Study Areas

• Bear’s Ears elk herd (DAU E-2)
• Trinchera elk herd (DAU E-33)
• Uncompahgre Plateau elk herd (DAU E-20)

Project Status

Research Objectives

• To estimate calf and adult female survival and cause-specific mortality rates
• To quantify pregnancy rates and nutritional condition
• To evaluate factors influencing survival, pregnancy rates, and nutritional condition

Project Description

In 2017, researchers began a 2-year pilot study to investigate factors influencing elk recruitment in two study areas in the state. The initial pilot project was expanded into a 3^rd study area in 2019 and the research continues to better understand elk population dynamics in Colorado and determine how predators, habitat, and weather conditions are impacting elk recruitment in Colorado.

Reports:

Evaluating Factors Influencing Elk Recruitment in Colorado (2019-current)

Publications:

Crews, S., N.D. Rayl, M.W. Alldredge, E.J. Bergman, C.R. Anderson Jr., and G. Bastille-Rousseau. 2025. Drivers of spring migration phenology in Rocky Mountain elk. Scientific Reports 15:7807.

Eric Bergman, Nathaniel Rayl

Study Areas

• Avalanche Creek elk herd (DAU E-15)
• Bear’s Ears elk herd (DAU E-2) 

Project Status: Ongoing

Research Objectives

• To assess how elk respond to human recreation
• To evaluate the influence of human recreation on elk distribution
• To estimate calf and adult female survival and cause-specific mortality rates
• To quantify pregnancy rates and nutritional condition
• To evaluate factors influencing survival, pregnancy rates, and nutritional condition

Project Description

In Colorado, outdoor recreation contributes 511,000 jobs, $62.5 billion in economic output, and $9.4 billion in local, state, and federal tax revenue. Thus, the economies of Colorado, its counties, and its communities, rely on managing the landscape for a multitude of outdoor recreational opportunities.

The sensitivity of elk to human presence and human activity has been a topic of interest for many decades. In Colorado, increasing public concerns over human recreational use have coincided with declines in elk productivity, but a direct relationship to this activity in Colorado remains unaddressed. This research aims to better understand the influence of human recreation on elk behavior and distribution.

Reports

• Wolf ecology
  • Wolf habitat selection and movement during re establishment in Colorado
• Wildlife management
  • Statistical estimation of wolf abundance
  • Quantifying the effects of management practices and disease on wildlife harvest
• Wolf-livestock dynamics
  • Identifying and examining potential nonconsumptive effects of wolves on cattle
• Predator-prey dynamics
  • Examining wolf and hunter effects on elk movement, space use, and aggregation patterns
  • Quantifying conditions under which predators influence prey diseases​

Areas of Interest and Expertise​

My background is in terrestrial wildlife ecology with a focus on carnivores (primarily gray wolves) and infectious diseases. I am interested in many facets of wildlife ecology, especially social behaviors, predator-prey interactions, population dynamics, and disease dynamics. My work intersects wildlife ecology, wildlife management, statistics, and disease ecology.​

Select Publications

• Draper, J. P., E. E. Brandell, J. Isabelle, C. Jacques, C. McCoy, E. Michel, D. J. Storm, C. Ott-Conn, B. Wojcik, W. C. Turner, and D. P. Walsh. 2025. Doe diligence: a regional analysis of antlerless deer harvest regulations in the Midwestern United States of America. PLoS One, 20(6): p.e0324708. https://doi.org/10.1371/journal.pone.0324708
• Inzalaco, H. N. t., E. E. Brandell, S. P. Wilson, M. Hunsaker, D. R. Stahler, K. Woelfel, D. P. Walsh, T. Nordeen, D. J. Storm, and S. S. Lichtenberg. 2024. Detection of prions from spiked and free-ranging carnivore feces. Scientific Reports 14:3804. doi.org/10.1038/s41598-023-44167-7
• Cubaynes, S., E. E. Brandell, D. R. Stahler, D. W. Smith, E. S. Almberg, S. Schindler, R. K. Wayne, A. P. Dobson, B. M. vonHoldt, D. R. MacNulty, P. D. Cross, P. J. Hudson, and T. Coulson. 2022. Disease outbreaks select for mate choice and coat color in wolves. Science 378:300–303; DOI: 10.1126/science.abi8745
• Meyer, C. J., K. A. Cassidy, E. E. Stahler, E. E. Brandell, C. B. Anton, D. R. Stahler, and D. W. Smith 2022. Parasitic infection increases risk-taking in a social, intermediate host carnivore. Communications Biology 5:1–10; doi.org/10.1038/s42003-022-04122-0
• Brandell, E. E., M. K. Jackson, P. C. Cross, A. J. Piaggio, D. R. Taylor, D. W. Smith, B. Boufana, D. R. Stahler, and P. J. Hudson. 2022. Evaluating noninvasive methods for estimating cestode prevalence in a wild carnivore population. PLoS ONE 17(11): e0277420; doi.org/10.1371/journal.pone.0277420
• Brandell, E. E., D. J. Storm, T. R. Van Deelen, D. P. Walsh, and W. C. Turner. 2022. A call to action: standardizing white-tailed deer harvest data in the Midwestern United States and implicationsfor quantitative analysis and disease management. Frontiers in Ecology and Evolution; https://doi.org/10.3389/fevo.2022.943411
• Gilbertson, M. L., E. E. Brandell, M. E. Pinkerton, N. M. Meaux, M.  Hunsaker, D. Jarosinski, W. Ellarson, D. P. Walsh, D. J. Storm, and W. C. Turner. 2022. Cause of death, pathology, and chronic wasting disease status of white-tailed deer (Odocoileus virginianus) mortalities in Wisconsin, USA. Journal of Wildlife Diseases 54:803–815; DOI: 10.7589/JWD-D-21-00202​​
• ​Brandell, E. E., P. C. Cross, D. W. Smith, W. Rogers, N. L. Galloway, D. MacNulty, D. R. Stahler, J. Treanor, and P. J. Hudson. 2022. Examination of the interaction between age‐specific predation and chronic disease in the Greater Yellowstone Ecosystem​. Journal of Animal Ecology. 91:1373-1384, https://doi.org/10.1111/1365-2656.13661​
• Rogers, W., E. E. Brandell, and P. C. Cross. 2022. Epidemiological differences between sexes affect management efficacy in simulated chronic wasting disease systems. Journal of Applied Ecology 59:1122-1133. https://doi.org/10.1111/1365-2664.14125​
• Brandell, E.E., D. J. Becker, L. Sampson, and K. M. Forbes. 2021. Demography, education, and research trends in the interdisciplinary field of disease ecology. Ecology and evolution 11:17581-17592, https://doi.org/10.1002/ece3.8466​
• Brandell, E. E., A. P. Dobson, P. J. Hudson, P. C. Cross, and D. W. Smith. 2021. A metapopulation model of social group dynamics and disease applied to Yellowstone wolves. Proceedings of the National Academy of Sciences, 118(10), https://doi.org/10.1073/pnas.2020023118
• Brandell, E.E., P. C. Cross, M. E. Craft, D. W. Smith, E. J. Dubovi, M. L.  Gilbertson, T. Wheeldon, J. A. Stephenson, S. Barber-Meyer, B. L. Borg, and M. Sorum. 2021. Patterns and processes of pathogen exposure in gray wolves across North America​​. Scientific reports 11(1):1-14.​
• DeCandia, A. L., E. C. Schrom, E. E. Brandell, D. R. Stahler, and B. M. vonHoldt. 2021. Sarcoptic mange severity is associated with reduced genomic variation and evidence of selection in Yellowstone National Park wolves (Canis lupus). Evolutionary applications, 14(2):429-445, https://doi.org/10.1111/eva.13127
• Brandell, E. E., N. M. Fountain‐Jones, M. L. Gilbertson, P. C. Cross, P. J. Hudson, D. W. Smith, D. R. Stahler, C. Packer, and M. E. Craft. 2021. Group density, disease, and season shape territory size and overlap of social carnivores. Journal of Animal Ecology, 90(1):87-101.

