rypes)

Percent vegetation cover

dense and extensive

Predatory fish and bullfrogs

none

Size of habitat

can be very small

Sunlight

high sunlight exposure

Vegetation height in adult foraging
wetlands

6-12 inches

Water depth for winter hibernation deep enough not to freeze to the bottom
Water depth in breeding wetlands

25-40 inches but can vary

Water quality

neutral pH. well-oxygenated, and unpolluted

Water temperature

54-73°F

Acknowledgements
Tina Jackson (Colorado Parks and Wildlife)
reviewed an earlier version and provided input on
preferred habitat conditions.
Suggested Reading and Citations

Management Recommendations

Corn. P.S., and L.J. Livo. 1989. Leopard frog and
wood frog reproduction in Colorado and
Wyoming. Nortl,westt m Nat11ralist 70: 1-9.

This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for leopard frogs.

CPW (Colorado Parks and Wildlife). 2015.
State Wildlife ACLion Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.

Hydrology
Maintain water depths to avoid water column freezing solid.
For breeding ponds, maintain depth of&gt;20 in. until metamorphosis completed.
After mid-July, draw down water containing predatory fish and bullfrog larvae.
Vegetation
Provide grass buffers arOlmd breeding ponds.
Contamination
Reduce nitrogen loading.
Reduce pesticides, chemicals and other toxins.
Reduce predatory fish.
•
Possibly reduce or change mosquito control.
Land Use
Eliminate livestock access to ponds.
Avoid or minimize clear-cutting.
Conservation
Translocate frogs to re-establish populations.
Promote conservation programs to provide grassland component in the landscape.
Promote native species in adjacent lands.

Hammerson. G. A. 1999. Amphibians and
Reptiles in Colorado: A Colorado Field
Guide. University Press of Colorado and
Colorado Division of Wildlife.
Nichols, ). 2006. Petition 10 list the western
United Stales population of northern leopard
frog (Rana pipiens) as threatened. Petitioners:
Center for Native Ecosystems, Biodiversity
Conservation Alliance, Defenders of
Black Hills, Forest Guardians, Center for
Biological Diversity, "The Ark Initiative, Native
Ecosystems Council, Rocky Mountain Clean
Air Action.
NDIS (Natural Diversity Information Source).
2014. On-line mapping tool formerly
available through Natural Resource Ecology
Laboratory, Colorado State University, Fort
Collins, Colorado.
Smith, B. E., and D. A. Keinath. 2004. Species
assessment for the northern leopard frog
(Rana pipie11s) in Wyoming. United States
Depar1111en1 of the Interior. Bureau of
Land Management, Wyoming State Office,
C heyenne. Wyoming.
Smith, B. E., and D. A. Kcinath. 2005. Plains
leopard frog (Ra11a hlairi): a technical
conservation assessment. USDA Forest
Sen~cc, Rocky Mountain Region.
Smith. B.E. and D.A. Kcinath. 2007. Northern
leopard frog (Ram, pipic11s): a technical
conservation assessment. USDA Forest
Service, Rocky Mountain Region.
USFS (Un ited States Forest Service). 2003.
Conservation assessment for plains leopard
frog (Rana hlairi). USDA Forest Service,
Eastern Region.

COLORADO PARKS &amp; W ILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-11 92 • cpw.state.co.us

�Habitat Scorecard for Leopard Frogs (v. Jan 2016)
Project Name:______________ __________ __ _ Date(s) of Assessment: ___ _ _ _ _ __
Instructions: Select appropriate checklist: (1) Breeding Wetlands (e.g., emergent marshes, playas, seeps, springs, moist soil units,
reservoirs, other impoundments), (2) Adult Foraging Wetlands (e.g., wet meadows, riparian areas), or (3) Over-wintering
Wetlands (oxbows, stream channels, warm water sloughs, gravel pits, and reservoirs). Enter one value that best describes each
habitat variable during the appropriate season, using the numbers in the value column. Habitat variables are in shaded boxes; ranges
of condition are directly below each variable. If condition is outside range or is not described, enter a zero.
Breeding Wetlands (e.g., emergent marshes, playas, seeps, springs, moist soil units, reservoirs, other impoundments)
Key habitat variable and conditions

