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Type: Fact Sheet
Subjects: Mule deer
Northwest Colorado
Wildlife management
Oil and gas development
Fact sheet
Type:Fact Sheet
Subject:Mule deer
Northwest Colorado
Wildlife management
Oil and gas development
Fact sheet
Description:Colorado Parks and Wildlife is concerned with the habitat needs and management of mule deer in the Piceance Basin, a 7,100 square mile area in northwest Colorado where natural gas resources are being developed extensively.
Description:

Wildlife Researcher, Ungulates/Carnivores

Current or Recent Research Projects

Areas of Interest and ExpertiseMy research focuses on large mammal ecology and management, with an emphasis on spatial ecology, population dynamics, and predator-prey interactions.

Streaming VideosEmblems of the West

Select Publications

Educ​ation

  • Ph.D., Wildlife, Fish & Conservation Biology – University of Massachusetts, 2017
  • M.S., Wildlife, Fish & Conservation Biology – University of Massachusetts, 2012
  • B.A., Philosophy – Haverford College, 2000

Current or Recent Positions

  • Wildlife Researcher – Colorado Parks and Wildlife, 2018-Present
  • Ecologist – U.S. Geological Survey, 2016-2018

Contact Information711 Independent AveGrand Junction, CO 81505

Email: nathaniel.rayl@state.co.usPhone: 970-255-6121
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Description:http://cpw.state.co.us/thingstodo/Pages/Statistics.aspxIn an effort to assist Colorado's big game hunters applying for the draw, Colorado Parks and Wildlife provides the following draw and hunting season information. These reports show how many licenses were available in all big game management units throughout Colorado, how many hunters applied for those limited licenses, how many of those hunters were successful drawing and how many preference points it took to be successful. [show more]
Type: Text
Subjects: Nuisance species
Wildlife management
Type:Text
Subject:Nuisance species
Wildlife management
Description:Following are summaries of Colorado’s most common regulations and statutes pertaining to nuisance wildlife questions. The laws and regulations in this handout are paraphrased for easier understanding and are intended only as a guide. Colorado’s wildlife statutes and regulations can be viewed in their entirety at Colorado Parks and Wildlife (CPW) offices or online at http://cpw.state.co.us/aboutus/Pages/Regulations.aspx. Please keep in mind that these laws note what is restricted by state statutes and permitted by CPW regulations; however, local laws and ordinances may be more specific about what is allowed at a certain location. It is your responsibility to determine whether it is legal to use a particular method of take in your city or county. CPW does not require nor request the removal of wildlife deemed a nuisance; the determination that a species has become a nuisance and should be removed is the prerogative of the landowner. There are many non-lethal methods for discouraging the presence of wildlife. For information about co-existing with wildlife, please contact your local CPW office or visit our website at cpw.state.co.us. [show more]
Type:Text
Subject:Aquatic nuisance species
ANS
Boating
Inspection
Description:The purpose of this workbook is to provide consistent guidance to State of Colorado Aquatic Nuisance Species (ANS) Trainers who are responsible for certification of individuals as Authorized Agents for the purpose of ANS Watercraft Inspection and Decontamination. It is of the highest importance that Authorized Locations (a.k.a. Watercraft Inspection and Decontamination Stations or WIDS) within Colorado are implemented and operated in a consistent fashion. The foundation of the WIDS network begins at certification training. Regardless of jurisdiction, Colorado Authorized Locations, Agents, and Operations should appear seamless to the boater because they are implemented with consistent messaging, education, inspection and decontamination procedures as taught in this course. [show more]
Type:Text
Subject:Aquatic nuisance species
ANS
Aquatic invasive species
AIS
Boating
Inspection
Description:Aquatic nuisance species (ANS) or aquatic invasive species (AIS) are also called non-native species, exotic species, non-indigenous species, noxious weeds, or pests. ANS can be plants or animals. Invasive aquatic plants are introduced plants that live either partially or completely submerged in the water and out-compete native species for light, space and nutrients creating a dense monoculture. Invasive aquatic animals also outcompete native species and require a watery habitat, but do not necessarily have to live entirely in water. [show more]
Type: Article
Subjects: Acoustic monitoring
Bayesian hierarchical model
Colorado
Foraging behavior
Herbivore foraging
Mule deer
<em>Odocoileus hemionus</em>
Spatial ecology
Type:Article
Subject:Acoustic monitoring
Bayesian hierarchical model
Colorado
Foraging behavior
Herbivore foraging
Mule deer
<em>Odocoileus hemionus</em>
Spatial ecology