Education

• Ph.D., Ecology – Pennsylvania State University, 2021
• B.S., Wildlife Biology –​ University of Montana, 2015 

Current or Recent Positions

• Wildlife Research Scientist – Colorado Parks and Wildlife, 2022-Present
• Postdoctoral Associate –​ Wisconsin Cooperative Wildlife Research Unit, Dept. of Forest and Wildlife Ecology, University of Wisconsin-Madison, 2020-2022

Contact Information

317 W. Prospect Rd.​
Fort Collins, CO 80526

Email: ellen.brandell@stat​e.co.us​

Phone: ​970-698-0252

Bobcat Ecology and Management

• Bobcat ecology
  • Where do bobcats occur?
    • Determining where bobcats select habitats and resources
  • What do bobcats eat?
    • Spatial and temporal variation in prey availability
    • Determining dietary composition and prey use/selection
  • Bobcat monitoring and management
    • How many bobcats are out there?
      • Evaluating statistical methods for estimating bobcat abundance/density
    • What are causes of bobcat mortality?
      • Estimating the relative magnitude of mortality sources
    • Bobcat reproduction
      • Spatial and temporal variation in bobcat family group occurrence
    • Understanding changes in bobcat population size
      • Determining potential important factors affecting bobcat population density and dynamics (e.g. habitat, prey availability, and harvest rate)
    • Improving bobcat data collection methods and population inference
      • Enhancing age classification through dentition and teeth measurements
      • Exploring cutting-edge methods in population size/density estimation

Areas of Interest and Expertise​

My background is in terrestrial wildlife ecology with a focus on carnivores (primarily bears and bobcats) and harvest, though my interests vary widely. I have interest in exploring both the lives and deaths of wildlife and how individual contexts scale up to have population level effects. This encapsulates both the contextual decisions of individual animals and how we manage and conserve their populations. I have researched several themes, including animal movement, connectivity, sociality, carcass ecology, and the effects of harvest. Moreover, I have utilized a multitude of data collection methods (e.g. location data, genetics, isotopes) and statistical approaches to answer research questions under these themes. My overarching aim is help provide pragmatic answers to (1) applied questions and (2) contribute to the ever-advancing suite of data collection and analytical tools available to practitioners. Moreover, I hope to (3) help bridge the knowledge gap between wildlife science, research findings, and people.