Value

Before

Water pH
6.1 -7

16.3

Water quality
No visual evidence of turbidity or ot her pollutant s
Some turbid it y or presence of other pollutants, but limited to small and localized areas wit hin the wetland; water
may be slightly clo udy
Water is cloudy or has unnatural oil sheen, but the bottom is still visible (note: if the sheen breaks apart when you
run your fi nger through it, it is a natural bacterial process and not water pollution)

16.3
10.8
5.4

Predominant depth of water
&gt;25 - 40 inches

16.3

&gt;40 - 80 inches

10.8

4-25 inches

5.4

Percent total canopy cover 6.6 feet
0 - 30%

15.4

&gt;30- 50%

10.3

&gt;50- 100%

5.1

Percent emergent vegetation
&gt;50- 90%

12.2

&gt;30-50%

8.1

10- 30%

4.1

Height of emergent vegetation
8-40 inches

12.2

&gt;40 - 80 inches

8.1

&gt;80 inches

4.1

Dominant vegetation
Robust wetland herbs (cattail, bulrush, reedgrass, etc.) tall sedges and rushes &gt;8 inches, aquatic vegetation
(submergent, floating leaves, algae)

11.3

Tall grasses (&gt;8 inches), open willows/shrubs

7.6

Total (of 100 possible): add all numbers in before or after columns

After

�Habitat Scorecard for Leopard Frogs (v.Jan 2016)
Project Name:_ _ __ _ __ _ __ __ __ __ _ __ _ __ _ _ _ _

Date(s) of Assessment: _ _ __ _ _ _ __

Instructions: Select appropriate checklist: (1 ) Breeding Wetlands (e.g., emergent m arshes, playas, seeps, springs, moist soil units,
reservoirs, other impoundments), (2) Adult Foraging Wetlands (e.g., wet meadows, riparian areas), or (3) Over-wintering
Wetlands (oxbows, stream channels, warm water sloughs, gravel pits, and reservoirs). Enter one value that best describes each
habitat variable during the appropriate season, using the numbers in the value column. Habitat variables are in shaded boxes; ranges
of condition are directly below each variable. If condition is outside range or is not described, enter a zero.
Adult Foraging Wetlands (e.g., wet meadows, riparian areas)
Key habitat variable and conditions

Value

Before

Height of herbaceous cover

2 - 20 inches

23.0

&gt;20 -40 inches

15.3

&gt;40 - 80 inches

7.7

Predominant depth of w ater

0-4 inches

23.0

&gt;4-8 inches

15.3

&gt;8 - 12 inches

7.7

Water pH

6.1 - 7

23.0

Percent of herbaceous cover

&gt;30 -90%

17.2

25 - 30% or &gt;90%

5.7

Dominant vegetation

Low grasses &lt;8 inches, annual forbs, perenn ial forbs
Total (of 100 possible): add all numbers in before or after columns

11.3

After

�Habitat Scorecard for Leopard Frogs (v.Jan 2016)
Project Name: _______ _ _______________ _ _ __ Date(s) of Assessment: _ _ _ _ _ _ _ __
Instructions: Select appropriate checklist: (1) Breeding Wetlands (e.g., emergent marshes, playas, seeps, springs, moist soil units,
reservoirs, other impoundments), (2) Adult Foraging Wetlands (e.g., wet meadows, riparian areas), or (3) Over-wintering
Wetlands (oxbows, stream channels, warm water sloughs, gravel pits, and reservoirs). Enter one value that best describes each
habitat variable during the appropriate season, using the numbers in the value column. Habitat variables are in shaded boxes; ranges
of condition are directly below each variable. If condition is outside range or is not described, enter a zero.
Over-wintering Wetlands (oxbows, stream channels, warm water sloughs, gravel pits, and reservoirs)
Key habitat variable and conditions

Value

Before

Percent of habitat covered by water
&gt;90-100%

19.4

&gt;60 - 90%

12.9

40 - 60%

6.5

Predominant depth of water
&gt;40 inches

19.4

35 - 40 inches

6.5

Water pH
6.1 - 7

23.0

Water quality
No visual evidence of turbidity or other pollutants.