Description:Foraging behavior underpins many ecological processes; however, robust assessments of this behavior for free-ranging animals are rare due to limitations to direct observations. We leveraged acoustic monitoring and GPS tracking to assess the factors influencing foraging behavior of mule deer (Odocoileus hemionus). We deployed custom-built acoustic collars with GPS radiocollars on mule deer to measure location-specific foraging. We quantified individual bites and steps taken by deer, and quantified two metrics of foraging behavior: the number of bites taken per step and the number of bites taken per unit time, which relate to foraging intensity and efficiency. We fit statistical models to these metrics to examine the individual, environmental, and anthropogenic factors influencing foraging. Deer in poorer body condition took more bites per step and per minute and foraged for longer irrespective of landscape properties. Other patterns varied seasonally with major changes in deer condition. In December, when deer were in better condition, they took fewer bites per step and more bites per minute. Deer also foraged more intensely and efficiently in areas of greater forage availability and greater movement costs. During March, when deer were in poorer condition, foraging was not influenced by landscape features. Anthropogenic factors weakly structured foraging behavior in December with no relationship in March. Most research on animal foraging is interpreted under the framework of optimal foraging theory. Departures from predictions developed under this framework provide insight to unrecognized factors influencing the evolution of foraging. Our results only conformed to our predictions when deer were in better condition and ecological conditions were declining, suggesting foraging strategies were state-dependent. These results advance our understanding of foraging patterns in wild animals and hig [show more]
Description:Short (1-5 page) leaflets that served as in-house scientific notes. The series was published from 1965-1996. The first 106 leaflets were published from 1965-1980; the last ten published sporadically until 1996. Leaflets are combined in one pdf file (with bookmarks). Leaflet numbers and titles are below. Print copies located in CPW Research Library: SK 351 .G21
# Title  Author
1 Upland game bird facts  Sandfort, Wayne W.
2 Care of big game hides  
3 Wild turkey hunting techniques  Burget, Martin L.
4 Productivity rates, age classes, and sex ratios of spring-caught beavers in Colorado  Rutherford, William H
5 Artificial revegetation on big game winter ranges White, Claude E., Jr.
6 Some effects of heavy use on browse plants  Shepherd, Harold R
7 Sex determination in dressed elk carcasses  Denney, Richard N.,
8 Warbles of cottontail rabbits  Pillmore, Richard E.
9 Lungworms in Colorado  Pillmore, Richard E.
10 Lungworm in bighorn sheep  Pillmore, Richard E.
11 Buffalo peaks bighorn seasons  Pillmore, Richard E.
12 Rabies and wildlife  Pillmore, Richard E.
13 Lungworm and lambs  Pillmore, Richard E.
14 Effects on pheasants of certain insecticides applied under modified field conditions in eastern Colorado : a thesis summary  Tigner, James R
15 Aging pheasant embryos  Sandfort, Wayne W.
16 Incubation, brooding, and care of pheasants  Mansfield, Willis G.
17 Relationship of pheasant weights to winter stress and starvation  Swope, Harold M
18 The history, function and establishment of exclosures in Colorado big game management  Baker, Bertram D.
19 Controlling rubber rabbitbrush with 2,4-D  Smith, Donald G
20 Colorado big game species as possible vectors of some livestock diseases  Denney, Richard N.,
21 Procedures in the appraisal of big game use of stacked alfalfa or native hay  Boyd, Raymond J
22 Duck wing surveys  Hopper, Richard M
23 Two condition indices of the Cache la Poudre mule deer herd and their application to management  Anderson, Allen E
24 Pellet group deposition rates for captive deer  McKean, William T.
25 Techniques for census of scaled quail  Hoffman, Donald M
26 Mourning dove migration in Colorado  Funk, Howard D.
27 Evaluation of a combined blue grouse-wild turkey hunting season  Myers, Gary T.
28 rev The effects of small rodents on deer winter range  Shepherd, Harold R
28 The effects of small rodents on deer winter range  Shepherd, Harold R.
29 Reliability of turkey sex and age ratio data based on hunter report card returns  Myers, Gary T.
30 Aerial waterfowl census techniques  Rutherford, William H
31 Browse range transects : purposes and guidelines for establishment  Baker, Bertram D.
32 An automatic camera recorder for census and identification of wildlife  Snyder, Warren D
33 Factors that should be considered in determining techniques for sexing and aging the southern white-tailed ptarmigan, lagopus leucurus altipetens  Rogers, Glenn E
34 Sex and age ratios of deer harvested under a hunters-choice, multiple-license hunting season  Rogers, Glenn E
35 Preparation of a potting medium for plants  Webster, Lawrence A.
36 Capturing big game animals by the use of drugs  Gordon, David F.
37 Weather and sage grouse productivity  Gill, R. Bruce
38 Yield and utilization of three deer winter range browse species  Medin, Dean E
39 General information on food and cover requirements of grouse in Colorado  Rogers, Glenn E
40 Chronology of waterfowl migration in Colorado  Rutherford, William H
41 Renovation of oakbrush habitat as a range improvement project for game clubs. Shepherd, Harold R
42 Survival of mountain mahogany transplants on two sites, Little Hills experiment station, Rio Blanco County, Colorado, 1965  McKean, William T.
43 A technique for mapping wildlife habitat in farmland areas  Snyder, Warren D
44 Deer mirrors : do they work? Boyd, Raymond J
45 Merriam's turkey roost preferences on mountain ranges  Hoffman, Donald M
46 Field methods for determination of sex and age of mourning doves  Funk, Howard D.
47 Use of paint for marking deer pellet-groups : a thesis summary  Kufeld, Roland C
48 Nesting structures for Canada geese  Grieb, Jack R