Select Publications

• Heeres, R. W., Leclerc, M., Frank, S., Kopatz, A., Pelletier, F., & Zedrosser, A. (2025). Flexibility in female spatiotemporal behavioral tactics to counter infanticide risk during the mating season. Movement Ecology, 13(1), 35.
• Hertel, A. G., Parres, A., Frank, S. C., Renaud, J., Selva, N., Zedrosser, A., Balkenhol, N., Maiorano, L., Fedorca, A., Dutta, T., Bogdanovic, N., Bragalanti, N., Chiriac, S., Cirovic, D., Ciucci, P., Domokos, C., Fedorca, M., Filacorda, S., Findo, S., . . . De Barba, M. (2025). Human Footprint and Forest Disturbance Reduce Space Use of Brown Bears (Ursus arctos) Across Europe. Glob Chang Biol, 31(1), e70011. https://doi.org/10.1111/gcb.70011
• Lucas, P. M., Thuiller, W., Talluto, L., Polaina, E., Albrecht, J., Selva, N., De Barba, M., Penteriani, V., Gueguen, M., Balkenhol, N., Dutta, T., Fedorca, A., Frank, S. C., Zedrosser, A., Afonso-Jordana, I., Ambarli, H., Ballesteros, F., Bashta, A. T., Bilgin, C. C., . . . Pollock, L. J. (2025). Trophic Interactions Are Key to Understanding the Effects of Global Change on the Distribution and Functional Role of the Brown Bear. Glob Chang Biol, 31(6), e70252. https://doi.org/10.1111/gcb.70252
• Zarzo-Arias, A., Heeres, R. W., Hertel, A. G., Leclerc, M., Frank, S., Steyaert, S. M., ... & Zedrosser, A. (2025). “The landscape of love”: sex-specific habitat-use during the mating season in a solitary large carnivore. Landscape Ecology, 40(12), 222.
• Hertel, A. G., Albrecht, J., Selva, N., Sergiel, A., Hobson, K. A., Janz, D. M., Mulch, A., Kindberg, J., Hansen, J. E., Frank, S. C., Zedrosser, A., & Mueller, T. (2024). Ontogeny shapes individual dietary specialization in female European brown bears (Ursus arctos). Nat Commun, 15(1), 10406. https://doi.org/10.1038/s41467-024-54722-z
• Heeres, R. W., Leclerc, M., Frank, S., Kopatz, A., Pelletier, F., & Zedrosser, A. (2024). Are nonsocial species more social than we think? Seasonal patterns in sociality in a solitary terrestrial carnivore. Animal Behaviour, 216, 107-130. https://doi.org/10.1016/j.anbehav.2024.07.022
• Zedrosser, A., Frank, S. C., Hansen, J. E., Steyaert, S. M. J. G., & Swenson, J. E. (2023). The social system of a "nonsocial" species, the brown bear. In Bear and Human (Vol. 3, pp. 121-128). Brepols Publishers. https://doi.org/doi:10.1484/M.TANE-EB.5.134329
• Hansen, J. E., Hertel, A. G., Frank, S. C., Kindberg, J., & Zedrosser, A. (2023). The role of familial conflict in home range settlement and fitness of a solitary mammal. Animal Behaviour, 202, 39-50. https://doi.org/10.1016/j.anbehav.2023.05.006
• Dutta, T., De Barba, M., Selva, N., Fedorca, A. C., Maiorano, L., Thuiller, W., Zedrosser, A., Signer, J., Pflüger, F., Frank, S., Lucas, P. M., & Balkenhol, N. (2023). An objective approach to select surrogate species for connectivity conservation. Frontiers in Ecology and Evolution, 11. https://doi.org/10.3389/fevo.2023.1078649
• Thorsen, N. H., Hansen, J. E., Stoen, O. G., Kindberg, J., Zedrosser, A., & Frank, S. C. (2022). Movement and habitat selection of a large carnivore in response to human infrastructure differs by life stage. Mov Ecol, 10(1), 52. https://doi.org/10.1186/s40462-022-00349-y
• Hansen, J. E., Hertel, A. G., Frank, S. C., Kindberg, J., & Zedrosser, A. (2022). Social environment shapes female settlement decisions in a solitary carnivore. Behav Ecol, 33(1), 137-146. https://doi.org/10.1093/beheco/arab118
• Arnberg, M. P., Frank, S. C., Blaalid, R., Davey, M. L., Eycott, A. E., & Steyaert, S. (2022). Directed endozoochorous dispersal by scavengers facilitate sexual reproduction in otherwise clonal plants at cadaver sites. Ecol Evol, 12(1), e8503. https://doi.org/10.1002/ece3.8503
• Frank, S. C., Pelletier, F., Kopatz, A., Bourret, A., Garant, D., Swenson, J. E., Eiken, H. G., Hagen, S. B., & Zedrosser, A. (2021). Harvest is associated with the disruption of social and fine-scale genetic structure among matrilines of a solitary large carnivore. Evol Appl, 14(4), 1023-1035. https://doi.org/10.1111/eva.13178
• Frank, S. C., Blaalid, R., Mayer, M., Zedrosser, A., & Steyaert, S. (2020). Fear the reaper: ungulate carcasses may generate an ephemeral landscape of fear for rodents. R Soc Open Sci, 7(6), 191644. https://doi.org/10.1098/rsos.191644
• Rivrud, I. M., Frank, S. C., Bischof, R., Mysterud, A., Steyaert, S., Hertel, A. G., Hagen, S. B., Eiken, H. G., Swenson, J. E., & Zedrosser, A. (2019). Heritability of head size in a hunted large carnivore, the brown bear (Ursus arctos). Evol Appl, 12(6), 1124-1135. https://doi.org/10.1111/eva.12786
• Badia, R., Blaalid, R., Puliti, S., Steyaert, S. M. J. G., & Frank, S. C. (2019). Reindeer carcasses provide foraging habitat for insectivorous birds of the alpine tundra. Ornis Norvegica, 42, 36-40. https://doi.org/10.15845/on.v42i0.2639
• Steyaert, S., Frank, S. C., Puliti, S., Badia, R., Arnberg, M. P., Beardsley, J., Okelsrud, A., & Blaalid, R. (2018). Special delivery: scavengers direct seed dispersal towards ungulate carcasses. Biol Lett, 14(8). https://doi.org/10.1098/rsbl.2018.0388
• Frank, S. C., Leclerc, M., Pelletier, F., Rosell, F., Swenson, J. E., Bischof, R., Kindberg, J., Eiken, H. G., Hagen, S. B., & Zedrosser, A. (2018). Sociodemographic factors modulate the spatial response of brown bears to vacancies created by hunting. J Anim Ecol, 87(1), 247-258. https://doi.org/10.1111/1365-2656.12767
• Leclerc, M., Frank, S. C., Zedrosser, A., Swenson, J. E., & Pelletier, F. (2017). Hunting promotes spatial reorganization and sexually selected infanticide. Sci Rep, 7, 45222. https://doi.org/10.1038/srep45222
• Frank, S. C., Ordiz, A., Gosselin, J., Hertel, A., Kindberg, J., Leclerc, M., Pelletier, F., Steyaert, S. M. J. G., Støen, O.-G., Van de Walle, J., Zedrosser, A., & Swenson, J. E. (2017). Indirect effects of bear hunting: a review from Scandinavia. Ursus, 28(2), 150-164. https://doi.org/10.2192/ursu-d-16-00028.1
• Steyaert, S. M. J. G., Zedrosser, A., Elfström, M., Ordiz, A., Leclerc, M., Frank, S. C., Kindberg, J., St⊘en, O. G., Brunberg, S., & Swenson, J. E. (2016). Ecological implications from spatial patterns in human‐caused brown bear mortality. Wildlife Biology, 22(4), 144-152. https://doi.org/10.2981/wlb.00165
• Frank, S. C., Steyaert, S. M., Swenson, J. E., Storch, I., Kindberg, J., Barck, H., & Zedrosser, A. (2015). A "clearcut" case? Brown bear selection of coarse woody debris and carpenter ants on clearcuts. For Ecol Manage, 348, 164-173. https://doi.org/10.1016/j.foreco.2015.03.051

Education

• Ph.D., Ecology – University of South-Eastern Norway (formerly Telemark University College), 2017
• M.S., Forest Ecology and Management – University of Freiburg, 2011
• B.S., Wildlife Biology –​ Colorado State University, 2003

Current or Recent Positions

• Wildlife Research Scientist – Colorado Parks and Wildlife, State of Colorado, 2022-Present
• Statistical Analyst / Quality Specialist Colorado Department of Public Health and Environment, 2020-2022, State of Colorado
• Postdoctoral Associate –​ “BearConnect”, Department of Nature, Health, and Environment, University of South-Eastern Norway, 2017-2020

Contact Information

346 Co Rd 362
Hot Sulphur Springs, CO 80451

Email: shane.frank@state.co.us

Phone: ​970-646-2961

Current or Recent Research Projects

Mitigating Impacts of Energy Development on Piceance Basin Mule Deer Populations

Areas of Interest and Expertise

My research focuses on large mammal ecology and management, with an emphasis on population dynamics, population monitoring, and population genetics.