18.4

Some turbidity or presence of other pollutants, but limited to small and localized areas within the wetland; water
may be slightly cloudy

12.3

Water is cloudy or has unnatural oil sheen, but the bottom is still visible (note: if the sheen b reaks apart when you
run your finger through it, it is a natural bacterial process and not water pollution)

6.1

Percent of submergent vegetation
&gt;30 - 60%

14.6

&gt; 10-30%

9.7

0-10%

4.9

Dominant vegetation
Robust wetland herbs (cattail, bulrush, reedgrass, etc.) tall sedges and rushes &gt;8 inches, aquatic vegetation
(submergent, floating leaves, algae)

Total (of 100 possible): add all numbers in before or after columns

9.7

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species
Distribution
Range
The red-sided garter snake is found
in Canada and the western United
States, with a disjunct distribution in
the western United States. In Colorado,
they are found in the northeast and
north-central part of the state.

Red-sided garter snakes (Thamnophis sirtalis parietalis, Family Colubridae) occur in wetlands
on Colorado's northeastern plains.

Species Description
Identification
Red-sided garter snakes, sometimes referred to as common garter snakes, can
grow as long as 49 inches. In Colorado,
they are usually considerably smaller
and may only grow as long as about 3
feet (36 inches). They have patches of
red between their lighter stripes.
Preferred Habitats
Garter snakes hibernate during the
winter, up to seven or eight months
in the more northern parts of their
range. While they are active, they are
wetland-dependent, occupying most
wetland habitat types within Colorado.
Diet
Red-sided garter snakes consume
primarily amphibians, fish, and
earthworm s.

Conservation Status
The abundance of red-sided garter
snakes is mostly unknown. In
Colorado, they are listed as a Species
of Concern and a Tier 2 Species of
Greatest Conservation Need (CPW
2015). Population declines of other
garter snakes, such as tl1e western
terrestrial garter snake (Tharnnophis
elegans) that consume mostly
amphibians, have apparently been
tied to amphibian declines. Because
the red-sided garter snake eats
primarily amphibians, it is possible
populations of red-sided garter snakes
are associated with populations of
amphibians.

-

Known occurence

Possible occurence

North America map used from Wild Animals of
Manitoba (http://,vw1v.gov.mb.ca/ conservation/wildlife/
mbsp/fs/rsgarter.html). Colorado map based on
Hammcrson (1999) and NDIS {2014).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.state.co.us

�Preferred Habitat Conditions
Dominant vegetation

emergent, sedges, grasses, and other vegetation
that provides cover

Landscape context

close and unfragmented connection between
upland hibernacula (hibernation sites) and
wetlands (foraging habitat)

Percent emergent cover

very dense

Size of habitat

not well understood, but larger is better

Water quality

pH = 6.1-7 with no visual evidence of turbidity or
other pollutants

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for red-sided garter snakes.
Hydrology
• For breeding ponds, maintain depth of &gt;20 inches until metamorphosis
completed.
• After mid-July, draw down water containing predatory fish and bullfrog larvae.
Vegetation
• Provide grass buffers around breeding ponds.
• Manage for dense emergent vegetation.
Containimation
Reduce nitrogen loading.
• Reduce pesticides, chemicals and other toxins.
• Reduce predatory fish.
• Possibly reduce or change mosquito control.
Land Use/ Other
• Eliminate livestock access to ponds.
• Avoid or minimize clear-cutting.

Acknowledgements
Tina Jackson (Colorado Parks and Wildlife)
reviewed an earlier version and provided inp ut
on preferred habitat conditions.
Suggested Reading and Citations
CPW (Colorado Park.~ and Wildlife). 2015.
State Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Hammerson, G. A. 1999. Amplribia11s n11d Reptiles
i11 Colorado. University Press of Colorado.
Niwot, Colorado.
Larsen, K. W.. P. T. Gregory, and R. Antoniak.
1993. Reproductive ecology of the common
garter snake '1/,amnoplris sirtalis at the
northern limic of its range. America11 Midland
Naturalist 129: 336-345.
Manitoba Wildlife Branch.20 15. Wild an imals of
Manitoba: Red-sided garccr snake fact sheet.
https://www.gov.mb.ca/conservation/wildlife/
mbsp/fs/rsgartcr. html.
Mallhews, K. R., It A. Knapp, and K. L. Pope.
2002. Garter snake d istributions in highelevation aquatic ecosystems: is there a link
with declining amphibian populations and
nonnative trout intro duc tio ns? Jourunl of

Conservation
Translocate frogs to re-establish populations that can serve as a food source.
Promote conservation programs to provide grassland component in the
landscape.
Promote native species in adjacent lands.