49 Sex and age determination of sage grouse from wing characteristics  Beck, Thomas D. I.
50 Photographic backdrop : a range study aid  Baker, Bertram D.
51 The Barbary sheep  Gordon, David F
52 Aids for the Rocky Mountain bighorn sheep hunter in Colorado. Tully, Robert J.
53 Effects of hunting on the selection of resting areas by mallards in west-central Colorado  Sandfort, Wayne W.
54 Determination of age and sex of the southern white-tailed ptarmigan  Braun, Clait E
55 Visual sex determination of mountain goats as related to selective hunting potential  Hibbs, L. Dale
56 The pheasant crowing count census and factors affecting its reliability  Swope, Harold M
57 Deer trapping and tagging results at the Little Hills Experiment Station. Bartmann, Richard M
58 Canada goose populations of Colorado  Grieb, Jack R
59 Pellet-plot robot : an aid in making pellet-group counts. Shepherd, Harold R
60 The breeding season in migratory mule deer  Anderson, Allen E
61 Air-ground comparison counts of breeding ducks in the San Luis Valley  Hopper, Richard M
62 Gestation periods, litter sizes, and breeding habits of Colorado furbearers  Rutherford, William H
63 A device for opening deer and elk jaws  Kufeld, Roland C.
64 Techniques for measuring shrub crown intercept when employing the line intercept method of sampling range vegetation  Baker, Bertram D.
65 A modified gallinaceous guzzler for scaled quail  Snyder, Warren D
66 Survival of transplanted cholla cacti  Hoffman, Donald M
67 Sagebrush vegetation before and six years after 2,4-D application  Anderson, Allen E
68 The microscopic technique in food habit studies  Bear, George D
69 Evaluation of aerial antelope census technique. Bear, George D
70 Antelope transplants in Colorado  Bear, George D
71 Antelope and net-wire fences  Bear, George D
72 Traps for controlling magpies  Kufeld, Roland C
73 Techniques for determining potentially critical deer highway crossings  Reed, Dale F.
74 Guidelines for range type-conversion projects in sage grouse range  Swope, Harold M
75 A pulsating light for identifying animals at night  Prenzlow, Edgar J.
76 A quadrat count system for estimating game population. Gill, R. Bruce
77 Deer-mirrors : a clearer picture  Gordon, David F
78 Guidelines for improving scaled quail habitat  Snyder, Warren D
79 A carrying crate for deer  Bartmann, Richard M.
80 Glossary of selected animal behavior definitions  Prenzlow, Edgar J.
81 Fencing for mule deer  McKean, William T.
82 Recommended practices in habitat management for pheasants in eastern Colorado  Snyder, Warren D.
82 rev. Recommended habitat management practices for pheasants in eastern Colorado  Snyder, Warren D
83 An effective turtle trap  Crawford, Gurney I.,
84 Evaluation of the Piceance Creek meadow deer count  Bartmann, Richard M
85 Use of ammonium nitrate-fuel oil mixtures in blasting potholes for wildlife  Hopper, Richard M
86 Determination of blue grouse sex and age from wing characteristics  Braun, Clait E
87 Stocking rates for mule deer and livestock on certain pinõn-juniper areas  McKean, William T.
88 Breeding dates of Colorado elk as estimated by fetal growth curves  Boyd, Raymond J
89 Influence of livestock and mule deer upon vigor of Indian ricegrass and tufted phlox on a pinõn-juniper range  McKean, William T.
90 Status of mountain goats in Colorado  [William H Rutherford] Rutherford, William H
91 Guidelines for evaluation of mountain goat transplant sites in Colorado  Rutherford, William H
92 Status of transplanted bighorn sheep in Colorado  Rutherford, William H
93 Guidelines for evaluating bighorn sheep transplanting sites in Colorado  Rutherford, William H
94 A video time-lapse system for wildlife surveillance  Reed, Dale F.
95 Techniques for supplementing diet, attracting and baiting bighorn sheep  Rutherford, William H
96 The relation of three physical condition indices of mule deer  Pojar, Thomas M
97 Guidelines for estimating deer numbers in connection with claims of damage to growing crops. Bartmann, Richard M
98 Postmortem thigh temperatures in mule deer  Reed, Dale F.
99 Seeding roadsides for pheasant nesting cover  Snyder, Warren D
100 Documenting lead poisoning in waterfowl  Adrian, William J.
101 A volunteer wing collection station  Hoffman, Richard W
102 Guideline to selecting rates of nitrogen fertilizer to increase herbage production on sagebrush winter ranges. Carpenter, Len H
103 A night-viewing device to monitor activities of wildlife  Carpenter, Len H
104 A life-table for managing deer populations  Freddy, David J.
105 Heart-rate measurements in mule deer research  Freddy, David J
106 Procedures for rearing wild ruminants in captivity  Neil, P. H.
107 Winter foods of mule deer in Piceance basin  Bartmann, Richard M
108 Shrub thicket establishment in Colorado's high plains  Snyder, Warren D
109 Appraisal of a quadrat census for mule deer in pinyon-juniper vegetation  Bartmann, Richard M
110 Portable computer system for field processing biotelemetry triangulation data  White, Gary C.
111 Identification of grouse species by hunters in northwestern Colorado : Implications for management  Giesen, Kenneth M
112 Expanding telemetry collar for elk calves  Bear, George D
113 Establishing switchgrass for wildlife in eastern Colorado  Snyder, Warren D
114 Bottle-raising wild ruminants in captivity. Wild, Margaret A.
115 Habitat management for upland game birds on eastern Colorado sandhill rangeland. Snyder, Warren D
116 Colorado pronghorn compatibility and conflicts with agriculture. Pojar, Thomas M.
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Description:

Led By

Fitzpatrick, Ryan

Project Status

Ongoing

Research Objective

Determine the amount and timing of native fish movement through the Owens-Hall fish passage structure.

Collaborators

Paul Foutz, Colorado Parks and Wildlife; David longrie and Kirsta Scherff-Norris, Colorado Springs Utilities

Project Description

Fountain Creek, Colorado, has a relatively intact native fish community that is dominated by the Species of Concern Flathead Chub, Platygobio gracilis and is also home to the state threatened Arkansas Darter, Etheostoma cragini. The native species community is strongest in the lower section of Fountain Creek, which is also the longest stretch of stream without a barrier (58-km). The first barrier that fish can encounter in this reach is the Owens-Hall diversion. To increase the stream reach available to native plains fishes, Colorado Springs Utilities installed a rock ramp fish passage structure on the diversion. This fishway was designed to act as a template to be used for barriers farther upstream. If this structure is effective at fish passage, it can be used as a relatively inexpensive template to increase connectivity within Fountain Creek and along the Front Range of Colorado. The purpose of this project is to evaluate the effectiveness of the Owens-Hall fish passage structure.

Three PIT tag reading arrays were used to be able to detect partial movement and direction of movement Flathead Chub were collected by electrofishing below the fish passage structure. All fish were checked for a PIT tag. Deploying of PIT tags consisted of sanitizing all scalpels, tags, and sutures in ethanol. An incision was made ventrally into the fish’s body cavity just posterior to the left pectoral fin. The 12-mm half-duplex Oregon RFID PIT tag was inserted and one suture was applied with two half-hitch knots.

Evaluation of the fish passage structure is ongoing, but Flathead Chub have been documented swimming through the fish passage structure. Future research will PIT tag additional species of fish focusing on the area directly downstream of the fish passage structure.

Technical Plans

See the Technical plans for the Owen’s Fish Passage Structure​.  ​

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