Streaming Videos

Wild Science: Mule Deer in the Piceance Basin

Select Publications

• Crews, S., N. D. Rayl, M. W. Alldredge, E. J. Bergman, C. R. Anderson Jr., E. H. VanNatta, J. D. Holbrook, and G. Bastille-Rousseau. 2025. Hierarchy in structuring of resource selection: understanding elk selection across space, time, and movement strategies. Ecology and Evolution 15:e71097. https://doi.org/10.1002/ece3.71097
• Crews, S., N. D. Rayl, M. W. Alldredge, E. J. Bergman, C. R. Anderson Jr., and G. Bastille-Rousseau. 2025. Drivers of spring migration phenology in Rocky Mountain elk. Scientific Reports 15:7807. https://doi.org/10.1038/s41598-025-91947-4
• Hurley, M. A., C. R. Anderson Jr., T. D. Forrester, and J. A. Gude. 2023. Carnivore-prey relationships. Pages 125-139 in J. R. Heffelfinger and P. R. Krausman, editors. Ecology and management of black-tailed and mule deer of North America. CRC Press, Boca Raton, Florida, USA.​
• ​Johnston, D. H., and C. R. Anderson Jr. 2023. Plant and mule deer responses to pinyon-juiper removal by three mechanical methods. Wildlife Society Bulletin 47:e1421​.  https://doi.org/10.1002/jwmg.22110​
• Bonar, M., S. J. Anderson, C. R. Anderson Jr, G. Wittemyer, J. M. Northrup, and A. B. Shafer. 2022. Genomic correlates for migratory direction in a free-ranging cervid. Proceedings of the Royal Society B 289:20221969. https://doi.org/10.1098/rspb.2022.1969​
• Johnston, D. H., C. R. Anderson Jr., R. D. Applegate, L. Bailey, E. Cooch, J. Feiberg, A. B. Franklin, R. J. Gutiérrez, K. V. Miller, J. D. Nichols, N. D. Niemuth, D. Otis, C. A. Ribic, M. M. Rowland, and T. L. Shaffer. 2021. Editor's Message: A perspective on the Journal of Wildlife Management. Journal of Wildlife Management 87:1305-1308.  https://doi.org/10.1002/jwmg.22110
• Swearingen, T. C., R. W. Klaver, C. R. Anderson Jr., and C. N. Jacques. 2023. Influence of camera model and alignment on the performance of paired camera stations. Wildlife Society Bulletin 47:e1422. doi.org/ 10.1002/wsb.1422
• Rheault H., C. R. Anderson Jr, M. Bonar, R. R. Marrotte, T. R. Ross, G. Wittem​yer and J. M. Northrup. 2021. Some memories never fade: inferring multi-scale memory effects on habitat selection of a migratory ungulate using step-selection functions. Frontiers in Ecology and Evolution 9:702818. https://doi.org/10.3389/fevo.2021.702818
• Northrup, J. M., ​C. R. Anderson Jr., B. D. Gerber, and G. Wittemyer. 2021. Behavioral and demographic responses of mule deer to energy development on winter range. Wildlife Monographs: 208:1–37. https://doi.org/10.1002/wmon.1060
• Bishop, C. J., M. W. Alldredge, D. P. Walsh, E. J. Bergman, C. R. Anderson Jr., D. Kilpatrick, J. Bakel, and C. Febvre. 2019. A noninvasive automated device for remotely collaring and weighing mule deer. Wildlife Society Bulletin 43:717-725. https://doi.org/10.1002/wsb.1034   
• Northrup, J. M., A. Arvin, C. R. Anderson Jr., E. Brown, and G. Whittemyer. 2019. On-animal acoustic monitoring provides insight to ungulate foraging behavior. Journal of Mammalogy 100:1479–1489. https://doi.org/10.1093/jmammal/gyz124
• Jacques, C. N., R. W. Klaver, T. S. Swearingen, E. D. Davis, C. R. Anderson Jr., J. A. Jenks, C. S. DePerno, and R. D. Bluett. 2019. Estimating density and detection of bobcats in a fragmented Midwestern landscapes using spatial capture-recapture data from camera traps. Wildlife Society Bulletin 43:256–264.  https://doi.org/10.1002/wsb.968
• Peterson, M. E., C. R. Anderson Jr., M. W. Alldredge, and P. F. Doherty Jr. 2018. Using maternal mule deer movements to estimate timing of parturition and assist fawn captures. Wildlife Society Bulletin 42:616–621. https://doi.org/10.1002/wsb.935
• Peterson, M.E., C. R. Anderson Jr., J. M. Northrup, and P. F. Doherty Jr. 2018. Mortality of mule deer fawns in a natural gas development area. The Journal of Wildlife Management 82:1135–1148. https://doi.org/10.1002/jwmg.21476
• Bergman, E. J., C. R. Anderson Jr., C. J. Bishop, A. A. Holland, and J. M. Northrup. 2017. Variation in ungulate body fat: individual versus temporal effects. The Journal of Wildlife Management 82:130–137. https://doi.org/10.1002/jwmg.21334
• Lendrum, P. E., J. M. Northrup, C. R. Anderson Jr.​, G. E. Liston, C. L. Aldridge, K. R. Crooks, and G. Wittemyer. 2017. Predation risk across a dynamic landscape: effects of anthropogenic land use, natural landscape features, and prey distribution. Landscape Ecology 33:151-170. https://doi.org/10.1007/s10980-017-0590-z
• Peterson, M. E., C. R. Anderson Jr., J. M. Northrup, and P. F. Doherty Jr. 2017. Reproductive success of mule deer in a natural gas development area. Wildlife Biology 4:1-5. https://doi.org/10.2981/wlb.00341
• Northrup, J. M., C. R. Anderson Jr., and G. Wittemyer. 2015. Quantifying spatial habitat loss from hydrocarbon development through assessing habitat selection patterns of mule deer. Global Change Biology 21:3961–3970. https://doi.org/10.1111/gcb.13037
• Lendrum, P. E., C. R. Anderson Jr., K. L. Monteith, J. A. Jenks, R. T. Bowyer. 2014. Relating the movement of a rapidly migrating ungulate to spatiotemporal patterns of forage quality. Mammalian Biology 79:369–375. https://doi.org/10.1016/j.mambio.2014.05.005
• Freeman, E. D., R. T. Larsen, M. E. Peterson, C. R. Anderson Jr., K. R. Hersey, B. R. McMillan. 2014. Effects of male‐biased harvest on mule deer: implications for rates of pregnancy, synchrony, and timing of parturition. Wildlife Society Bulletin 38:806–811. https://doi.org/10.1002/wsb.450
• Northrup, J. M., M. B. Hooten, C. R. Anderson Jr., and G. Wittemyer. 2013. Practical guidance on characterizing availability in resource selection functions under a use–availability design. Ecology 94:1456‐1463. https://doi.org/10.1890/12–1688.1
• Lendrum, P. E., C. R. Anderson Jr., K. L. Monteith, J. A. Jenks and R. T. Bowyer. 2013. Mig​rating mule deer: effects of anthropogenically altered landscapes. PLoS ONE 8(5): e64548.  https://doi.org/10.1371/journal.pone.0064548
• Lendrum, P. E., C. R. Anderson Jr., R. A. Long, J. G. Kie, and R. T. Bowyer. 2012. Habitat se​lection by mule deer during migration: effects of landscape structure and natural–gas development. Ecosphere 3:1–19. https://doi.org/10.1890/ES12–00165.1
• Anderson, C. R. Jr., F. G. Lindzey, K. H. Knopff, M. G. Jalkotzy, and M. S. Boyce. 2010. Cougar management in North America. Pages 41-54 in M. Hornocker and S. Negri, Editors. Cougar: Ecology and Conservation. University of Chicago Press, Chicago, IL, USA.
• Anderson, C. R. Jr., and F. G. Lindzey. 2005. Expe​rimental evaluation of population trend and harvest composition in a Wyoming cougar population. Wildlife Society Bulletin 33:179-188.  https://doi.org/10.2193/0091-7648(2005)33[179:EEOPTA]2.0.CO;2
• Anderson, C. R. Jr., and F. G. Lindzey. 2003. Estimating cougar predation rates from GPS location clusters. The Journal of Wildlife Management 67:307-316. https://www.jstor.org/stable/3802772
• Anderson, C. R. Jr., M. A. Ternent, and D. S. Moody. 2002. Grizzly bear-cattle interactions on two cattle allotments in northwest Wyoming. Ursus 13:247-256. https://www.jstor.org/stable/3873205