Herpetology 36: 16-22.
NDIS (Natural Diversity Information Source).
2014. O n-line mapping tool formerly
available through Natural Resource Ecology
Laboratory, Colorado State University, Fort
Collins, Colorado.
Shine, R., M. J. Elphick, P. S. Harlow, I. 'C
Moore, M. P. l.eMaster, and R. T. i\llason.
200 I. Movements, mating, a nd dispersal
of red-sided gartersnakes (111011111ophis
sirtalis parictnlis) from a communal den in
Manitoba. Copcia, Vol. 2001. No. I (Feb. 16,
2001 ): 82-91.
License for Zooplan and J Hazard photos: h11ps://
creativecommons.org/licenses/by-sa/3.0/
deed.en.

COLO RADO PARKS &amp; W ILDLIFE • 6060 Broadway, Denver, CO 802 16 • (303) 297- 1192 • cpw.state.co. us

�Habitat Scorecard for Red-sided Garter Snakes (v. Jan 2016)
Project Name: _ _ _ _ _ _ _ _ _ _ _ _ __ _ _ _ _ __ _ _ __ _

Date(s) of Assessment: _ _ _ _ _ _ _ __

Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value colum n. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Percent of emergent vegetation
&gt;60- 100%

36.4

&gt;40-60%

24.2

20-40%

12.1

Amphibians in same habitat
Abundant amphibians present

36.4

Only occasional amphibians noted

12. 1

Water quality
No visual evidence of turbidity or other pollutants

27.2

Some turbidity or p resence of other pollutants, but limited to sma ll and localized areas within the wetland; water
may be slig htly cloudy

18.2

Water is cloudy or has unnatural oil sheen, but the bottom is still visible (note: if the sheen breaks apart when you
run your finger t hrough it, it is a natural bacterial process and not water pollution)

9.1

Total (of 100 possible): add all numbers in before or after columns

After

�ASSESSING HABITAT QUALITY FOR PRIORITY WILDLIFE SPECIES IN COLORADO WETLANDS

Species Distribution
Range
The range of yellow mud turtles
extends from parts of Nebraska
through northern Mexico and from
far souilieastern Arizona to disjointed
populations in Illinois, Missouri, and
Iowa. In Colorado, yellow mud turtles
are known only in the far eastern
counties.
North America m ap used from U.S. Geologic

Kinostemon flavescens

Yellow mud turtles (Kinosternon flavescens, Family Kinosternidae) are found in wetlands on
the far eastern plains of Colorado. They lay their eggs in sandy soil.

Species Description
Identification
The carapace of an adult yellow mud
turtles is typically no longer than 6-7
inches, dome shaped and flat on top.
The marginal shields are often yellow.
TI1e turtles have relatively large heads
and yellow throats.
Preferred Habitats
Yellow mud turtles require iliree basic
habitats to complete th eir life cycle:
(1 ) an upland hibernation site, (2)
aquatic habitats, often wetlands and
other bottomlands, and (3) a sandy site
for aestivation (summer dormancy,
similar to winter hibernation) and
egg-laying. Additionally, iliey require
suitable habitat to m ove among iliese
sites. They will occupy almost any
wetland type iliat has adjacent sandy
areas, such as sand ridges and sand
hills.

Diet
Yellow mud trutles are primarily
carnivorous, though they do consume
duckweed, Their diet consists of
snails, fish, crayfish, and oilier macroinvertebrates.

r7.

Conservation Status
Due to low population numbers, in
Colorado, yellow mud turtles are listed
as a Species of Concern and Tier 2
Species of Greatest Conservation Need
(CPW 2015). Severe declines in Iowa,
Illinois, and Missouri are attributed to
lower water tables and wiilidrawal of
water from aquifers.

Survey (http://nas.cr.usgs.gov). Colorado m ap based
Hammerson (1999) and N DIS (2014).