Education

• Ph.D., Zoology and Physiology — University of Wyoming, 2003
• M.S., Zoology and Physiology — University of Wyoming, 1994
• B.S., Wildlife Biology — Colorado State University, 1990

Current or Recent Positions

• Mammals Researcher Section Leader — Colorado Parks and Wildlife, 2013-Present
• Wildlife Researcher — Colorado Parks and Wildlife, 2006-2013
• Affiliate Faculty — Idaho State University, 2009-Present
• Affiliate Faculty — Colorado State University, 2012-Present

Current or Recent Research Projects

• Snowshoe Hare Response to Silvicultural Treatments
• Monitoring Canada Lynx in Colorado
• Mammal and Breeding Bird Response to Bark Beetle Outbreaks

Areas of Interest and Expertise

My research focuses on estimating wildlife population parameters and forest carnivore ecology, conservation and monitoring. 

Streaming Videos

• Catching pine
• Lynx: shadows of the forest
• Snowshoe hare
• Snowshoe hare releases
• Snowshoe hare trapping, collaring and release

Select Publications

• Bosco-Lauth, A. M., S. M. Porter, R. M. Maison, K. A. Fox, J. M. Marano, J. S. Ivan, K. Griffin, M. Quilici, and J. J. Root. 2025. Rabbit Hemorrhagic Disease Virus 2 experimental infection in snowshoe hares (Lepus americanus). Journal of Wildlife Diseases 61:674-684. doi: 10.7589/JWD-D-24-00141
• Devarajan, K. [numerous authors], J. S. Ivan, [numerous authors]. 2025. When the wild things are: defining mammalian diel activity and plasticity. Science Advances 11:eado3843. DOI: 10.1126/sciadv.ado3843
• Ivan, J. S. 2024. Canada Lynx (lynx Canadensis). Pages 129-152 in J. E. Cartron and J. K. Frey, Eds. Wild Carnivores of New Mexico. University of New Mexico Press.
• Hooten, M. B., M. R. Schwob, D. S. Johnson and J. S. Ivan. 2024. Geostatistical capture-recapture models​​. Spatial Statistics 59:100817. doi.org/10.1016/j.spasta.2024.100817​
• Squires, J. R., Olson, L. E., Ivan, J. S., McDonald, P. M., and Holbrook, J. D. 2024. Anthropogenically protected but naturally disturbed: a specialist carnivore at its southern range periphery. Biodiversity and Conservation. https://doi.org/10.1007/s10531-024-02978-8
• Hooten, M. B., M. R. Schwob, D. S. Johnson, and J. S. Ivan. 2023. Multistage hierarchical capture-recapture models. Environmetrics 34:e2799. doi.org/10.1002/env.2799
• Ivan, J. S., E. S. Newkirk, and B. D. Gerber. 2023. Differential impacts of spruce beetle outbreaks on snowshoe hares and red squirrels in the southern Rocky Mountains​. Forest Ecology and Management 544:121147. doi.org/10.1016/j.foreco.2023.121147
• Squire​s, J. R., J. S. Ivan, K. E. Paolini, L. E. Olson, G. M. Jones, and J. D. Holbrook​​. 2022. Keystone structures maintain forest function for Canada lynx after large-scale spruce beetle outbreak​​. Environmental Research: Ecology 2:011001. https://doi.org/10.1088/2752-664X/ac8eb7
• Van Ee, J. J., J. S. Ivan, and M. B. Hooten. 2022. Community confounding in joint species distribution models. Scientific Reports 12:1–14. https://doi.org/10.1038/s41598-022-15694-6​
• Kraberger, S., L. E. K. Serieys, C. Richet, N. M. Fountain-Jones, G. Baele, J. M. Bishop, M. Nehring, J. S. Ivan, E. S. Newkirk, J. R. Squires, M. C. Lund, S. P. D. Riley, C. C. Wilmers, P. D. van Helden, K. Van Doorslaer, M. Culver, S. VandeWoude, D. P. Martin, and A. Varsani. 2021. Complex evolutionary history of felid anelloviruses. Virology 562:176–189. https://doi.org/10.1016/j.virol.2021.07.013
• Olson, L. E., N. Bjornlie, G. Hanvey, J. D. Holbrook, J. S. Ivan, S. Jackson, B. Kertson, T. King, M. Lucid, D. Murray, R. Naney, J. Rohrer, A. Scully, D. Thornton, Z. Walker, and J. R. Squires. 2021. Improved prediction of Canada lynx distribution through regional model transferability and data efficiency. Ecology and Evolution 11:1667–1690. https://doi.org/10.1002/ece3.7157
• Squires, J. R., J. D. Holbrook, L. E. Olson, J. S. Ivan, R. W. Ghormley, and R. L. Lawrence. 2020. A specialized forest carnivore navigates landscape-level disturbance: Canada lynx in spruce-beetle impacted forests. Forest Ecology and Management 475:118400. https://doi.org/10.1016/j.foreco.2020.118400
• Tabak, M. A., M. S. Norouzzadeh, D. W. Wolfson, E. J. Newton, R. K. Boughton, J. S. Ivan, E. A. Odell, E. S. Newkirk, R. Y. Conrey, J. Stenglein, F. Iannarilli, J. Erb, R. K. Brook, A. J. Davis, J. Lewis, D. P. Walsh, J. C. Beasley, K. C. VerCauteren, J. Clune, and R. S. Miller. 2020. Improving the accessibility and transferability of machine learning algorithms for identification of animals in camera trap images: MLWIC2. Ecology and Evolution 10:10374-10383. https://doi.org/10.1002/ece3.6692
• Latif, Q. S., J. S. Ivan, A. E. Seglund, D. L. Pavlacky, and R. L. Truex. 2020. Avian relationships with bark beetle outbreaks and underlying mechanisms in lodgepole pine and spruce-fir forests of Colorado. Forest Ecology and Management 464:118043. https://doi.org/10.1016/j.foreco.2020.118043
• Lukacs, P. M., D. Evans Mack, R. Inman, J. A. Gude, J. S. Ivan, R. P. Lanka, J. C. Lewis, R. A. Long, R. Sallabanks, Z. Walker, S. Courville, S. Jackson, R. Kahn, M. K. Schwartz, S. C. Torbit, J. S. Waller, and K. Carroll. 2020. Wolverine occupancy, spatial distribution, and monitoring design. The Journal of Wildlife Management 84:841–851. https://doi.org/10.1002/jwmg.21856
• Zimova, M., A. P. Sirén, J. J. Nowak, A. M. Bryan, J. S. Ivan, T. L. Morelli, S. L. Suhrer, J. Whittington, and L. S. Mills. 2019. Local climate determines vulnerability to camouflage mismatch in snowshoe hares. Global Ecology and Biogeography 29:503–515.  https://doi.org/10.1111/geb.13049
• Gigliotti, L. C., N. D. Berg, R. Boonstra, S. M. Cleveland, D. R. Diefenbach, E. M. Gese, J. S. Ivan, K. Kielland, C. J. Krebs, A. V. Kumar, L. S. Mills, J. N. Pauli, H. B. Underwood, E. C. Wilson, and M. J. Sheriff. 2019. Latitudinal variation in snowshoe hare (Lepus americanus) body mass: a test of Bergmann’s rule. Canadian Journal of Zoology 98:88–95. https://doi.org/10.1139/cjz-2019-0184
• Squires, J. R., L. E. Olson, E. K. Roberts, J. S. Ivan, and M. Hebblewhite. 2019. Winter recreation and Canada lynx: reducing conflict through niche partitioning. Ecosphere 10:e02876. https://doi.org/10.1002/ecs2.2876
• Ivan, J. S. 2019. Density estimation. Pages 20.1–20.19 in E. G. Cooch and G. C. White, editors. Program MARK - 'a gentle introduction'.
• Ayers, C. R., K. C. Hanson-Dorr, K. Stromborg, T. W. Arnold, J. S. Ivan, and B. S. Dorr. 2019. Survival, fidelity, and dispersal of double-crested cormorants on two Lake Michigan Islands. The Auk 136:1-10. https://doi.org/10.1093/auk/ukz040​
• Magee, P. A., J. D. Coop, and J. S. Ivan. 2019. Thinning alters avian occupancy in piñon–juniper woodlands. The Condor 121:1-17. https://doi.org/10.1093/condor/duy008
• Tabak, M. A., M. S. Norouzzadeh, D. W. Wolfson, S. J. Sweeney, K. C. Vercauteren, N. P. Snow, J. M. Halseth, P. A. Di Salvo, J. S. Lewis, M. D. White, B. Teton, J. C. Beasley, P. E. Schlichting, R. K. Boughton, B. Wight, E. S. Newkirk, J. S. Ivan, E. A. Odell, R. K. Brook, P. M. Lukacs, A. K. Moeller, E. G. Mandeville, J. Clune, and R. S. Miller. 2018. Machine learning to classify animal species in camera trap images: applications in ecology. Methods in Ecology and Evolution 10:585–590.  https://doi.org/10.1111/2041-210X.13120 
• Buderman, F. E., M. B. Hooten, M. W. Alldredge, E. M. Hanks, and J. S. Ivan. 2018. Time-varying predatory behavior is primary predictor of fine-scale movement of wildland-urban cougars. Movement Ecology 6:22. https://doi.org/10.1186/s40462-018-0140-6 
• Ivan, J. S., A. E. Seglund, R. L. Truex, and E. S. Newkirk. 2018. Mammalian responses to changed forest conditions resulting from bark beetle outbreaks in the southern Rocky Mountains. Ecosphere 9:e02369. https://doi.org/10.1002/ecs2.2369
• Olson, L. E., J. R. Squires, E. K. Roberts, J. S. Ivan, and M. Hebblewhite. 2018. Sharing the same slope: behavioral responses of a threatened mesocarnivore to motorized and nonmotorized winter recreation. Ecology and Evolution 8:8555-8572. https://doi.org/10.1002/ece3.4382 
• Olson, L. E., J. R. Squires, E. K. Roberts, A. D. Miller, J. S. Ivan, and M. Hebblewhite. 2017. Modeling large-scale winter recreation terrain selection with implications for recreation management and wildlife. Applied Geography 86:66-91.  https://doi.org/10.1016/j.apgeog.2017.06.023
• Buderman, F. E., M. B. Hooten, J. S. Ivan, and T. M. Shenk. 2018. Large-scale movement behavior in a reintroduced predator population. Ecography 41:126-139.  https://onlinelibrary.wiley.com/doi/ftr/10.1111/ecog.03030
• Baigas, P. E., J. R. Squires, L. E. Olsen, J. S. Ivan, and E. K. Roberts. 2017. Using environmental features to model highway crossing behavior of Canada lynx in the Southern Rocky Mountains. Landscape and Urban Planning 157:200-213.  https://doi.org/10.1016/j.landurbplan.2016.06.007
• Hooten, M. B., F. E. Buderman, B. M. Brost, E. M. Hanks, and J. S. Ivan. 2016. Hierarchical animal movement models for population-level inference. Environmetrics 27:322-333.  https://doi.org/10.1002/env.2402
• Ivan, J. S., and T. M. Shenk. 2016. Winter diet and hunting success of Canada lynx in Colorado. The Journal of Wildlife Management 80:1049-1058. https://doi.org/10.1002/jwmg.21101
• Ivan, J. S., and E. S. Newkirk. 2016. CPW Photo Warehouse: a custom database to facilitate archiving, identifying, summarizing, and managing photo data collected from camera traps. Methods in Ecology and Evolution 7:499-504. https://doi.org/10.1111/2041-210X.12503
• Buderman, F. E., M. B. Hooten, J. S. Ivan, and T. M. Shenk. 2016. A functional model for characterizing long-distance movement behavior. Methods in Ecology and Evolution 7:264-273. https://doi.org/10.1111/2041-210X.12465
• Ellis, M. M., J. S. Ivan, J. M. Tucker, and M. K. Schwartz. 2015. rSPACE: Spatially based power analysis for conservation and ecology. Methods in Ecology and Evolution 6:621-625. https://doi.org/10.1111/2041-210X.12369
• Ivan, J. S., G. C. White, and T. M. Shenk. 2014. Density and demography of snowshoe hares in central Colorado. The Journal of Wildlife Management 78:580–594. https://doi.org/10.1002/jwmg.695
• Gerber, B. D., J. S. Ivan, and K. P. Burnham. 2014. Estimating the abundance of rare and elusive carnivores from photographic-sampling data when the population size is very small. Population Ecology 56:463-470. https://doi.org/10.1007/s10144-014-0431-8
• Trainor, A. M., O. J. Schmitz, J. S. Ivan, and T. M. Shenk. 2014. Enhancing species distribution modeling by characterizing predator–prey interactions. Ecological Applications 24:204–216. https://doi.org/10.1890/13-0336.1
• Ellis, M. M., J. S. Ivan, and M. K. Schwartz. 2014. Spatially explicit power analyses for occupancy-based monitoring of wolverine in the U.S. Rocky Mountains. Conservation Biology 28:52–62. https://doi.org/10.1111/cobi.12139
• Ivan J. S., G. C. White, and T. M. Shenk. 2013. Using simulation to compare methods for estimating density from capture-recapture data. Ecology 94:817–826. https://doi.org/10.1890/12-0102.1
• Ivan, J. S., G. C. White, and T. M. Shenk. 2013. Using auxiliary telemetry information to estimate animal density from capture-recapture data. Ecology 94:809–816. https://doi.org/10.1890/12-0101.1
• Stromborg, K. L., J. S. Ivan, J. K. Netto, and C. R. Courtney. 2012. Survivorship and mortality patterns of double-crested cormorants at Spider Island, Wisconsin, 1988–2006. Waterbirds 35:31–39. https://doi.org/10.1675/063.035.sp105