COLORADO PARKS &amp; WILDLIFE • 6060 Broadway, Denver, CO 80216 • (303) 297-1192 • cpw.stale.co.us

�Preferred Habitat Conditions
Aestivation depth

2-l0 inches

Aestivation site relative to wetland

16 feet above elevation of wetland

Dominant vegetation

aquatic (submergent, floating, algae)

Hydrology of wetland

ephemeral wetlands; occasionally farm ponds

Landscape context

wetland adjacency to sand ridges, sandy rises, or
sand hills

Water depth

3-10 inches

Water temperature

64-68°F

Management Recommendations
This fact sheet contains easy-to-use guidelines for understanding habitat needs of
Colorado Parks and Wildlife priority wetland-dependent wildlife. A number of
practical steps can be taken to improve habitat for yellow mud turtles.
Hydrology

•

In spring, maintain water depth that limits access by terrestrial predators but
allows turtle hydration, reproduction and feeding.
In later summer and fall, reduce depth to promote kill of predatory fish and
predatory aquatic turtles.

Conservation

•
•
•
•

Maintain sand habitat between wetlands to facilitate seasonal movements and
gene flow.
Maintain mosaic of wetland diversity.
Remove barriers that prevent movement an1ong wetlands.
Control predators.

Acknowledgements
Tina Jackson (Colorado Parks and Wildlife)
reviewed an earlier version and provided input
on preferred habitat conditions.
Suggested Reading and Citations
Bernstein. N. P., and J. L. Christiansen. 201 I.
Response of a yellow mud turtle (Ki11ostemo11
flavescens Agassiz) community to habitat
change: management implications for a
nature preserve. Natural Areas /011rnal 31:
414-419.
Christiansen, ). L., J. A. Cooper, J. W. Bickham, B.
). Gallaway, and M. D. Springer. I 985. Aspects
of the natural history of the yellow mud
tu rtle Ki11ostemo11 flavescens (Kinoslernidae)
in Iowa: a proposed endangered species.
Sout/Jwestern Naturalist 30: 4 13-425.
CPW (Colorado Parks and Wildlife). 2015.
Stale Wildlife Action Plan: A Strategy for
Conserving Wildlife in Colorado. Denver,
Colorado.
Degenhardt, W. G., and ). L.. C hristiansen. 1974.
Distri bution and habitats of tu rtles in New
Mexico. Soutlnvestem Na turalist 19: 21-46.
Gregoire, D. R. 2015. Ki11osterno11 jlavesce11s.
USGS Nonindigenous Aquatic Species
Database, Gainesville, FL.
Hammerson, G. A. 1999. Amphibians and
Reptiles in Colorado: A Colorado Field
Guide. University Press of Colorado and
Colorado Division of Wildlife.
Iverson, J. B. 199 1. Life history and demography
of the yellow mnd turtle, Ki11ostemon
flavesce11s. Herpetologicn 47: 373-395.
NDIS (Natural Diversity Information Source).
20 I 4. On-line mapping tool for merly
available through Natural Resource Ecology
Laboratory. Colorado State University, Fort
Collins, Colorado.
Tuma, M. W. 2006. Range, habitat use, and
seasonal activity of the yellow mud turtle
(Ki11ostemon flavescens) in Northwestern
Illinois: implications for site-specific
conservation and management. Clielo11in11
Conservation and Biology 5: I 08-120.
Webster, C. 1986. SubsIrate preference and
activity in the turtle, Kinosternon flavescens.
Jou ma/ of Herpetology 20: 477-482.

COLORADO PARKS &amp; WTLDLU:E • 6060 Broadway, Denver, CO 80216 • {303) 297- 1192 • cpw.state.co.us

�Habitat Scorecard for Yellow Mud Turtles (v. Jan 2016)
Project Name:_ _ _ __ __ _ _ _ __ _ __ _ _ _ __ __ _ _ __

Date(s) of Assessment: _ _ __ _ __ __

Instructions: Enter one value that best describes early to mid-summer conditions of each habitat variable, using the numbers in the
value column. Habitat variables are in shaded boxes; ranges of condition are directly below each variable. If condition is outside
range or is not described, enter a zero.