Education

• Ph.D., Fish, Wildlife and Conservation Biology — Colorado State University, 2011
• M.S., Wildlife Biology — University of Montana, 2000
• B.S., Wildlife Science — Purdue University, 1997

Current or Recent Positions

• Wildlife Researcher — Colorado Parks and Wildlife, 2010–Present
• Refuge Biologist — U.S. Fish and Wildlife Service, 2003-2005
• Piping Plover Recovery Biologist — The Nature Conservancy, 2000-2003

Current or Recent Research Projects

• Using Non-Invasive Genetic Sampling to Estimate Mountain Lion and Bobcat Abundance, Age Structure and Diet
• Mountain Lion Demographics and Human Interactions Along the Urban-Exurban Front Range

Areas of Interest and Expertise

My research interest focuses on large carnivore and ungulate ecology, population parameter estimation, habitat use, avian ecology and population sampling. 

Streaming Videos

Mountain lions in Colorado – a 4 part series

• Series Trailer
• Part 1: Mountain Lion Biology & Historical Perspective 
• Part 2: Mountain Lion Habitat & Human Expansion 
• Part 3: Hunting
• Part 4: What to do if you encounter a mountain lion
• Colorado Mountain Lions (all 4 videos combined)

Select Publications

• Crews, S., N. D. Rayl, M. W. Alldredge, E. J. Bergman, C. R. Anderson Jr., E. H. VanNatta, J. D. Holbrook, and G. Bastille-Rousseau. 2025. Hierarchy in structuring of resource selection: understanding elk selection across space, time, and movement strategies. Ecology and Evolution 15:e71097. https://doi.org/10.1002/ece3.71097
• Crews, S., N. D. Rayl, M. W. Alldredge, E. J. Bergman, C. R. Anderson Jr., and G. Bastille-Rousseau. 2025. Drivers of spring migration phenology in Rocky Mountain elk. Scientific Reports 15:7807. https://doi.org/10.1038/s41598-025-91947-4
• Venumière-Lefebvre, C. C., H. E. Johnson, S. W. Breck, M. W. Alldredge, and K. R. Crooks. 2025. Bears avoid residential neighborhoods in response to the experimental reduction of anthropogenic attractants. Frontiers in Ecology and Evolution 13:1657106. doi: 10.3389/fevo.2025.1657106
• Fountain-Jones, N. M., S. Kraberger, R. B. Gagne, D. R. Trumbo, P. E. Salerno, W. C. Funk, K. Crooks, R. Biek, M. W. Alldredge, K. Logan, G. Baele, S. Dellicour, H. B. Ernest, S. VandeWoude, S. Carver, and M. E. Craft. 2021. Host relatedness and landscape connectivity shape pathogen spread in the puma, a large secretive carnivore​. Communications Biology 4:12. doi: 10.1038/s42003-020-01548-2.
• Gagne, R. B., S., Kraberger, R. McMinn, D. R. Trumbo, C. R. Anderson Jr, K. A. Logan, M. W. Alldredge, K. Griffin and S. VandeWoude. 2021. Viral sequen​ces recovered from puma tooth DNA reconstruct statewide viral phylogenies. Frontiers in Ecology and Evolution 9:734462. doi: 10.3389/fevo.2021.734462​
• Kraberger, S., N. M. Fountain-Jones, R. B. Gagne, J. Malmberg, N. G. Dannemiller, K. Logan, M. Alldredge, A. Varsani, K. R. Crooks, M. Craft, S. Carver, and S. VandeWoude. 2020. Frequent cross-species transmissions of foamy virus between domestic and wild felids. Virus Evolution 6(1):vez058. https://doi.org/10.1093/ve/vez058
• Bishop, C. J., M. W. Alldredge, D. P. Walsh, E. J. Bergman, C. R. Anderson Jr, D. Kilpatrick, J. Bakel, and C. Febvre. 2019. A noninvasive automated device for remotely collaring and weighing mule deer. Wildlife Society Bulletin 43:717–725. https://doi.org/10.1002/wsb.1034
• Trumbo, D. R., P. E. Salerno, K. A. Logan, M. W. Alldredge, R. B. Gagne, C. P. Kozakiewicz, S. Kraberger, N. M. Fountain‐Jones, M. E. Craft, S. Carver, H. B. Ernest, K. R. Crooks, S. VandeWoude, and W. C. Funk. 2019. Urbanization impacts apex predator gene flow but not genetic diversity across an urban‐rural divide. Molecular Ecology 28:4926–4940. https://doi.org/10.1111/mec.15261
• Alldredge, M. W., F. E. Buderman, and K. A. Blecha. 2019. Human–Cougar interactions in the wildland–urban interface of Colorado's front range. Ecology and Evolution 9:10415–10431.  https://doi.org/10.1002/ece3.5559
• Alldredge, M. W., T. Blecha, and J. H. Lewis. 2019. Less invasive monitoring of cougars in Colorado's Front Range. Wildlife Society Bulletin 43:222–230. https://doi.org/10.1002/wsb.971 
• Kechejian, S. R., N. Dannemiller, S. Kraberger, C. Ledesma-Feliciano, J. Malmberg, M. Roelke Parker, M. Cunningham, R. McBride, S. P. Riley, W. T. Vickers, K. Logan, M. Alldredge, K. Crooks, M. Löchelt, S. Carver, and S. VandeWoude. 2019. Feline foamy virus is highly prevalent in free-ranging puma concolor from Colorado, Florida and Southern California. Viruses 11:359. https://doi.org/10.3390/v11040359
• Kirby, R., H. E. Johnson, M. W. Alldredge, and J. N. Pauli. 2019. The cascading effects of human food on hibernation and cellular aging in free-ranging black bears. Scientific Reports 9:2197. https://doi.org/10.1038/s41598-019-38937-5
• Peterson, M. E., C. R. Anderson Jr, M. W. Alldredge, and P. F. Doherty Jr. 2018. Using maternal mule deer movements to estimate timing of parturition and assist fawn captures. Wildlife Society Bulletin 42:616–621. https://doi.org/10.1002/wsb.935
• Buderman, F. E., M. B. Hooten, M. W. Alldredge, E. M. Hanks, and J. S. Ivan. 2018. Time-varying predatory behavior is primary predictor of fine-scale movement of wildland-urban cougars. Movement Ecology 6:22. https://doi.org/10.1186/s40462-018-0140-6
• Blecha, K. A., R. B. Boone, and M. W. Alldredge. 2018. Hunger mediates apex predator's risk avoidance response in wildland-urban interface. Journal of Animal Ecology 87:609–622. https://doi.org/10.1111/1365-2656.12801
• Lewis, J. S., K. A. Logan, M. W. Alldredge, S. Carver, S. N. Bevins, M. Lappin, S. VandeWoude, and K. R. Crooks. 2017. The effects of demographic, social, and environmental characteristics on pathogen prevalence in wild felids across a gradient of urbanization. PLoS One 12(11):e0187035. https://doi.org/10.1371/journal.pone.0187035
• Lewis, J. S., K. A. Logan, M. W. Alldredge, D. M. Theobald, S. VandeWoude, and K. R. Crooks. 2017. Contact networks reveal potential for interspecific interactions of sympatric wild felids driven by space use. Ecosphere 8(3):e01707. https://doi.org/10.1002/ecs2.1707
• Kirby, R., M. W. Alldredge, and J. N. Pauli. 2017. Environmental, not individual, factors drive markers of biological aging in black bears. Evolutionary Ecology 31:571–584. https://doi.org/10.1007/s10682-017-9885-4
• Moss, W. E., M. W. Alldredge, K. A. Logan, and J. N. Pauli. 2016. Human expansion precipitates niche expansion for an opportunistic apex predator (Puma concolor). Scientific Reports 6:39639. https://doi.org/10.1038/srep39639
• Kirby, R., M. W. Alldredge, and J. N. Pauli. 2016. The diet of black bears tracks the human footprint across a rapidly developing landscape. Biological Conservation 200:51–59. https://doi.org/10.1016/j.biocon.2016.05.012
• Moss, W. E., M. W. Alldredge, and J. N. Pauli. 2015. Quantifying risk and resource use for a large carnivore in an expanding urban-wildland interface. Journal of Applied Ecology 53:371–378. https://doi.org/10.1111/1365-2664.12563
• Lewis, J. S., K. A. Logan, M. W. Alldredge, L. L. Bailey, S. VandeWoude, and K. R. Crooks. 2015. The effects of urbanization on population density, occupancy, and detection probability of wild felids. Ecological Applications 25:1880–1895. https://doi.org/10.1890/14-1664.1
• Blecha, K. A., and M. W. Alldredge. 2015. Improvements on GPS location cluster analysis for the prediction of large carnivore feeding activities: ground-truth detection probability and inclusion of activity sensor measures. PLoS One 10(9): e0138915. https://doi.org/10.1371/journal.pone.0138915 
• Alldredge, M. W., D. P. Walsh, L. L. Sweanor, R. B. Davies, and A. Trujillo. 2015. Evaluation of translocation of black bears involved in human-bear conflicts in South-central Colorado. Wildlife Society Bulletin 39:334–340. https://doi.org/10.1002/wsb.526 
• Hanks, E. M., M. B. Hooten, and M. W. Alldredge. 2015. Continuous-time discrete-space models for animal movement. The Annals of Applied Statistics 9:145-165. https://doi.org/10.1214/14-AOAS803
• Hooten, M. B., E. M. Hanks, D. S. Johnson, and M. W. Alldredge. 2013. Reconciling resource utilization and resource selection functions. Journal of Animal Ecology 82:1146–1154. https://doi.org/10.1111/1365-2656.12080 
• Hooten, M. B., E. M. Hanks, D. S. Johnson, and M. W. Alldredge. 2014. Temporal variation and scale in movement-based resource selection functions. Statistical Methodology 17:82–98. https://doi.org/10.1016/j.stamet.2012.12.001
• Bevins, S. N., S. Carver, E. E. Boydston, L. M. Lyren, M. Alldredge, K. A. Logan, S. P. D. Riley, R. N. Fisher, T. W. Vickers, W. Boyce, M. Salman, M. R. Lappin, K. R. Crooks, and S. VandeWoude. 2012. Three pathogens in sympatric populations of pumas, bobcats, and domestic cats: implications for infectious disease transmission. PLoS One 7(2):e31403. https://doi.org/10.1371/journal.pone.0031403 
• Horne, J. S., K. M. Strickler, and M. Alldredge. 2011. Quantifying the importance of patch-specific changes in habitat to metapopulation viability of an endangered songbird. Ecological Applications 21:2478–2486. https://doi.org/10.1890/10-2328.1 
• Alldredge, M. W. 2011. Cougars on the edge. The Wildlife Professional 5:72–76. 
• Simons, T. R., K. H. Pollock, J. M. Wettroth, M. W. Alldredge, K. Pacifici, and J. Brewster. 2009. Sources of measurement error, misclassification error, and bias in auditory avian point count data. Pages 237–254 in Thomson D.L., E.G. Cooch, M.J. Conroy, editors. Modeling demographic processes in marked populations. Environmental and ecological statistics. Volume 3. Springer, Boston, USA. https://doi.org/10.1007/978-0-387-78151-8_10 
• Bevins, S. N., J. A. Tracey, S. P. Franklin, V. L. Schmit, M. L. Macmillan, K. L. Gage, M. E. Schriefer, K. A. Logan, L. L. Sweanor, M. W. Alldredge, C. Krumm, W. M. Boyce, W. Vickers, S. P. Riley, L. M. Lyren, E. E. Boydston, R. N. Fisher, M. E. Roelke, M. Salman, K. R. Crooks, and S. VandeWoude. 2009. Wild felids as hosts for human plague, Western United States. Emerging Infectious Diseases 15:2021–2024. https://doi.org/10.3201/eid1512.090526
• Alldredge, M. W., T. R. Simons, K. H. Pollock, and K. Pacifici. 2007. A field evaluation of the effectiveness of distance sampling and double independent observers to estimate detection probability in aural avian point counts. Avian Conservation and Ecology - Écologie et conservation des oiseaux 2:13. http://www.ace-eco.org/vol2/iss2/art13/

Education

• Ph.D., Zoology and Biomathematics — North Carolina State University, 2004
• M.S., Biomathematics — North Carolina State University, 2002
• M.S., Wildlife Resources — University of Idaho, 1999
• B.S., Mechanical Engineering — University of Colorado, 1994

Current or Recent Positions

• Wildlife Researcher — Colorado Parks and Wildlife, 2006-Present
• Post-Doctoral Researcher — USGS Cooperative Fish and Wildlife Research Unit, North Carolina State University, 2004-2006
• Carnivore Biologist —Wyoming Game and Fish Department, 1995