Key habitat variable and conditions

Value

Before

Water

Ephemeral

13.7

Permanent

9.1

Substrate of wetland

Soft mud or muddy sand

12.2

Adjacent aestivation site (aestivation= dormancy to avoid summer heat, site must have sandy soil}

At least 5.5 yards above elevation of wetla nd and easy to access

12.2

Sandy soil depth of adjacent aestivation site

2- 10 inches

12.2

Percent emergent vegetation

&gt;70-1 00%

12.2

&gt;40 - 70%

8.2

20 - 40%

4.1

Predominant depth of water

3 - 10 inches

10.7

&gt; 10-16inches

7.2

Water temperature

64-68°F

10.7

&gt;68 - 79°F

7.2

&gt;79-91 °F

3.6

Percent of landscape within 100 yard dominated by grassland

&gt;70 - 100%

10.7

&gt;40-70%
0 - 40%

7.2
3.6

Percent of landscape within 100 yard dominated by grassland

Aquatic vegetation (submergent, floating, algae)
Total (of 100 possible): add all numbers in before or after columns

5.4

After

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              <text>Wildlife species profiles: Assessing habitat quality for priority wildlife species in Colorado wetlands</text>
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              <text>Wildlife species profiles, compiled in 2016, for priority wetland-dependent birds, mammals, fish, amphibians, and reptiles. The purpose of the profiles is to provide landowners and land managers with easy-to-use guidelines for understanding habitat needs of priority wetland-dependent wildlife.&amp;nbsp;&lt;br /&gt;&lt;br /&gt;&lt;a href="https://cpw.state.co.us/species-profiles"&gt;Find current species profiles on CPW website&lt;/a&gt;&lt;br /&gt;&lt;br /&gt;Species profiles included:&lt;br /&gt;&lt;strong&gt;BIRDS&lt;/strong&gt;&lt;br /&gt;&#13;
&lt;ul&gt;&#13;
&lt;li&gt;American bittern&lt;/li&gt;&#13;
&lt;li&gt;Bald eagle&amp;nbsp;&lt;/li&gt;&#13;
&lt;li&gt;Black rail&lt;/li&gt;&#13;
&lt;li&gt;Dabbling ducks&lt;/li&gt;&#13;
&lt;li&gt;Least tern&lt;/li&gt;&#13;
&lt;li&gt;Lewis's woodpecker&lt;/li&gt;&#13;
&lt;li&gt;Long-billed curlew&lt;/li&gt;&#13;
&lt;li&gt;Piping plover&lt;/li&gt;&#13;
&lt;li&gt;Sandhill crane&lt;/li&gt;&#13;
&lt;li&gt;Short-eared owl&lt;/li&gt;&#13;
&lt;/ul&gt;&#13;
&lt;strong&gt;MAMMALS&lt;br /&gt;&lt;/strong&gt;&#13;
&lt;ul&gt;&#13;
&lt;li&gt;Preble's jumping mouse&lt;/li&gt;&#13;
&lt;li&gt;River otter&lt;strong&gt;&lt;/strong&gt;&lt;/li&gt;&#13;
&lt;/ul&gt;&#13;
&lt;strong&gt;FISH&lt;br /&gt;&lt;/strong&gt;&#13;
&lt;ul&gt;&#13;
&lt;li&gt;Arkansas darter&amp;nbsp;&lt;/li&gt;&#13;
&lt;li&gt;Brassy minnow&lt;/li&gt;&#13;
&lt;li&gt;N. redbelly dace&lt;/li&gt;&#13;
&lt;li&gt;S. redbelly dace&lt;/li&gt;&#13;
&lt;/ul&gt;&#13;
&lt;strong&gt;HERPS&lt;br /&gt;&lt;/strong&gt;&#13;
&lt;ul&gt;&#13;
&lt;li&gt;N. leopard frog&lt;/li&gt;&#13;
&lt;li&gt;Plains leopard frog&lt;/li&gt;&#13;
&lt;li&gt;Red-sided garter snake&lt;/li&gt;&#13;
&lt;li&gt;Yellow mud turtle&lt;/li&gt;&#13;
&lt;/ul&gt;&#13;
&lt;br /&gt;Print copy located in CPW Research Library: FILE Fact Sheets</text>
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          <name>Creator</name>
          <description>An entity primarily responsible for making the resource</description>
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              <text>Colorado Parks &amp; WIldlife</text>
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          <name>Date Created</name>
          <description>Date of creation of the resource.</description>
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              <text>2016</text>
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